''Camponotus herculeanus'' (or Hercules ant) is a species of

ant Ants are Eusociality, eusocial insects of the Family (biology), family Formicidae and, along with the related wasps and bees, belong to the Taxonomy (biology), order Hymenoptera. Ants evolved from Vespoidea, vespoid wasp ancestors in the Cre ...

in the genus ''

Camponotus Carpenter ants (''Camponotus'' spp.) are a genus of large ants (workers ) indigenous to many parts of the world. True carpenter ants build nests inside wood, consisting of galleries chewed out with their mandibles or jaws, preferably in dead, ...

'', the carpenter ants, occurring in Northern Eurasia, from Norway to Eastern Siberia, and North America. First described as ''

Formica ''Formica'' is a genus of ants of the subfamily Formicinae, including species commonly known as wood ants, mound ants, thatching ants, and field ants. ''Formica'' is the type genus of the Formicidae, and of the subfamily Formicinae. The type ...

herculeana'' by

Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...

1758 Events January–March * January 1 – Swedish biologist Carl Linnaeus (Carl von Linné) publishes in Stockholm the first volume (''Animalia'') of the 10th edition of ''Systema Naturae'', the starting point of modern zoologic ...

, the species was moved to ''

'' by

Mayr Mayr is a German surname. Notable people with the surname include: * Andrea Mayr (born 1979), Austrian female long-distance runner * Ernst Mayr (1904–2005), German American evolutionary biologist * Franz Xaver Mayr (1875–1965), Austrian gastro ...

in 1861.

Description

The colony of ''Camponotus herculeanus'' consists of one or several wingless females (queens), some fertile males, and three castes of sterile workers, known as majors, intermediates, and minors, in decreasing order of size. The queens are large, about in length, and are blackish in colour. The males are a similar colour but about half the size of the queens. The workers usually have blackish heads and gasters, and dark reddish-brown

mesosoma The mesosoma is the middle part of the body, or tagma, of arthropods whose body is composed of three parts, the other two being the prosoma and the metasoma. It bears the legs, and, in the case of winged insects, the wings. Wasps, bees and a ...

s, petioles and legs. In majors, the scapes (the long segments of the antenna, before the elbow) are shorter than the length of the head; in intermediates they are about the same length, and in minors, they extend well beyond the back of the head. The head and the dorsal surfaces of the mesosoma and gaster of the largest majors are bristly.

Distribution and habitat

''Camponotus herculeanus'' has a widespread distribution in the Northern Hemisphere, being present in most of Europe, Central and Northern Asia, Canada and the United States. It is common in mountainous regions and is the dominant ant species in mountainous and northerly parts of North America. It occupies a range of habitats including various types of conifer and hardwood forests, clearings, oak scrubland, disturbed areas, pastures and seashore grassland.

Ecology

Nests of ''Camponotus herculeanus'' are built in timber, living or rotting trees, stumps, fallen logs and occasionally the structural timbers of buildings. The ants use their strong jaws to excavate galleries and chambers under the bark or in the wood, with a preference for damp wood or timber with fungal decay. In standing trees, their tunnels sometimes extend for above the ground. Satellite colonies, linked to the original nest by underground tunnels, may develop nearby, often in warmer, drier locations. These house older larvae, pupae, winged reproductives and workers, with the eggs and younger larvae remaining in the main nest. A colony of ''Camponotus herculeanus'' contains several wingless females, which may be unrelated. Winged reproductives are produced in late summer and overwinter in the colony, emerging to fly in swarms on warm spring days. The workers become active in spring and forage in the vicinity of the nest. They tend

aphid Aphids are small sap-sucking insects in the Taxonomic rank, family Aphididae. Common names include greenfly and blackfly, although individuals within a species can vary widely in color. The group includes the fluffy white Eriosomatinae, woolly ...

s, and the larvae of the silvery blue butterfly (''Glaucopsyche lygdamus''), which often feeds on the lupine ''

Lupinus bakeri ''Lupinus'', commonly known as lupin, lupine, or regionally bluebonnet, is a genus of plants in the legume family Fabaceae. The genus includes over 199 species, with centres of diversity in North and South America. Smaller centres occur in No ...

''. The diet consists of the honeydew produced by sap-sucking insects and the ants also consume any insect larvae that they encounter. The ant cricket ''

Myrmecophilus pergandei ''Myrmecophilus pergandei'', the eastern ant cricket, is a species of ant cricket in the family Myrmecophilidae. It is found in North America. It is a wingless cricket that is an obligate kleptoparasite of ants living in their nests. They lack ...

'' sometimes lives in the colony, where it is tolerated by the ants.

References

External links

* {{Taxonbar, from=Q162870 herculeanus Hymenoptera of Asia Hymenoptera of Europe Hymenoptera of North America Insects described in 1758 Taxa named by Carl Linnaeus