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The Calabar angwantibo (''Arctocebus calabarensis''), also known as the Calabar potto, is a
strepsirrhine Strepsirrhini or Strepsirhini (; ) is a Order (biology), suborder of primates that includes the Lemuriformes, lemuriform primates, which consist of the lemurs of Fauna of Madagascar, Madagascar, galagos ("bushbabies") and pottos from Fauna of A ...
primate Primates are a diverse order (biology), order of mammals. They are divided into the Strepsirrhini, strepsirrhines, which include the lemurs, galagos, and lorisids, and the Haplorhini, haplorhines, which include the Tarsiiformes, tarsiers and ...
of the family
Lorisidae Lorisidae (or sometimes Loridae) is a family of strepsirrhine primates. The lorisids are all slim arboreal animals and comprise the lorises, pottos and angwantibos. Lorisids live in tropical, central Africa as well as in south and southeast A ...
. It shares the genus '' Arctocebus'' with the golden angwantibo (''Arctocebus aureus''). It is closely related to the
potto The pottos are three species of strepsirrhine primate in the genus ''Perodicticus'' of the family Lorisidae. In some English-speaking parts of Africa, they are called "softly-softlys". Etymology The common name "potto" may be from Wolof (a t ...
(''Perodicticus potto'') and to the various
loris Loris is the common name for the strepsirrhine mammals of the subfamily Lorinae (sometimes spelled Lorisinae) in the family Lorisidae. ''Loris'' is one genus in this subfamily and includes the slender lorises, ''Nycticebus'' is the genus con ...
es. The Calabar angwantibo lives in the
rainforest Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainfo ...
s of west Africa, particularly in tree-fall zones. In areas where the forest has been cleared, it has been known to live on farmland. Its range covers
Cameroon Cameroon (; french: Cameroun, ff, Kamerun), officially the Republic of Cameroon (french: République du Cameroun, links=no), is a country in west- central Africa. It is bordered by Nigeria to the west and north; Chad to the northeast; th ...
,
Nigeria Nigeria ( ), , ig, Naìjíríyà, yo, Nàìjíríà, pcm, Naijá , ff, Naajeeriya, kcg, Naijeriya officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf of G ...
and Equatorial Guinea. The species takes its name from the Nigerian city of
Calabar Calabar (also referred to as Callabar, Calabari, Calbari and Kalabar) is the capital city of Cross River State, Nigeria. It was originally named Akwa Akpa, in the Efik language. The city is adjacent to the Calabar and Great Kwa rivers and ...
. The Calabar angwantibo weighs between 266 and 465 grams. It has orangish-yellow fur on its back, grey or white fur on its belly, and a distinctive white line on its forehead and nose. Like other lorids, this
angwantibo Angwantibos are two species of strepsirrhine primates classified in the genus ''Arctocebus'' of the family Lorisidae. They are also known as golden pottos because of their yellow or golden coloration. Angwantibos live in tropical Africa and thei ...
has a very short index finger, which allows it to get a strong grip on tree branches. The second toe on each foot has a specialised claw that the angwantibo uses for grooming. The Calabar angwantibo is the only primate to have a functioning
nictitating membrane The nictitating membrane (from Latin '' nictare'', to blink) is a transparent or translucent third eyelid present in some animals that can be drawn across the eye from the medial canthus to protect and moisten it while maintaining vision. All An ...
(third eyelid).Montagna, W., Machida, H., and Perkins, E. M. 1966. "The skin of primates XXXIII.: The skin of the angwantibo". ''American Journal of Physical Anthropology''. Vol. 25, 277-290. The Calabar angwantibo is
nocturnal Nocturnality is an ethology, animal behavior characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnality, diurnal meaning the opposite. Nocturnal creatures generally have ...
and
arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally, but others are exclusively arboreal. The habitats pose num ...
. It stays considerably lower in the trees than the other nocturnal strepsirrhines in its range, and is typically found between 5 and 15 metres above ground. It moves by climbing very slowly through the trees, always grasping branches with at least three of its limbs at a time. During the day the angwantibo sleeps under dense foliage, hanging from a branch. The Calabar angwantibo's diet consists mainly of insects, especially
caterpillar Caterpillars ( ) are the larva, larval stage of members of the order Lepidoptera (the insect order comprising butterfly, butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawfly ...
s, but it also eats some fruit. It will eat strong-smelling insects that other animals reject. Before eating a caterpillar, the angwantibo wipes it carefully with its hands to remove any poisonous barbs. When confronted by a
predator Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill ...
, the Calabar angwantibo will roll up into a ball, but keep its mouth open beneath its armpit. If the attacker persists, the angwantibo will bite it and not let go. Calabar angwantibos forage for food alone, but each male's territory overlaps that of several females. Angwantibos reinforce social bonds through
mutual grooming Social grooming is a behavior in which social animals, including humans, clean or maintain one another's body or appearance. A related term, allogrooming, indicates social grooming between members of the same species. Grooming is a major socia ...
and scent-marking. Mating takes place only in the final phase of the female's
estrous cycle The estrous cycle (, originally ) is the set of recurring physiological changes that are induced by reproductive hormones in most mammalian therian females. Estrous cycles start after sexual maturity in females and are interrupted by anestr ...
, and is performed hanging upside-down from a branch. The female gives birth to a single infant after a gestation period of 131 to 136 days; the young are normally born between January and April. Infants are born with their eyes open and can cling to their mother's fur right away.


In literature

The Calabar angwantibo, along with its close relative the potto, makes an appearance in
Patrick O'Brian Patrick O'Brian, CBE (12 December 1914 – 2 January 2000), born Richard Patrick Russ, was an English novelist and translator, best known for his Aubrey–Maturin series of sea novels set in the Royal Navy during the Napoleonic Wars, and cen ...
's Aubrey–Maturin novels. Stephen Maturin acquires a Calabar angwantibo on his travels and becomes 'absurdly attached' to it. The search for an angwantibo is also a minor focus of
Gerald Durrell Gerald Malcolm Durrell, (7 January 1925 – 30 January 1995) was a British naturalist, writer, zookeeper, conservation movement, conservationist, and television presenter. He founded the Durrell Wildlife Conservation Trust and the Jersey Zoo o ...
's first book, '' The Overloaded Ark''.


References


External links


Calabar potto (''Arctocebus calabarensis'')
at The Primata site {{Taxonbar, from=Q1153493 Calabar angwantibo Mammals of Cameroon Mammals of West Africa Primates of Africa Calabar angwantibo