''Calathus'' is a genus of

ground beetle Ground beetles are a large, cosmopolitan distribution, cosmopolitan family (biology), family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in North America and 2,700 in Europe. As of 2015, it ...

native to the

Palearctic The Palearctic or Palaearctic is a biogeographic realm of the Earth, the largest of eight. Confined almost entirely to the Eastern Hemisphere, it stretches across Europe and Asia, north of the foothills of the Himalayas, and North Africa. Th ...

(including

Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...

, the

Near East The Near East () is a transcontinental region around the Eastern Mediterranean encompassing the historical Fertile Crescent, the Levant, Anatolia, Egypt, Mesopotamia, and coastal areas of the Arabian Peninsula. The term was invented in the 20th ...

, and

North Africa North Africa (sometimes Northern Africa) is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region. However, it is sometimes defined as stretching from the Atlantic shores of t ...

). There are at least 190 described species in ''Calathus''. The genus includes species that present different forms in relation to wing length. In some species, this phenotype is determined by a single

locus Locus (plural loci) is Latin for "place". It may refer to: Mathematics and science * Locus (mathematics), the set of points satisfying a particular condition, often forming a curve * Root locus analysis, a diagram visualizing the position of r ...

gene, in which short wings are dominant. Other species, however, have lost this polymorphism, expressing either the short-winged morph or the long-winged morph. One species, ''Calanthus melanocephalus'', still presents both forms, but they're determined by the environment, specifically temperature and food conditions, instead of depending on the genotype of the individual, in what's called

polyphenism A polyphenic trait is a trait for which multiple, discrete phenotypes can arise from a single genotype as a result of differing environmental conditions. It is therefore a special case of phenotypic plasticity. There are several types of polyphen ...

Species

These 198 species belong to the genus Calathus ; Subgenus Amphyginus Haliday, 1841 : '' Calathus rotundicollis'' Dejean, 1828 ; Subgenus Bedelinus Ragusa, 1885 : '' Calathus circumseptus'' Germar, 1823 ; Subgenus Calathus (beetle), Calathus Bonelli, 1810 : '' Calathus acuticollis'' Putzeys, 1873 : '' Calathus alternans'' Faldermann, 1836 : '' Calathus arcuatus'' Gautier des Cottes, 1870 : '' Calathus atitari'' Novoa, 1999 : '' Calathus baeticus'' Rambur, 1837 : '' Calathus bosnicus'' Ganglbauer, 1891 : '' Calathus brevis'' Gautier des Cottes, 1866 : '' Calathus busii'' F.Battoni, 1984 : '' Calathus casalei'' F.Battoni, 1986 : '' Calathus distinguendus'' Chaudoir, 1846 : '' Calathus ellipticus'' Reitter, 1889 : '' Calathus erzeliki'' Schweiger, 1977 : '' Calathus focarilei'' Schatzmayr, 1947 : '' Calathus fracassii'' Heyden, 1908 : '' Calathus fuscipes'' (Goeze, 1777) : '' Calathus giganteus'' Dejean, 1828 : '' Calathus glabricollis'' Dejean, 1828 : '' Calathus granatensis'' Vuillefroy, 1866 : '' Calathus heinzianus'' F.Battoni, 1986 : '' Calathus hispanicus'' Gautier des Cottes, 1866 : '' Calathus jakupicensis'' B.V.Gueorguiev, 2008 : '' Calathus kirschenhoferianus'' F.Battoni, 1986 : '' Calathus korax'' Reitter, 1889 : '' Calathus libanensis'' Putzeys, 1873 : '' Calathus lissoderus'' Putzeys, 1873 : '' Calathus longicollis'' Motschulsky, 1865 : '' Calathus luctuosus'' (Latreille, 1804) : '' Calathus malacensis'' Nègre, 1966 : '' Calathus minutus'' Gautier des Cottes, 1866 : '' Calathus mirei'' Nègre, 1966 : '' Calathus moralesi'' Nègre, 1966 : '' Calathus muchei'' Jedlicka, 1961 : '' Calathus oertzeni'' Jeanne & F.Battoni, 1987 : '' Calathus opacus'' Lucas, 1846 : '' Calathus ordinatus'' Gautier des Cottes, 1870 : '' Calathus oreades'' Nègre, 1966 : '' Calathus pirazzolii'' Putzeys, 1873 : '' Calathus ravasinii'' G.Müller, 1935 : '' Calathus reflexicollis'' Faldermann, 1839 : '' Calathus reflexus'' Schaum, 1858 : '' Calathus roccai'' F.Battoni, 1984 : '' Calathus rubripes'' Dejean, 1831 : '' Calathus sirentensis'' D'Amore Fracassi, 1908 : '' Calathus syriacus'' Chaudoir, 1863 : '' Calathus uniseriatus'' Vuillefroy, 1866 : '' Calathus vignatagliantii'' F.Battoni, 1986 : '' Calathus vivesi'' Nègre, 1966 : '' Calathus vuillefroyi'' Gautier des Cottes, 1867 : '' Calathus zabroides'' Putzeys, 1873 ; Subgenus Certocalathus J.Schmidt & Will, 2020 : '' Calathus advena'' (LeConte, 1846) ; Subgenus Denticalathus J.Schmidt, 2018 : '' Calathus aethiopicus'' Alluaud, 1918 : '' Calathus aethiops'' Alluaud, 1937 : '' Calathus afroalpinus'' Lassalle, 2016 : '' Calathus ankoberensis'' Lassalle, 2016 : '' Calathus baleensis'' Lassalle, 2016 : '' Calathus balli'' Novoa; Gañan & Baselga, 2015 : '' Calathus carballalae'' Novoa & Gañán, 2014 : '' Calathus chioriae'' Lassalle, 2016 : '' Calathus chokeensis'' Lassalle, 2016 : '' Calathus dubaulti'' Lassalle, 2016 : '' Calathus gugheensis'' Basilewsky, 1953 : '' Calathus gunaensis'' Lassalle, 2016 : '' Calathus juan'' Novoa & Gañán, 2014 : '' Calathus kebedei'' Novoa; Gañan & Baselga, 2015 : '' Calathus montanus'' Alluaud, 1937 : '' Calathus nitidus'' Basilewsky, 1957 : '' Calathus ollivieri'' Lassalle, 2016 : '' Calathus oreobius'' Alluaud, 1937 : '' Calathus orthomoides'' Alluaud, 1932 : '' Calathus parvicollis'' Fairmaire, 1882 : '' Calathus patrizii'' Basilewsky, 1953 : '' Calathus queinneci'' Lassalle, 2016 : '' Calathus ras'' Basilewsky, 1957 : '' Calathus reebae'' Lassalle, 2016 : '' Calathus scotti'' Alluaud, 1937 : '' Calathus scottianus'' Basilewsky, 1953 : '' Calathus shoanus'' Alluaud, 1932 : '' Calathus simienensis'' Basilewsky, 1957 : '' Calathus theodori'' Ancey, 1882 : '' Calathus trapezicollis'' Alluaud, 1937 : '' Calathus vagestriatus'' Fairmaire, 1882 : '' Calathus wolloensis'' Lassalle, 2016 ; Subgenus Iberocalathus Toribio, 2006 : '' Calathus rotundatus'' Jacquelin du Val, 1855 ; Subgenus Indocalathus J.Schmidt, 2018 : '' Calathus kirschenhoferi'' F.Battoni, 1982 ; Subgenus Lauricalathus Machado, 1992 : '' Calathus abaxoides'' Brullé, 1839 : '' Calathus amplius'' Escalera, 1921 : '' Calathus angularis'' Brullé, 1839 : '' Calathus angustulus'' Wollaston, 1862 : '' Calathus appendiculatus'' Wollaston, 1862 : '' Calathus ascendens'' Wollaston, 1862 : '' Calathus auctus'' Wollaston, 1862 : '' Calathus canariensis'' Harold, 1868 : '' Calathus carinatus'' Brullé, 1839 : '' Calathus ciliatus'' Wollaston, 1862 : '' Calathus cognatus'' Wollaston, 1864 : '' Calathus depressus'' Brullé, 1836 : '' Calathus freyi'' Colas, 1941 : '' Calathus gomerensis'' Colas, 1943 : '' Calathus laureticola'' Wollaston, 1865 : '' Calathus marcellae'' Colas, 1943 : '' Calathus rectus'' Wollaston, 1862 : '' Calathus rufocastaneus'' Wollaston, 1862 : '' Calathus spretus'' Wollaston, 1862 ; Subgenus Lindrothius Kurnakov, 1961 : '' Calathus aeneocupreus'' (Heinz, 1971) : '' Calathus aequistriatus'' (Kurnakov, 1961) : '' Calathus caucasicus'' Chaudoir, 1846 : '' Calathus grandiceps'' (Kurnakov, 1961) : '' Calathus horsti'' (Reitter, 1888) : '' Calathus laticaudis'' (Kurnakov, 1961) : '' Calathus mandibularis'' (Kurnakov, 1961) : '' Calathus praestans'' (Heyden, 1885) : '' Calathus pseudopraestans'' (Kurnakov, 1961) : '' Calathus recticaudis'' (Kurnakov, 1961) : '' Calathus robustus'' (Kurnakov, 1961) : '' Calathus sotshiensis'' (Zamotajlov, 1997) : '' Calathus stricticaudis'' (Kurnakov, 1961) : '' Calathus subpraestans'' (Kurnakov, 1961) ; Subgenus Neocalathus Ball & Nègre, 1972 : '' Calathus albanicus'' Apfelbeck, 1906 : '' Calathus ambigens'' Bates, 1891 : '' Calathus ambiguus'' (Paykull, 1790) : '' Calathus anistschenkoi'' Berlov & Ippolitova, 2005 : '' Calathus asturiensis'' Vuillefroy, 1866 : '' Calathus aztec'' Ball & Nègre, 1972 : '' Calathus bolivari'' Nègre, 1968 : '' Calathus calceus'' Ball & Nègre, 1972 : '' Calathus cinctus'' Motschulsky, 1850 : '' Calathus clauseni'' Ball & Nègre, 1972 : '' Calathus deplanatus'' Chaudoir, 1843 : '' Calathus deyrollei'' Gautier des Cottes, 1870 : '' Calathus durango'' Ball & Nègre, 1972 : '' Calathus erratus'' (C.R.Sahlberg, 1827) : '' Calathus erwini'' Ball & Nègre, 1972 : '' Calathus extensicollis'' Putzeys, 1873 : '' Calathus femoralis'' Chaudoir, 1846 : '' Calathus gonzalezi'' Mateu, 1956 : '' Calathus gregarius'' (Say, 1823) : '' Calathus hyrcanus'' Heinz, 1970 : '' Calathus ingratus'' Dejean, 1828 : '' Calathus kollari'' Putzeys, 1873 : '' Calathus leechi'' Ball & Nègre, 1972 : '' Calathus leptodactylus'' Putzeys, 1873 : '' Calathus lundbladi'' Colas, 1938 : '' Calathus marmoreus'' Ball & Nègre, 1972 : '' Calathus melanocephalus'' (Linnaeus, 1758) : '' Calathus metallicus'' Dejean, 1828 : '' Calathus mexicanus'' Chaudoir, 1837 : '' Calathus micropterus'' (Duftschmid, 1812) : '' Calathus mollis'' (Marsham, 1802) : '' Calathus opaculus'' LeConte, 1854 : '' Calathus peltatus'' Kolenati, 1845 : '' Calathus peropacus'' Casey, 1920 : '' Calathus pommeranzi'' J.Schmidt, 1999 : '' Calathus potosi'' Ball & Nègre, 1972 : '' Calathus relictus'' J.Schmidt & Tian, 2013 : '' Calathus rhaticus'' Antoine, 1941 : '' Calathus rotgeri'' Ball & Nègre, 1972 : '' Calathus ruficollis'' Dejean, 1828 : '' Calathus semisericeus'' Fairmaire, 1879 : '' Calathus simplicicollis'' Wollaston, 1862 : '' Calathus solieri'' Bassi, 1834 : '' Calathus subfuscus'' Wollaston, 1865 : '' Calathus vicenteorum'' Schatzmayr, 1937 : '' Calathus whiteheadi'' Ball & Nègre, 1972 : '' Calathus wittmerianus'' Deuve; Lassalle & Queinnec, 1985 : †'' Calathus elpis'' Ortuño & Arillo, 2009 ; Subgenus Spinocalathus J.Schmidt, 2018 : '' Calathus annapurnae'' J.Schmidt, 1999 : '' Calathus deliae'' Morvan, 1999 : '' Calathus fabigi'' J.Schmidt, 2018 : '' Calathus ganeshi'' J.Schmidt, 2018 : '' Calathus gorkhaorum'' J.Schmidt, 2018 : '' Calathus hastatus'' J.Schmidt, 2018 : '' Calathus heinertzi'' Deuve; Lassalle & Queinnec, 1985 : '' Calathus holzschuhi'' Kirschenhofer, 1990 : '' Calathus kryzhanovskii'' J.Schmidt, 1999 : '' Calathus manasluensis'' J.Schmidt, 1999 : '' Calathus shikaensis'' J.Schmidt, 2018 : '' Calathus vulneratus'' J.Schmidt, 2018 ; Subgenus Synuchidius Apfelbeck, 1908 : '' Calathus ganglbaueri'' (Apfelbeck, 1908) ; Subgenus Tachalus Ball & Nègre, 1972 : '' Calathus ovipennis'' Putzeys, 1873 ; Subgenus Trichocalathus Bolivar y Pieltain, 1940 : '' Calathus obliteratus'' Wollaston, 1865 : '' Calathus pilosipennis'' Machado, 1992 : '' Calathus refleximargo'' Machado, 1992 ; Subgenus unassigned : '' Calathus algens'' Andrewes, 1934 : '' Calathus anatolicus'' Jedlicka, 1969 : '' Calathus carvalhoi'' Serrano & Borges, 1986 : '' Calathus colasianus'' Mateu, 1970 : '' Calathus complanatus'' Dejean, 1828 : '' Calathus fimbriatus'' Wollaston, 1858 : '' Calathus gelascens'' Andrewes, 1934 : '' Calathus himalayae'' Bates, 1891 : '' Calathus montanellus'' Heinz, 2000 : '' Calathus montivagus'' Dejean, 1831 : '' Calathus pecoudi'' Colas, 1938 : '' Calathus suffuscus'' Andrewes, 1934 : '' Calathus tombesii'' F.Battoni, 1976 : '' Calathus vividus'' (Fabricius, 1801)

References

External links

*
''Calathus'' at Fauna Europaea
* * Platyninae Palearctic insects Taxa named by Franco Andrea Bonelli Carabidae genera Long stubs with short prose {{Platyninae-stub