Byttneria Irwinii
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''Ayenia'' is a genus of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
s in the mallow family,
Malvaceae Malvaceae (), or the mallows, is a family of flowering plants estimated to contain 244 genera with 4225 known species. Well-known members of economic importance include Theobroma cacao, cacao, Cola (plant), cola, cotton, okra, Hibiscus sabdariffa, ...
. It includes 216 species of subshrubs, shrubs, small trees, and lianas.Whitlock, B. A., & Hale, A. M. (2011)
The phylogeny of ''Ayenia'', ''Byttneria'', and ''Rayleya'' (Malvaceae ''s.l.'') and its implications for the evolution of growth forms.
''Systematic Botany'', 36(1), 129-136.
They are native to the tropical
Americas The Americas, sometimes collectively called America, are a landmass comprising the totality of North America and South America.''Webster's New World College Dictionary'', 2010 by Wiley Publishing, Inc., Cleveland, Ohio. When viewed as a sing ...
and southwestern United States, tropical Africa, and tropical Asia.


Description

''Ayenia'' includes subshrubs, shrubs, small trees, and lianas. The genus is distinguished by its tiny yet extraordinarily complex flowers.


Ecology

Species of ''Ayenia'' grow in diverse habitats, from open areas in dry and seasonally-dry regions to humid forests, river banks, and from lowlands to high elevations. ''Byttneria'' species are host plants to insects such as beetles of the genus '' Lonchophorellus''.


Taxonomy

''Ayenia'', ''Byttneria'', and ''Rayleya'' were formerly described as separate genera, with ''Ayenia'' having a Neotropical distribution, ''Byttneria'' as a
Pantropical A pantropical ("all tropics") distribution is one which covers tropical regions of both the Eastern and Western hemispheres. Examples of species include caecilians, modern sirenians and the plant genera ''Acacia'' and ''Bacopa''. ''Neotropical' ...
genus, and ''Rayleya'' as a monotypic genus with a single Brazilian species, ''Rayleya bahiensis''. A phylogenetic analysis found that ''Byttneria'' was
paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...
and ''Ayenia'' was nested within it, and ''Rayleya'' was a sister clade to the others. In 2018 Christenhusz and Byng transferred the ''Byttneria'' and ''Rayleya'' names to ''Ayenia'', which had nomenclatural priority.Dorr, L.J. 2023. New combinations in Ayenia (Malvaceae, Byttnerioideae) for the flora of Ecuador. ''Phytoneuron'' 2023-04: 1– 5. Published 17 February 2023. ISSN 2153-733X


Species

217 species are currently accepted. Selected species include: *'' Ayenia abutilifolia'' *'' Ayenia abutiloides'' *'' Ayenia acalyphifolia'' *'' Ayenia aculeata'' *'' Ayenia acuminata'' *'' Ayenia aemulata'' *'' Ayenia affinis'' *'' Ayenia albiflora'' *'' Ayenia aliculata'' *'' Ayenia ambongensis'' *'' Ayenia ancistrodonta'' *'' Ayenia andamensis'' *'' Ayenia angulata'' *'' Ayenia angustifolia'' *'' Ayenia aprica'' *'' Ayenia ardua''
Cristóbal Cristóbal or Cristobal, the Spanish version of Christopher, is a masculine given name and a surname which may refer to: Given name *Cristóbal Balenciaga (1895–1972), Spanish fashion designer *Cristóbal Cobo (born 1976), Chilean academic * Cr ...
– Wingstem ayenia *'' Ayenia arenesii'' *'' Ayenia aristeguietae'' *'' Ayenia asplundii'' – Ecuador and Peru *'' Ayenia asterotricha'' *''
Ayenia atrata ''Ayenia'' is a genus of flowering plants in the mallow family, Malvaceae. It includes 216 species of subshrubs, shrubs, small trees, and lianas.Whitlock, B. A., & Hale, A. M. (2011)The phylogeny of ''Ayenia'', ''Byttneria'', and ''Rayleya'' (Mal ...
'' *'' Ayenia attenuatifolia'' *'' Ayenia aurantiaca'' *'' Ayenia australis'' *'' Ayenia bahiensis'' *'' Ayenia baronii'' *'' Ayenia beccarii'' *'' Ayenia benensis'' *'' Ayenia berlandieri'' *'' Ayenia bernieri'' *'' Ayenia besalampensis'' *'' Ayenia beyrichiana'' *'' Ayenia biloba'' *'' Ayenia blanchetiana'' *'' Ayenia boliviana'' *'' Ayenia byttnerioides'' *'' Ayenia cajalbanensis'' *'' Ayenia campicola'' *'' Ayenia capillata'' *'' Ayenia caripensis'' *'' Ayenia catalpifolia'' *'' Ayenia celebica'' *'' Ayenia celtoides'' *'' Ayenia compacta''
Rose A rose is either a woody perennial plant, perennial flowering plant of the genus ''Rosa'' (), in the family Rosaceae (), or the flower it bears. There are over three hundred Rose species, species and Garden roses, tens of thousands of cultivar ...
– California ayenia *'' Ayenia conciliata'' *'' Ayenia corchorifolia'' *'' Ayenia cordata'' *'' Ayenia cordobensis'' *'' Ayenia coriacea'' *'' Ayenia corylifolia'' *'' Ayenia crenulata'' *'' Ayenia cristobaliana'' *'' Ayenia crotonoides'' *'' Ayenia cuatrecasasii'' *'' Ayenia cubensis'' *'' Ayenia curtisii'' *'' Ayenia dahomensis'' *'' Ayenia decaryana'' *'' Ayenia dentata'' *'' Ayenia divaricata'' *'' Ayenia donatica'' *'' Ayenia ekmanii'' *'' Ayenia elegans'' *'' Ayenia eliae'' *'' Ayenia elliptica'' *'' Ayenia ellipticifolia'' *'' Ayenia erecta'' *'' Ayenia erosa'' *'' Ayenia euphrasiifolia'' Griseb. – Eyebright ayenia *'' Ayenia fasiculata'' *'' Ayenia fernandesii'' *'' Ayenia filiformis'' S.Watson – Trans-Pecos ayenia *'' Ayenia filipes'' *'' Ayenia flaccida'' *'' Ayenia flexuosa'' – Ecuador *'' Ayenia fluvialis'' *'' Ayenia fontis'' *'' Ayenia fruticosa'' *'' Ayenia fulva'' *'' Ayenia gayana'' *'' Ayenia geminifolia'' *'' Ayenia genistella'' *'' Ayenia glabra'' S.Watson – Smooth ayenia *'' Ayenia glabrescens'' *'' Ayenia glazioui'' *'' Ayenia gracilipes'' *'' Ayenia grandifolia'' *'' Ayenia grisea'' *'' Ayenia guineensis'' *'' Ayenia hatschbachii'' *'' Ayenia herbacea'' *'' Ayenia heteromorpha'' *'' Ayenia heterophylla'' *'' Ayenia hirsuta'' *'' Ayenia hirta'' *'' Ayenia humbertiana'' *'' Ayenia idroboi'' *'' Ayenia implicabilis'' *'' Ayenia incallida'' *'' Ayenia insulicola'' Cristóbal – Dwarf ayenia *'' Ayenia integrifolia'' *'' Ayenia irwinii'' *'' Ayenia ivorensis'' *'' Ayenia jackiana'' *'' Ayenia jaculifolia'' *'' Ayenia jaliscana'' *'' Ayenia jaramilloana'' *'' Ayenia jussieui'' *'' Ayenia juzepczukii'' *'' Ayenia klugii'' *'' Ayenia krapovickasii'' *'' Ayenia laevigata'' *'' Ayenia lancifolia'' *'' Ayenia lasiophylla'' *'' Ayenia latifolia'' *'' Ayenia latipetala'' *'' Ayenia limitaris'' Cristóbal – Rio Grande ayenia, Texas ayenia *'' Ayenia lingulata'' *'' Ayenia lobata'' *'' Ayenia lopez-mirandae'' *'' Ayenia loxensis'' *'' Ayenia lucida'' *'' Ayenia luyensis'' *'' Ayenia macrantha'' *'' Ayenia macrophylla'' *'' Ayenia magna'' *'' Ayenia maingayi'' *'' Ayenia mansfeldiana'' *'' Ayenia mastatalensis'' *'' Ayenia mastersii'' *'' Ayenia melantha'' *'' Ayenia melastomifolia'' *'' Ayenia melleri'' *'' Ayenia micrantha'' *'' Ayenia microphylla'' A.Gray – Dense ayenia, shrubby ayenia *'' Ayenia minytricha'' *'' Ayenia mirandae'' *'' Ayenia morifolia'' *'' Ayenia morii'' *'' Ayenia nervosa'' *'' Ayenia nitidula'' *'' Ayenia noblickii'' *'' Ayenia nossibeensis'' *'' Ayenia nummularia'' *'' Ayenia obcordata'' *'' Ayenia obliqua'' *'' Ayenia oblongata'' *'' Ayenia oblongifolia'' *'' Ayenia obtusata'' *'' Ayenia odonellii'' *'' Ayenia oligantha'' *'' Ayenia oranensis'' *'' Ayenia ostenii'' *'' Ayenia ovata'' *'' Ayenia ovatifolia'' *'' Ayenia palmeri'' *'' Ayenia palustris'' *'' Ayenia paniculata'' *'' Ayenia parviflora'' *'' Ayenia pedersenii'' *'' Ayenia peninsularis'' *'' Ayenia peregrina'' *'' Ayenia perrieri'' *'' Ayenia pescapraeifolia'' *'' Ayenia petiolata'' *'' Ayenia pilosa'' Cristóbal – Hairy ayenia *'' Ayenia piresii'' *'' Ayenia praecipua'' *'' Ayenia praeclara'' *'' Ayenia purpusii'' *'' Ayenia pusilla'' *'' Ayenia ramosissima'' *'' Ayenia reinwardtii'' *'' Ayenia rhamnifolia'' *'' Ayenia rojasii'' *'' Ayenia rotundifolia'' *'' Ayenia rubriflora'' *'' Ayenia sagasteguii'' *'' Ayenia sagittifolia'' *'' Ayenia saligna'' *'' Ayenia sambiranensis'' *'' Ayenia scabra'' *'' Ayenia scabrida'' *'' Ayenia scalpellata'' *'' Ayenia schumanniana'' *'' Ayenia schumannii'' *'' Ayenia schunkei'' *'' Ayenia scorpiura'' *'' Ayenia sidifolia'' *'' Ayenia simulatrix'' *'' Ayenia sparrei'' *'' Ayenia spinosa'' *'' Ayenia spinulosa'' *'' Ayenia standleyi'' *'' Ayenia stenophylla'' *'' Ayenia stipularis'' *'' Ayenia subsessilis'' *'' Ayenia subtilis'' *'' Ayenia tenuicaulis'' *'' Ayenia tomentosa'' *'' Ayenia tortilis'' *'' Ayenia triadenia'' *'' Ayenia truncata'' *'' Ayenia tucumanensis'' *'' Ayenia uaupensis'' *'' Ayenia urosepala'' *'' Ayenia urticifolia'' *'' Ayenia vargasii'' *'' Ayenia velutina'' *'' Ayenia violacea'' *'' Ayenia virgata'' *'' Ayenia vitifolia'' *'' Ayenia voulily'' *'' Ayenia weberbaueri'' *'' Ayenia wrightii'' *'' Ayenia wygodzinskyi'' *'' Ayenia xochipilliae''


References


External links

* *
Jepson Manual TreatmentUSDA Plants Profile
{{Authority control Byttnerioideae Malvaceae genera Flora of the Afrotropical realm Flora of the Indomalayan realm Flora of the Neotropical realm Plants described in 1756 Taxa named by Carl Linnaeus