Burchell's Courser
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Burchell's courser (''Cursorius rufus'') is a
wader 245px, A flock of Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflats in order to foraging, ...
in the pratincole and courser family,
Glareolidae Glareolidae is a family of birds in the wader suborder Lari. It contains two distinct groups, the pratincoles and the coursers. The atypical Egyptian plover (''Pluvianus aegyptius''), traditionally placed in this family, is now known to be o ...
. The name of this bird commemorates the English naturalist
William John Burchell __NOTOC__ William John Burchell (23 July 1781 – 23 March 1863) was an English explorer, naturalist, traveller, artist, and author. His thousands of plant specimens, as well as field journals from his South African expedition, are held by Kew ...
. Native to Africa, Burchell's courser is a small, diurnal, and terrestrial bird that lives in the western parts of southern
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
. Although classed as
wader 245px, A flock of Red_knot.html" ;"title="Dunlins and Red knot">Dunlins and Red knots Waders or shorebirds are birds of the order Charadriiformes commonly found wikt:wade#Etymology 1, wading along shorelines and mudflats in order to foraging, ...
s, these are
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s of dry open country, preferably semi-desert, where they typically hunt their
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
prey (usually
Harvester Termites The Hodotermitidae (from Greek ὁδός (hodós), travelling; Latin '' termes'', woodworm) are a basal Old World family of termites known as the harvester termites. They are distinguished by the serrated inner edge of their mandibles, and the ...
) by running on the ground. It principally feeds off of
insects Insects (from Latin ') are hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs of jointed ...
and lives in open, short
grasslands A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes such as clover, and other herbs. Grasslands occur ...
and burnt
veld Veld ( or , Afrikaans language, Afrikaans and Dutch language, Dutch: ''veld'', field), also spelled veldt, is a type of wide-open, rural landscape in Southern Africa. Particularly, it is a flat area covered in grass or low scrubland, scrub, ...
. It grows up to and on average weighs about as an adult.


Description

Burchell's courser has a graceful figure and an upright posture. It has a blue-grey hind crown which is mainly how it differs from the similar Temminck's courser. It also has a horizontal black band demarcating a white underbelly, and an overall pale
rufous Rufous () is a color that may be described as reddish-brown or brownish- red, as of rust or oxidised iron. The first recorded use of ''rufous'' as a color name in English was in 1782. However, the color is also recorded earlier in 1527 as a d ...
colour. Its face features white supercilia above black eye lines, all ending at the back of the neck to form a double V. In terms of feathers, its secondaries are mostly white with a black underwing and brown to grey
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are small feathers behind t ...
. Typical of coursers, its bill is long, arched down, and of a dark colour like its eyes. Similar to the underbelly, the legs are of a distinctive creamy white. While flying, the Burchell's courser's feet extend past its tail. Females and males look similar, while juveniles lack the rufous colouring, the facial stripes, and the grey hind crown of the adults. They also are mottled, with black and beige barring. Overall, the Burchell's courser, with its camouflage appearance, blends in with the landscape making it challenging to spot.


Taxonomy

Burchell's courser belongs to the order
Charadriiformes Charadriiformes (, from '' Charadrius'', the type genus of family Charadriidae) is a diverse order of small to medium-large birds. It includes about 390 species and has members in all parts of the world. Most charadriiform birds live near water ...
which mainly refers to birds living near water. Characterized by a curved beak and nostrils located at the base of the nose, the Burchell's courser is part of the
Glareolidae Glareolidae is a family of birds in the wader suborder Lari. It contains two distinct groups, the pratincoles and the coursers. The atypical Egyptian plover (''Pluvianus aegyptius''), traditionally placed in this family, is now known to be o ...
family, specifically belonging to the courser group. Its genus, ''
Cursorius '' Cursorius '' is a genus of coursers, a group of wading birds. The genus name derive from Latin ''cursor'' meaning "runner". There are five species which breed in Africa and South Asia. They have long legs, short wings and long pointed bill ...
,'' contains five species, all of which live in the arid parts of the
Old World The "Old World" () is a term for Afro-Eurasia coined by Europeans after 1493, when they became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia in the Eastern Hemisphere, previously ...
and display distinctive features such as long legs and short wings. Burchell's courser and its close relative, '' C. cursor'', have at times been referred to as the same species. However, although they have a direct common ancestor, they should not be considered as such. ''C. rufus'' is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
and does not have any subspecies.


Distribution and habitat

Endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to Africa, Burchell's courser prefers the warm and dry areas of
Southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed upon, but some groupings include the United Nations geoscheme for Africa, United Nations geoscheme, the intergovernmental Southern African Development Community, and ...
. Specifically, it is found in most of
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
and
Namibia Namibia, officially the Republic of Namibia, is a country on the west coast of Southern Africa. Its borders include the Atlantic Ocean to the west, Angola and Zambia to the north, Botswana to the east and South Africa to the south; in the no ...
, and in parts of
Angola Angola, officially the Republic of Angola, is a country on the west-Central Africa, central coast of Southern Africa. It is the second-largest Portuguese-speaking world, Portuguese-speaking (Lusophone) country in both total area and List of c ...
(Southwest) and
Botswana Botswana, officially the Republic of Botswana, is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory part of the Kalahari Desert. It is bordered by South Africa to the sou ...
( Kalahari Basin). Its distribution range occasionally coincides with that of the Temminck's courser. Burchell's courser inhabits
deserts A desert is a landscape where little precipitation occurs and, consequently, living conditions create unique biomes and ecosystems. The lack of vegetation exposes the unprotected surface of the ground to denudation. About one-third of the l ...
or semi deserts with gravel rather than just sand, but also open grasslands with short grass, and sparse, burnt, or overgrazed vegetation. It is often referred as locally nomad based on its movements at the landscape level, possibly travelling due to fluctuations in rainfall. Although it is classified as
least concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
, Its population is declining, potentially due to its habitat loss on arable lands, inadequate farming practices such as intense
irrigation Irrigation (also referred to as watering of plants) is the practice of applying controlled amounts of water to land to help grow crops, landscape plants, and lawns. Irrigation has been a key aspect of agriculture for over 5,000 years and has bee ...
and overuse of
pesticides Pesticides are substances that are used to pest control, control pest (organism), pests. They include herbicides, insecticides, nematicides, fungicides, and many others (see table). The most common of these are herbicides, which account for a ...
, and other anthropogenic disturbances.


Behaviour

The adults have developed a diversionary display as a strategy against predators. They perform "displacement
brooding Broodiness is the action or behavioral tendency to sit on a clutch of eggs to Egg incubation, incubate them, often requiring the non-expression of many other behaviors including feeding and drinking.Homedes Ranquini, J. y Haro-García, F. Zoogen ...
" to distract any potential threats from the nesting area. Even though the chicks are considered
precocial Precocial species in birds and mammals are those in which the young are relatively mature and mobile from the moment of birth or hatching. They are normally nidifugous, meaning that they leave the nest shortly after birth or hatching. Altricial ...
, they lack the heat-loss mechanisms present in the adults to survive the hot climate, and thus need shade to avoid overheating.


Voice

Burchell's courser often prefers to run rather than fly, and will most likely take to the air only when the disturbance is significant. Often referred to as inconspicuous, this bird is mostly quiet on the ground, but becomes vocal when flying, producing a loud flight call. Thus far, three types of vocalizations it produces have been identified: one throaty "chuk" as it launches in the air, a few "chuk chuk" grunts while flying, and a contact call consisting of a "kwirrt-kwirrt" or a "kok-kok-kwich". If threatened by a predator, the Burchell's courser may run away from its nest, emitting an
alarm call "Alarm Call" is a song recorded by Icelandic singer Björk for her third studio album ''Homogenic'' (1997). It was released as the fourth single from the album, peaking at number 33 in the United Kingdom. The sped-up radio edit of the song was us ...
in order to divert the predator.


Feeding

Burchell's courser is predominantly an
insectivore file:Common brown robberfly with prey.jpg, A Asilidae, robber fly eating a hoverfly An insectivore is a carnivore, carnivorous animal or plant which eats insects. An alternative term is entomophage, which can also refer to the Entomophagy ...
. It uses its long bill to forage the ground and dig up insects. This bird also runs to catch its prey. It has been found to frequently eat from the
Coleoptera Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
group, especially
curculionidae The Curculionidae are a family of weevils, commonly called snout beetles or true weevils. They are one of the largest animal families with 6,800 genera and 83,000 species described worldwide. They are the sister group to the family Brentidae. Th ...
beetles as its main food source. However, the Burchell's courser tends to have a preference for
harvester termites The Hodotermitidae (from Greek ὁδός (hodós), travelling; Latin '' termes'', woodworm) are a basal Old World family of termites known as the harvester termites. They are distinguished by the serrated inner edge of their mandibles, and the ...
(''Hodotermes'' ''mossambicus'') when available, which can constitute over 50% of its diet. It almost never chooses seeds as a source of nutrition. The yearly frequency of rainfalls affects the food availability and quality, which is the main reason the Burchell's courser travels around.


Breeding

Burchell's coursers may occur in small flocks but are mostly solitary,
monogamous Monogamy ( ) is a relationship of two individuals in which they form a mutual and exclusive intimate partnership. Having only one partner at any one time, whether for life or serial monogamy, contrasts with various forms of non-monogamy (e.g. ...
birds. They have developed an all-year-round breeding strategy due to their nomadic movements, although they often breed just before the
wet season The wet season (sometimes called the rainy season or monsoon season) is the time of year when most of a region's average annual rainfall occurs. Generally, the season lasts at least one month. The term ''green season'' is also sometimes used a ...
between July and December. With a small
clutch A clutch is a mechanical device that allows an output shaft to be disconnected from a rotating input shaft. The clutch's input shaft is typically attached to a motor, while the clutch's output shaft is connected to the mechanism that does th ...
size, the Burchell's courser only lays two eggs directly on the ground, sometimes surrounded by stools of antelopes, small rocks, or dried up
organic matter Organic matter, organic material or natural organic matter is the large source of carbon-based compounds found within natural and engineered, terrestrial, and aquatic environments. It is matter composed of organic compounds that have come fro ...
. The eggs have an oval shape, and look black from afar, but they are actually of a buff colour with many dark markings. There is a lot of uncertainty when it comes to the
hatching Hatching () is an artistic technique used to create tonal or shading effects by drawing (or painting or scribing) closely spaced parallel lines. When lines are placed at an angle to one another, it is called cross-hatching. Hatching is als ...
and fledging times, but it is believed that both parents participate in the incubation period.


References


External links


Burchell's courser
in The Atlas of Southern African Birds
Burchell's courser ''Cursorius rufus''
at The Cornell Lab of Ornithology

at VIREO * ttps://ebird.org/media/catalog?taxonCode=burcou2&sort=rating_rank_desc&mediaType=p®ionCode= Burchell's courser photo galleryat eBird {{Taxonbar, from=Q263815 Burchell's courser Birds of Southern Africa Burchell's courser Taxa named by John Gould