crayfish
Crayfish are freshwater crustaceans belonging to the clade Astacidea, which also contains lobsters. In some locations, they are also known as crawfish, craydids, crawdaddies, crawdads, freshwater lobsters, mountain lobsters, rock lobsters, mu ...
from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the second largest freshwater crayfish genus inhabiting the Northern Hemisphere, with only sixty fewer species than the genus ''
Procambarus
''Procambarus'' is a genus of crayfish in the family Cambaridae, all native to North and Central America. It includes a number of troglobitic species, and the marbled crayfish (''marmorkrebs''), which is parthenogenetic. Originally described as a ...
''. Though ''Cambarus'' are varied across species, the two terminal elements that make up the male form I gonopod form ninety degree angles with the central appendage, allowing for their identification. Unlike the genus ''
Procambarus
''Procambarus'' is a genus of crayfish in the family Cambaridae, all native to North and Central America. It includes a number of troglobitic species, and the marbled crayfish (''marmorkrebs''), which is parthenogenetic. Originally described as a ...
'' whose first
pleopod
The decapod ( crustaceans such as a crab, lobster, shrimp or prawn) is made up of 20 body segments grouped into two main body parts: the cephalothorax and the pleon (abdomen). Each segment may possess one pair of appendages, although in various ...
tends to have three processes at the tip, ''Cambarus'' has only one or two. ''Cambarus'' reach 17–26 mm
carapace
A carapace is a Dorsum (biology), dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tor ...
lengths in their first year, while average adult carapace length ranges from 55–62 mm.
As a genus containing nearly 100 species, ''Cambarus's'' coloration is variable. ''
Cambarus bartonii
''Cambarus bartonii'' is a species of crayfish native to eastern North America, where it is called the common crayfish or Appalachian brook crayfish.
''C. bartonii'' was the first crayfish to be described from North America, when Johan Christian ...
'' is dark brown, while species like ''
Cambarus pauleyi
''Cambarus'' is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the se ...
'' range from subtle to vibrant blues and reds. Other species are light green or grayish in color.
Biogeography
Most species of ''Cambarus'' are restricted to the United States and Canada. They are distributed along the eastern coast, extending from New Brunswick to northern Florida. However, the genus extends as far westward as the Rocky Mountains of Wyoming and Colorado, inhabiting a variety of
freshwater
Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...
environments.
Habitat
''Cambarus'' occupy a range of freshwater environments including streams, rivers, lakes, and burrows. Burrowing species of the genus include ''
Cambarus dubius
''Cambarus'' is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the se ...
''. ''Cambarus'' also include many cave-dwelling species, both stygobites and stygophiles. While salinity and temperature changes minimally affect ''Cambarus'', the genus has shown an intolerance to pollution.
Ecology
Diet
Like other crayfish, ''Cambarus'' are foragers. Diets are largely plant-based, though ''Cambarus'' also consume small marine organisms like molluscs, larvae, tadpoles and amphibian eggs. ''Cambarus'' consume small rodents or birds when available. In their first year, ''Cambarus'' typically consume 1-4% of their overall body-weight each day. The genus is central to many freshwater food webs as they help maintain water quality through consumption of
algae
Algae (; singular alga ) is an informal term for a large and diverse group of photosynthetic eukaryotic organisms. It is a polyphyletic grouping that includes species from multiple distinct clades. Included organisms range from unicellular mic ...
.
Vulnerability
One of the largest crayfish genera, ''Cambarus'' includes a sizable number of vulnerable species. Cave-dwellers like '' Cambarus jonesi'' are at risk due to their lack of
genetic diversity
Genetic diversity is the total number of genetic characteristics in the genetic makeup of a species, it ranges widely from the number of species to differences within species and can be attributed to the span of survival for a species. It is dis ...
and low population count. Other species like ''Cambarus veteranus'' are at risk due to human practices like logging and mining, which increase sediment amounts in freshwater environments. Increased sediment causes these freshwater environments to be uninhabitable, and ''Cambarus'' are forced to relocate as a result.
Behavior
Growth
Molting
In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is the manner in which an animal routinely casts off a part of its body (often, but not always, an outer ...
occurs among ''Cambarus'' approximately 5-10 times during their first year, and 3-5 times during subsequent years. ''Cambarus'' remain relatively inactive during periods of molting, as the shedding of
chitinous
Chitin ( C8 H13 O5 N)n ( ) is a long-chain polymer of ''N''-acetylglucosamine, an amide derivative of glucose. Chitin is probably the second most abundant polysaccharide in nature (behind only cellulose); an estimated 1 billion tons of chiti ...
exoskeletons leaves them more vulnerable to predation and injury. Many species of ''Cambarus'' continue to grow well into
adult
An adult is a human or other animal that has reached full growth. In human context, the term ''adult'' has meanings associated with social and legal concepts. In contrast to a " minor", a legal adult is a person who has attained the age of major ...
hood.
Reproduction
''Cambarus'' typically mate in the early spring. Both ''
Cambarus bartonii
''Cambarus bartonii'' is a species of crayfish native to eastern North America, where it is called the common crayfish or Appalachian brook crayfish.
''C. bartonii'' was the first crayfish to be described from North America, when Johan Christian ...
'' and ''
Cambarus robustus
''Cambarus robustus'', known generally as the robust crayfish or Big Water crayfish, is a species of crayfish in the family Cambaridae
The Cambaridae are the largest of the four families of freshwater crayfish, with over 400 Species. Most of ...
'' only mate once during their three-year life span, with females of both carrying fewer eggs than those of the genus '' Orconectes''.
Gallery
Classification
The genus ''Cambarus'' contains around 100 species, many of which are listed on the IUCN Red List. Species in the genus were formerly divided among 12
subgenera
In biology, a subgenus (plural: subgenera) is a taxonomic rank directly below genus.
In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between t ...
. In a 2017 study, these subgenera were found to lack any phylogenetic validity and were therefore eliminated.
*''
Cambarus andersoni
''Cambarus'' is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the sec ...
Cambarus bartonii
''Cambarus bartonii'' is a species of crayfish native to eastern North America, where it is called the common crayfish or Appalachian brook crayfish.
''C. bartonii'' was the first crayfish to be described from North America, when Johan Christian ...
Cambarus batchi
''Cambarus batchi'', the bluegrass crayfish, is a species of crayfish in the family Cambaridae. It is endemic to Kentucky, known as the "Bluegrass State" which lead to the common name.
The IUCN conservation status of ''Cambarus batchi'' is "L ...
'' Schuster, 1973 – bluegrass crayfish
*'' Cambarus bouchardi'' Hobbs, 1970 – Big South Fork crayfish
*''
Cambarus brachydactylus
''Cambarus'' is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the se ...
'' Hobbs, 1953
*''
Cambarus brimleyorum
''Cambarus'' is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the se ...
'' Cooper, 2006
*'' Cambarus buntingi'' R. W. Bouchard, 1973 – Bunting crayfish
*''
Cambarus callainus
The Big Sandy crayfish, ''Cambarus callainus'', are freshwater, tertiary burrowing crustaceans of the family Cambaridae. They are found in the streams and rivers of Appalachia in Virginia, West Virginia, and Kentucky, in what is known as the ...
Cambarus carolinus
''Cambarus'' is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the se ...
Cambarus clairitae
''Cambarus'' is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the se ...
Cambarus coosae
''Cambarus'' is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the se ...
Cambarus diupalma
''Cambarus'' is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the se ...
'' Jones and Eversole, 2015
*'' Cambarus doughertyensis'' Cooper & Skelton, 2003 - Dougherty Burrowing Crayfish
*''
Cambarus dubius
''Cambarus'' is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the se ...
Cambarus fasciatus
''Cambarus fasciatus'', the Etowah crayfish, is a species of crayfish in the family Cambaridae. It is endemic
Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country ...
'' Hobbs, 1981 – Etowah crayfish
*''
Cambarus friaufi
''Cambarus'' is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the se ...
'' Hobbs, 1953 – hairy crayfish
*''
Cambarus gentryi
''Cambarus gentryi'', the linear cobalt crayfish, is small burrowing species of crayfish in the family Cambaridae, notable for its blue carapace
A carapace is a Dorsum (biology), dorsal (upper) section of the exoskeleton or shell in a number ...
Cambarus hystricosus
''Cambarus'' is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the se ...
Cambarus loughmani
''Cambarus loughmani'', the blue Teays mudbug, is a species of burrowing crayfish endemic to the pre-glacial Teays River Valley in West Virginia. The species was previously considered to be part of the Cambarus dubius
''Cambarus'' is a lar ...
Cambarus monongalensis
''Cambarus monongalensis'', the blue crayfish or Monongahela crayfish, is a species of burrowing crayfish native to Pennsylvania and West Virginia. It has also been found recently in Ohio. The common name refers to the Monongahela River, with ...
'' Ortmann, 1905
*''
Cambarus nerterius
''Cambarus nerterius'', the Greenbrier cave crayfish, is a species of crayfish in the family Cambaridae. It is Endemism, endemic to the state of West Virginia in the United States. It is found only in or immediately adjacent to caves in Greenbrier ...
'' Hobbs, 1964 – Greenbrier cave crayfish
*''
Cambarus nodosus
''Cambarus nodosus'' is a species of crayfish in the family Cambaridae. It is found in North America.
The IUCN
The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural R ...
'' R. W. Bouchard & Hobbs, 1976
*''
Cambarus obeyensis
''Cambarus obeyensis'', the Obey crayfish, is a species of crayfish in the family Cambaridae. It is found in North America.
The IUCN
The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of ...
Cambarus ortmanni
''Cambarus'' is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the se ...
'' Williamson, 1907 – Ortmann mudbug
*''
Cambarus parrishi
''Cambarus parrishi'', the Hiwassee headwaters crayfish, is a species of crayfish
Crayfish are freshwater crustaceans belonging to the clade Astacidea, which also contains lobsters. In some locations, they are also known as crawfish, crayd ...
'' Hobbs, 1981 – Hiwassee headwater crayfish
*''
Cambarus parvoculus
''Cambarus'' is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the sec ...
Cambarus pauleyi
''Cambarus'' is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the se ...
Cambarus pyronotus
''Cambarus pyronotus'', the fireback crayfish, is a species of crayfish
Crayfish are freshwater crustaceans belonging to the clade Astacidea, which also contains lobsters. In some locations, they are also known as crawfish, craydids, crawd ...
'' R. W. Bouchard, 1978 – fireback crayfish
*'' Cambarus reburrus'' Prins, 1968 – French Broad crayfish
*'' Cambarus reduncus'' Hobbs, 1956
*''
Cambarus reflexus
''Cambarus'' is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the sec ...
'' Hobbs, 1981
*''
Cambarus robustus
''Cambarus robustus'', known generally as the robust crayfish or Big Water crayfish, is a species of crayfish in the family Cambaridae
The Cambaridae are the largest of the four families of freshwater crayfish, with over 400 Species. Most of ...
'' Girard, 1852 – big water crayfish
*'' Cambarus rusticiformis'' Rhoades, 1944 – Depression crayfish
*''
Cambarus sciotensis
''Cambarus'' is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the sec ...
Cambarus smilax
''Cambarus smilax'', the Greenbrier crayfish, is a crayfish in the order Decapoda, family Cambaridae, genus Cambarus. It is most closely related to ''Cambarus robustus''. It is endemic to the Greenbrier River watershed in Pocahontas, Greenbrie ...
Cambarus speleocoopi
''Cambarus speleocoopi'', the Sweet Home Alabama cave crayfish, is a small, freshwater crayfish endemic to Marshall County, Alabama
Marshall County is a county of the state of Alabama, United States. As of the 2020 census the population was ...
Cambarus sphenoides
''Cambarus'' is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the se ...
Cambarus striatus
''Cambarus'' is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the se ...
Cambarus theepiensis
''Cambarus'' is a large and diverse genus of crayfish from the United States and Canada. The adults range in size from about 5 centimeters (2 in) up to approximately 15 centimeters (6 in).
Description
The genus ''Cambarus'' is the sec ...
'' Loughman, Foltz, Garrison and Welsh, 2013
*'' Cambarus truncatus'' Hobbs, 1981 – Oconee Burrowing Crayfish
*'' Cambarus tuckasegee'' Cooper & Schofield, 2002
*'' Cambarus unestami'' Hobbs & Hall, 1969
*'' Cambarus veitchorum'' J. E. Cooper & M. R. Cooper, 1997 – White Spring cave crayfish
*'' Cambarus veteranus'' Faxon, 1914 – Big Sandy crayfish
*'' Cambarus williami'' R. W. Bouchard & J. W. Bouchard, 1995 – Brawleys Fork crayfish
*''
Cambarus zophonastes
''Cambarus zophanastes'', also known as the Hell Creek Cave Crayfish, is named for its original location of discovery, Hell Creek Cave. It is also found in other similar habitats in Stone County and Marion County, Arkansas. These habitats i ...