
A caiman ( (also spelled cayman) from
Taíno
The Taíno are the Indigenous peoples of the Caribbean, Indigenous peoples of the Greater Antilles and surrounding islands. At the time of European contact in the late 15th century, they were the principal inhabitants of most of what is now The ...
''kaiman'') is an
alligatorid belonging to the
subfamily
In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
Caimaninae, one of two primary lineages within the
Alligatoridae
The family (biology), family Alligatoridae of crocodylians includes alligators, caimans and their extinct relatives.
Phylogeny
The superfamily Alligatoroidea includes all crocodilians (fossil and extant) that are more closely related to the Am ...
family
Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
, the other being
alligator
An alligator, or colloquially gator, is a large reptile in the genus ''Alligator'' of the Family (biology), family Alligatoridae in the Order (biology), order Crocodilia. The two Extant taxon, extant species are the American alligator (''A. mis ...
s. Caimans are native to
Central and
South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
and inhabit
marshes,
swamp
A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
s,
lake
A lake is often a naturally occurring, relatively large and fixed body of water on or near the Earth's surface. It is localized in a basin or interconnected basins surrounded by dry land. Lakes lie completely on land and are separate from ...
s, and
mangrove
A mangrove is a shrub or tree that grows mainly in coastal saline water, saline or brackish water. Mangroves grow in an equatorial climate, typically along coastlines and tidal rivers. They have particular adaptations to take in extra oxygen a ...
rivers. They have scaly skin and live a fairly nocturnal existence. They are relatively small-sized
crocodilia
Crocodilia () is an order of semiaquatic, predatory reptiles that are known as crocodilians. They first appeared during the Late Cretaceous and are the closest living relatives of birds. Crocodilians are a type of crocodylomorph pseudosuchia ...
ns with an average maximum weight of depending on species, with the exception of the
black caiman
The black caiman (''Melanosuchus niger'') is a crocodilian reptile endemic to South America. With a maximum length of around and a mass of over , it is the largest living species of the family Alligatoridae, and the third-largest crocodilian in ...
(''Melanosuchus niger''), which can grow more than in length and weigh in excess of 450 kg (1,000 Ib). The black caiman is the largest caiman species in the world and is found in the slow-moving rivers and lakes that surround the Amazon basin. The smallest species is the
Cuvier's dwarf caiman (''Paleosuchus palpebrosus''), which grows to long. There are six different species of caiman found throughout the watery jungle habitats of Central and Southern America. The average length for most of the other caiman species is about long.
Caimans are distinguished from alligators, their closest relatives, by a few defining features: a lack of a bony septum between the nostrils, ventral armor composed of overlapping bony scutes formed from two parts united by a suture, and longer and sharper teeth than alligators, plus caimans tend to be more agile and crocodile-like in their movements.
The calcium rivets on caiman scales make their hides stiffer.
Several extinct forms are known, including ''
Purussaurus'', a giant
Miocene
The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
genus that grew to
and the ''
Mourasuchus'', which had a wide
duck
Duck is the common name for numerous species of waterfowl in the family (biology), family Anatidae. Ducks are generally smaller and shorter-necked than swans and goose, geese, which are members of the same family. Divided among several subfam ...
-like snout.
Behavior
Caimans are predators and, like alligators and crocodiles, their diet largely consists of fish. Caimans also hunt insects, birds, small mammals and reptiles.
Due to their large size and ferocious nature, caimans have few natural predators within their environments. Humans are their main predators, because the animals have been hunted for their meat and skin.
Jaguar
The jaguar (''Panthera onca'') is a large felidae, cat species and the only extant taxon, living member of the genus ''Panthera'' that is native to the Americas. With a body length of up to and a weight of up to , it is the biggest cat spe ...
s,
anacondas and
crocodile
Crocodiles (family (biology), family Crocodylidae) or true crocodiles are large, semiaquatic reptiles that live throughout the tropics in Africa, Asia, the Americas and Australia. The term "crocodile" is sometimes used more loosely to include ...
s are the only other predators of caimans, although they usually prey on the smaller specimens or specific species of caiman such as the
Spectacled Caiman and
Yacare caiman. During summer or droughts, caimans may dig a burrow and go into a form of summer hibernation called
aestivation
Aestivation ( (summer); also spelled estivation in American English) is a state of animal dormancy, similar to hibernation, although taking place in the summer rather than the winter. Aestivation is characterized by inactivity and a lowered m ...
.
Female caimans build a large nest in which to lay their eggs. The nests can be more than wide. Female caimans lay between 10 and 50 eggs, which hatch within about six weeks. Once they have hatched, the mother caiman takes her young to a shallow pool of water, where they can learn how to hunt and swim. The juveniles of spectacled caiman have been shown to stay together in pods for up to 18 months.
Phylogeny
Caimaninae is
cladistically defined as ''Caiman crocodylus'' (the
spectacled caiman) and all species closer to it than to ''Alligator mississippiensis'' (the
American alligator).
This is a
stem-based definition for caimaninae, and means that it includes more
basal extinct
Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
caimanine ancestors that are more closely related to living caimans than to
alligators. The clade
Jacarea includes the most derived caimans, being defined as the last common ancestor of ''
Caiman latirostris'' (Broad-snouted caiman), ''
Caiman crocodilus'' (Spectacled caiman), ''
Caiman yacare'' (Yacare caiman), ''
Melanosuchus niger'' (Black caiman), and all its descendants.
Below is a
cladogram
A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
showing the
phylogeny
A phylogenetic tree or phylogeny is a graphical representation which shows the evolutionary history between a set of species or Taxon, taxa during a specific time.Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, M ...
of Caimaninae, modified from Hastings ''et al.'' (2013).
Here is an alternative cladogram from Bona ''et al.'' 2018.
The
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
taxa ''
Stangerochampsa'', ''
Brachychampsa'' and ''
Albertochampsa'' have been previously referred to as stem-group caimans,
but Walter et al. (2022) recovered them as the basalmost
alligatorines based on phylogenetic analysis and claimed that the earliest definitive stem-group caimans are known from the
earliest Paleocene.
A different study by Adam Cossette and David Tarailo in 2024 recovered ''Brachychampsa'' and relatives in a clade at the base of Caimaninae. They named this clade Brachychampsini, defining it as "the largest clade of alligatorids more closely related to ''
Brachychampsa montana'' than to ''
Caiman crocodilus'' or ''
Alligator mississippiensis''".
See also
*
Chinese alligator
The Chinese alligator (; ), also known as the Yangtze alligator (), China alligator, or historically the muddy dragon, is a crocodilian endemic to China. It and the American alligator (''A. mississippiensis'') are the only living species in th ...
References
{{Taxonbar, from=Q478759
Alligatoridae
Selandian first appearances
Extant Selandian first appearances
mk:Кајмани