The soldier beetles (Cantharidae) are relatively soft-bodied, straight-sided
beetle
Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
s. They are
cosmopolitan
Cosmopolitan may refer to:
Internationalism
* World citizen, one who eschews traditional geopolitical divisions derived from national citizenship
* Cosmopolitanism, the idea that all of humanity belongs to a single moral community
* Cosmopolitan ...
in distribution. One of the first described species has a color pattern reminiscent of the
red coats of early British soldiers, hence the common name. They are also known commonly as leatherwings because of their soft
elytra
An elytron (; ; : elytra, ) is a modified, hardened forewing of beetles (Coleoptera), though a few of the true bugs (Hemiptera) such as the family Schizopteridae are extremely similar; in true bugs, the forewings are called hemelytra (sometime ...
.
Historically, these beetles were placed in a superfamily "Cantharoidea", which has been subsumed by the superfamily
Elateroidea
The Elateroidea are a large superfamily of beetles. It contains the familiar click beetles, fireflies, and soldier beetles and their relatives. It consists of about 25,000 species.
Description
Elateroidea is a morphologically diverse group ...
; the name is still sometimes used as a rankless grouping, including the families Cantharidae,
Lampyridae
The Lampyridae are a family of elateroid beetles with more than 2,000 described species, many of which are light-emitting. They are soft-bodied beetles commonly called fireflies, lightning bugs, or glowworms for their conspicuous production ...
,
Lycidae,
Omethidae (which includes
Telegeusidae),
Phengodidae, and
Rhagophthalmidae.
Soldier beetles often feed on nectar and pollen as well as predating other small insects. The larvae are caterpillar like, dark colored, active and covered in fine hairs, earning them the name velvet worms. They feed on the ground and in foliage hunting eggs, small insects, snails and other small creatures.
Evolutionary history
The oldest described member of the family is ''
Molliberus'' from the
Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
(early
Albian
The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...
) aged El Soplao amber from Cantabria, Spain, belonging to the tribe
Cantharini in the subfamily
Cantharinae. Other described genera include 6 from the early
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
(early
Cenomanian
The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
) aged
Burmese amber
Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. Th ...
, with 5 belonging to Cantharinae and one to
Malthininae
The soldier beetles (Cantharidae) are relatively soft-bodied, straight-sided beetles. They are Cosmopolitan distribution, cosmopolitan in distribution. One of the first described species has a color pattern reminiscent of the Red coat (Bri ...
, and ''
Katyacantharis'', from the Cenomanian aged Agdzhakend amber of Azerbaijan, suggested to belong to Cantharinae. Indeterminate specimens have been reported from the
Aptian
The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...
aged
Koonwarra fossil bed of the
Strzelecki Group
The Halibut Field is an oil field, within the Gippsland Basin offshore of the Australian state of Victoria (Australia), Victoria. The oil field is located approximately 64 km offshore of southeastern Australia. The total area of this field ...
, Australia and the
Barremian
The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
aged
Lebanese amber
Lebanese amber is fossilized resin found in Lebanon and its surroundings. It dates back approximately 130-125 million years to the Barremian of the Early Cretaceous. It formed on what was then the northern coast of Gondwana, believed to be a tropic ...
.
Subfamilies, tribes and selected genera
Five subfamilies are normally accepted:
Cantharinae
*
tribe
The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...
Cantharini
** ''
Cantharis''
** ''
Rhagonycha''
* tribe Podabrini
** ''
Podabrus''
Chauliognathinae
* tribe Chauliognathini
** ''
Belotus''
** ''
Chauliognathus''
* tribe Ichthyurini
** ''
Trypherus''
Dysmorphocerinae
* ''
Afronycha''
Wittmer, 1949 - central-southern Africa
* ''
Asilis''
Broun, 1893 - New Zealand
* ''
Compsonycha''
* ''
Dysmorphocerus''
Solier, 1849
* ''
Flabelloontelus''
* ''
Geigyella''
Wittmer, 1972 - New Guinea
* ''
Hansasilis''
* ''
Heteromastix''
Boheman, 1858 - Australia
* ''
Hyponotum''
* ''
Micronotum''
* ''
Neoontelus''
Wittmer, 1972 - New Zealand
* ''
Oontelus''
Solier, 1849 - S. America
* ''
Plectocephalon''
* ''
Plectonotum''
Gorham, 1891 -
Americas
The Americas, sometimes collectively called America, are a landmass comprising the totality of North America and South America.''Webster's New World College Dictionary'', 2010 by Wiley Publishing, Inc., Cleveland, Ohio. When viewed as a sing ...
Malthininae
* tribe Malchinini
** ''
Macrocerus''
Motschulsky, 1845 - Europe (synonym ''Malchinus'')
* tribe Malthinini
** ''
Caccodes''
Sharp, 1885 - Central America, Pacific islands
** ''
Malthinellus''
Kiesenwetter, 1874 - Japan
** ''
Malthinus''
Latreille, 1805 - Japan, Europe, N. America
* tribe Malthodini
** ''
Frostia''
Bert. ex Guill.
** ''
Malthodes''
Kiesenwetter, 1852 - mostly Europe, N. America & New Zealand
** †''
Archaeomalthodes''
Hsiao et al. 2016 Burmese amber, Myanmar, Cenomanian
Silinae
* tribe Silini
** ''
Cordylocera''
Guérin-Méneville, 1830
** ''
Silis''
Charpentier, 1825
* tribe Tytthonyxini
** ''
Tytthonyx''
LeConte, 1851
Reproduction
Large males of the soldier beetle exercise choice for larger females. Body size correlates with the abilities of males to secure females, and of females to evade males.
See also
*
List of Cantharidae genera
References
External links
Delta keyKeys for the identification of British Cantharidae*
*
{{Authority control
Biological pest control beetles
*
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