Buxolestes
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''Buxolestes'' is an extinct
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of semi-aquatic, non-
placental Placental mammals (infraclass Placentalia ) are one of the three extant subdivisions of the class Mammalia, the other two being Monotremata and Marsupialia. Placentalia contains the vast majority of extant mammals, which are partly distinguished ...
eutherian Eutheria (from Greek , 'good, right' and , 'beast'; ), also called Pan-Placentalia, is the clade consisting of placental mammals and all therian mammals that are more closely related to placentals than to marsupials. Eutherians are distingu ...
mammals A mammal () is a vertebrate animal of the class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three middle e ...
belonging to the family
Pantolestidae Pantolestidae is an extinct family of semi-aquatic, non-placental eutherian mammals. Forming the core of the equally extinct suborder Pantolesta, the pantolestids evolved as a series of increasingly otter-like forms, ranging from the Middle Pa ...
. Species in this genus were part of the first placental
evolutionary radiation An evolutionary radiation is an increase in taxonomic diversity that is caused by elevated rates of speciation, that may or may not be associated with an increase in morphological disparity. A significantly large and diverse radiation within ...
during the
Middle Eocene The Eocene ( ) is a geological epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''Ēṓs'', ' Dawn') a ...
(48-40 mya) and found in the
Bracklesham Group The Bracklesham Group (formerly Bracklesham Beds), in geology, is a series of clays and marls, with sandy and lignitic beds, in the middle Eocene of the Hampshire Basin and London Basin of England. The type section of the Bracklesham Group is th ...
and Wittering Formation of
England England is a Countries of the United Kingdom, country that is part of the United Kingdom. It is located on the island of Great Britain, of which it covers about 62%, and List of islands of England, more than 100 smaller adjacent islands. It ...
, at the
Messel Pit The Messel Formation is a Formation (geology), geologic formation in Hesse, central Germany, dating back to the Eocene Epoch (geology), epoch (about 47 Ma). Its geographic range is restricted to the Messel pit. There it unconformably overlie ...
in
Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
and in Bouxwiller, France.


Description

''Buxolestes'' were otter-like freshwater fish predators with a body length reaching about and a tail about long. They were significantly smaller than most living species of otters.
Fossilized A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
stomach contents confirm their semiaquatic freshwater habits. The anatomy of these archaic "
insectivorous A robber fly eating a hoverfly An insectivore is a carnivorous animal or plant which eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects. The first vertebrate insectivores we ...
" mammals is known through well-preserved Middle
Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
specimens found at
Messel Messel is a municipality in the district of Darmstadt-Dieburg in Hesse near Frankfurt am Main in Germany. The village is first mentioned, as ''Masilla'', in the Lorsch codex. Messel was the property of the lords of Groschlag from ca. 1400 to 1799 ...
in
Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
. Their structure evidences a clear adaptation to a semiaquatic way of life. The forelimbs and hindlimbs are powerful and show strong claws. The tail is clearly fit for swimming. The skull is long, with large molars that appear to be adapted to a diet of molluscs with shells (
freshwater clam Freshwater bivalves are molluscs of the order Bivalvia that inhabit freshwater ecosystems. They are one of the two main groups of freshwater molluscs, along with freshwater snails. The majority of bivalve molluscs are saltwater species that liv ...
s and
freshwater snail Freshwater snails are gastropod mollusks that live in fresh water. There are many different families. They are found throughout the world in various habitats, ranging from ephemeral pools to the largest lakes, and from small seeps and springs t ...
s), but the predilection postulated from the dentition has not been confirmed.


Species

* ''Buxolestes minor'' Pfretzschner, 1999 * ''Buxolestes piscator'' Koenigswald, 1980


References


External links


The Paleobiology DatabaseGlobal Names
{{Taxonbar, from=Q3647762 Cimolestans Lutetian genera Eocene mammals of Europe Paleogene England Fossils of England Eocene France Fossils of France Eocene Germany Fossils of Germany Fossil taxa described in 1970 Prehistoric mammal genera