''Bunodes'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...

synziphosurine Synziphosurina is a paraphyletic group of chelicerate arthropods previously thought to be basal horseshoe crabs (Xiphosura). It was later identified as a grade composed of various basal euchelicerates, eventually excluded form the monophyletic Xi ...

, a

paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...

group of fossil

chelicerate The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, ticks, ...

arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arth ...

s. ''Bunodes'' was regarded as part of the clade Planaterga.

Fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

s of the single and

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...

, ''B. lunula'', have been discovered in

deposits A deposit account is a bank account maintained by a financial institution in which a customer can deposit and withdraw money. Deposit accounts can be savings accounts, current accounts or any of several other types of accounts explained below. ...

of the

Silurian The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago (Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoi ...

period in

Ludlow Ludlow () is a market town in Shropshire, England. The town is significant in the history of the Welsh Marches and in relation to Wales. It is located south of Shrewsbury and north of Hereford, on the A49 road (Great Britain), A49 road which ...

England England is a country that is part of the United Kingdom. It shares land borders with Wales to its west and Scotland to its north. The Irish Sea lies northwest and the Celtic Sea to the southwest. It is separated from continental Europe ...

(in the

United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the European mainland, continental mainland. It comprises England, Scotlan ...

). ''Bunodes'' is the type genus of the

family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...

Bunodidae, the other genera of the same family being '' Limuloides''. ''Bunodes'' is characterized by a vaulted

carapace A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the und ...

with radiated hump-like ridges. Within the 10-segmented

opisthosoma The opisthosoma is the posterior part of the body in some arthropods, behind the prosoma ( cephalothorax). It is a distinctive feature of the subphylum Chelicerata (arachnids, horseshoe crabs and others). Although it is similar in most respects ...

, the first opisthosomal

tergite A ''tergum'' (Latin for "the back"; plural ''terga'', associated adjective tergal) is the dorsal ('upper') portion of an arthropod segment other than the head. The anterior edge is called the 'base' and posterior edge is called the 'apex' or 'mar ...

is greatly reduced and always covered by the posterior region of preceding carapace, while the second tergite is significantly well-developed. the last 3 opisthosomal segments specialized into a narrow postabdomen and lacking tergopleurae (lateral extension of tergites). Tubercles of various sizes covering most of the dorsal surface of both carapace and tergites. Similar to '' Limuloides'', ''Bunodes'' was once though to have lateral eyes on the second pair of radiated ridges of its carapace, but later investigation did not find any evidence of it.

References

Synziphosurina Planaterga Silurian first appearances Silurian arthropods Fossils of England Fossil taxa described in 1854 {{Chelicerata-stub