The bull trout (''Salvelinus confluentus'') is a
char of the family
Salmonidae
Salmonidae is a family of ray-finned fish that constitutes the only currently extant family in the order Salmoniformes . It includes salmon (both Atlantic and Pacific species), trout (both ocean-going and landlocked), chars, freshwater whitefish ...
native to northwestern
North America.
Historically, ''S. confluentus'' has been known as the "
Dolly Varden" (''S. malma''), but was reclassified as a separate species in 1980.
Bull trout are listed as a
threatened species
Threatened species are any species (including animals, plants and fungi) which are vulnerable to endangerment in the near future. Species that are threatened are sometimes characterised by the population dynamics measure of ''critical depensa ...
under the U.S.
Endangered Species Act
The Endangered Species Act of 1973 (ESA or "The Act"; 16 U.S.C. § 1531 et seq.) is the primary law in the United States for protecting imperiled species. Designed to protect critically imperiled species from extinction as a "consequence of e ...
(1998)
[ and as vulnerable on the IUCN Red List of Threatened Species.
]
Description
Like other species of char, the fins of a bull trout have white leading edges. Its head and mouth are unusually large for salmonids, giving it its name. Bull trout have been recorded measuring up to in length and weighing .[ Bull trout may be either migratory, moving throughout large river systems, lakes, and the ocean, or they may be resident, remaining in the same stream their entire lives.][ Migratory bull trout are typically much larger than resident bull trout, which rarely exceed .][ Bull trout can be differentiated from brook trout (''S. fontinalis'') by the absence of distinct spots on the ]dorsal fin
A dorsal fin is a fin located on the back of most marine and freshwater vertebrates within various taxa of the animal kingdom. Many species of animals possessing dorsal fins are not particularly closely related to each other, though through c ...
, as well as yellow, orange, or salmon-colored spots on the back as opposed to red spots with blue halos on the brook trout. Bull trout lack the deeply forked tail fin of lake trout
The lake trout (''Salvelinus namaycush'') is a freshwater char living mainly in lakes in northern North America. Other names for it include mackinaw, namaycush, lake char (or charr), touladi, togue, and grey trout. In Lake Superior, it can als ...
(''S. namaycush'', another char).[
]
Distribution and habitat
''S. confluentus'' is found in the cold, clear waters of the high mountains and coastal rivers of northwestern North America, including Yukon
Yukon (; ; formerly called Yukon Territory and also referred to as the Yukon) is the smallest and westernmost of Canada's three territories. It also is the second-least populated province or territory in Canada, with a population of 43,964 as ...
, British Columbia
British Columbia (commonly abbreviated as BC) is the westernmost Provinces and territories of Canada, province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include ...
, Washington, Oregon
Oregon () is a state in the Pacific Northwest region of the Western United States. The Columbia River delineates much of Oregon's northern boundary with Washington, while the Snake River delineates much of its eastern boundary with Idah ...
, Idaho
Idaho ( ) is a state in the Pacific Northwest region of the Western United States. To the north, it shares a small portion of the Canada–United States border with the province of British Columbia. It borders the states of Montana and W ...
, and western Montana
Montana () is a U.S. state, state in the Mountain states, Mountain West List of regions of the United States#Census Bureau-designated regions and divisions, division of the Western United States. It is bordered by Idaho to the west, North ...
, as well as the Jarbidge River of northern Nevada
Nevada ( ; ) is a state in the Western region of the United States. It is bordered by Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. Nevada is the 7th-most extensive, ...
. A population of bull trout exists east of the Continental Divide
A continental divide is a drainage divide on a continent such that the drainage basin on one side of the divide feeds into one ocean or sea, and the basin on the other side either feeds into a different ocean or sea, or else is endorheic, no ...
in Alberta
Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...
, where it is the provincial fish. The historical range of bull trout also included northern California
California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the ...
, but they are likely extirpated
Local extinction, also known as extirpation, refers to a species (or other taxon) of plant or animal that ceases to exist in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinct ...
.[Washington's Native Chars]
, Washington Department of Fish and Wildlife
Bull trout have exacting habitat demands, requiring water temperatures generally below 55 °F (13 °C), clean gravel
Gravel is a loose aggregation of rock fragments. Gravel occurs naturally throughout the world as a result of sedimentary and erosive geologic processes; it is also produced in large quantities commercially as crushed stone.
Gravel is classif ...
beds, deep pools, complex cover such as snags and cut banks, and large systems of interconnected waterways to accommodate spawning migrations. Thus, they favor the deep pools of cold lakes and large rivers, as well as high, cold mountain headwaters. Bull trout may be anadromous
Fish migration is mass relocation by fish from one area or body of water to another. Many types of fish migrate on a regular basis, on time scales ranging from daily to annually or longer, and over distances ranging from a few metres to thousan ...
in coastal rivers, and individual bull trout have been found to have migrated from one coastal river to another by the ocean.[, King County, Washington]
Feeding
Young bull trout feed on zooplankton
Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...
and zoobenthos, especially chironomid
The Chironomidae (informally known as chironomids, nonbiting midges, or lake flies) comprise a family of nematoceran flies with a global distribution. They are closely related to the Ceratopogonidae, Simuliidae, and Thaumaleidae. Many species ...
s. As they grow larger, they begin to feed heavily upon other fish. In coastal Washington, some of the southernmost populations of bull trout feed heavily on salmon
Salmon () is the common name
In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of ...
eggs and fry, as well as fish.
Conservation
The bull trout is listed as a threatened species under the U.S. Endangered Species Act throughout its range in the contiguous United States
The contiguous United States (officially the conterminous United States) consists of the 48 adjoining U.S. states and the Federal District of the United States of America. The term excludes the only two non-contiguous states, Alaska and Hawai ...
.[ It is used as a management indicator species for several national forests, including Boise National Forest and ]Sawtooth National Forest
Sawtooth National Forest is a National Forest that covers 2,110,408 acres (854,052 ha) in the U.S. states of Idaho (~96 percent) and Utah (~4 percent). Managed by the U.S. Forest Service in the U.S. Department of Agriculture, it was originall ...
(Sawtooth National Recreation Area
The Sawtooth National Recreation Area (SNRA) is a national recreation area in central Idaho, United States that is managed as part of Sawtooth National Forest. The recreation area, established on August 22, 1972, is managed by the U.S. Forest S ...
). They can also be found in the Glacier National Park. Bull trout reproduction requires cold water and very low amounts of silt
Silt is granular material of a size between sand and clay and composed mostly of broken grains of quartz. Silt may occur as a soil (often mixed with sand or clay) or as sediment mixed in suspension with water. Silt usually has a floury feel wh ...
, both of which are negatively impacted by road building and logging. Additionally, its need to migrate throughout river systems may be hindered by impassible fish barriers, such as dams.[ Bull trout populations are also in danger from hybridization with non-native brook trout.][ Several of these issues were raised in a long-running lawsuit where in 2003, the Oregon Natural Desert Association and the Center for Biological Diversity sued the U.S. Forest Service claiming they violated the National Forest Management Act and ]Wild and Scenic Rivers Act
The National Wild and Scenic Rivers System was created by the Wild and Scenic Rivers Act of 1968 (Public Law 90-542), enacted by United States Congress, the U.S. Congress to preserve certain rivers with outstanding natural, cultural, and recreat ...
by approving grazing plans in Oregon's Malheur National Forest
The Malheur National Forest is a National Forest in the U.S. state of Oregon. It contains more than in the Blue Mountains of eastern Oregon. The forest consists of high desert grasslands, sage, juniper, pine, fir, and other tree species. ...
. In April, 2018, U.S. District Judge Michael W. Mosman
Michael Wise Mosman (born December 23, 1956) is a Senior United States district judge of the United States District Court for the District of Oregon. He served as Chief Judge for the U.S. District Court for the District of Oregon from February 1 ...
dismissed the complaint.
They are a prized game fish
Game fish, sport fish or quarry refer to popular fish pursued by recreational anglers, and can be freshwater or saltwater fish. Game fish can be eaten after being caught, or released after capture. Some game fish are also targeted commercia ...
in northern Canada
Northern Canada, colloquially the North or the Territories, is the vast northernmost region of Canada variously defined by geography and politics. Politically, the term refers to the three territories of Canada: Yukon, Northwest Territories and ...
. It was once maligned out of fear they threatened populations of other native species more prized by anglers. Some jurisdictions publicize the requirement to release with the slogan "No black, put it back".
Historical names
"Dolly Varden" in California
Historically, confusion has existed between ''S. confluentus'' and ''Salvelinus malma malma
The Dolly Varden trout (''Salvelinus malma'') is a species of salmonid fish native to cold-water tributaries of the Pacific Ocean in Asia and North America. It belongs to the genus ''Salvelinus'', or true chars, which includes 51 recognized spec ...
'', today commonly called the Dolly Varden trout
The Dolly Varden trout (''Salvelinus malma'') is a species of salmonid fish native to cold-water tributaries of the Pacific Ocean in Asia and North America. It belongs to the genus ''Salvelinus'', or true chars, which includes 51 recognized spe ...
. This was likely due to overlapping ranges and similar appearances among members of the two species.
The first recorded use of the name "Dolly Varden" for a fish species was applied to members of ''S. confluentus'' caught in the McCloud River
The McCloud River is a longU.S. Geological Survey. National Hydrography Dataset high-resolution flowline dataThe National Map, accessed March 10, 2011 river that flows east of and parallel to the upper Sacramento River, in Siskiyou County and ...
in northern California in the early 1870s. In his book ''Inland Fishes of California'', Peter Moyle recounted a letter from Mrs. Valerie Masson Gomez:
In 1874, Livingston Stone, a naturalist working for the U.S. government, wrote of this fish:
It is currently unknown whether the name "Dolly Varden" was later applied to ''S. m. malma'' because of its similar appearance to ''S. confluentus''; the two may have even been believed to be the same species. The name "Dolly Varden" may have also been given to ''S. m. malma'' independent of the McCloud River fish.
Ironically, the original "Dolly Varden" trout (i.e., ''S. confluentus'') apparently likely became extirpated
Local extinction, also known as extirpation, refers to a species (or other taxon) of plant or animal that ceases to exist in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinct ...
in the McCloud River in the 1970s, although reports continue of its being caught. Other fish species, typically introduced trout, outcompete ''S. confluentus'', and can interbreed with them, resulting in sterile hybrids. An attempt to reintroduce ''S. confluentus'' to the McCloud was unsuccessful, and no additional attempts are expected.
Other uses of "Dolly Varden"
The "Dolly Varden" name is also applied to the other subspecies of ''S. malma'', the ''S. m. krascheninnikova'', and ''S. m. miyabei'', found in Lake Shikaribetsu
A lake is an area filled with water, localized in a basin, surrounded by land, and distinct from any river or other outlet that serves to feed or drain the lake. Lakes lie on land and are not part of the ocean, although, like the much larger ...
on the island of Hokkaidō in Japan.Use of Dolly Varden name in Japan
/ref>
The name has also been applied to ''S. alpinus'', today more commonly known as Arctic char
The Arctic char or Arctic charr (''Salvelinus alpinus'') is a cold-water fish in the family Salmonidae, native to alpine lakes and arctic and subarctic coastal waters. Its distribution is Circumpolar North. It spawns in freshwater and popu ...
.
"Bull trout" in Europe
The name "bull trout" was also given in the past to some of the large sea trout that run the River Tweed
The River Tweed, or Tweed Water ( gd, Abhainn Thuaidh, sco, Watter o Tweid, cy, Tuedd), is a river long that flows east across the Border region in Scotland and northern England. Tweed cloth derives its name from its association with the R ...
and other rivers in Scotland
Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to th ...
and North East England
North East England is one of nine official regions of England at the first level of ITL for statistical purposes. The region has three current administrative levels below the region level in the region; combined authority, unitary authori ...
. Victorian anglers and others classified these as a separate race, but today they are biologically classified along with all other UK brown and sea trout as ''Salmo trutta
The brown trout (''Salmo trutta'') is a European species of salmonid fish that has been widely introduced into suitable environments globally. It includes purely freshwater populations, referred to as the riverine ecotype, ''Salmo trutta'' Morph ...
''. This does not deny that populations of ''S. trutta'' can differ appreciably in habits, size, and appearance from place to place, or indeed in the same river or lake.
References
{{Taxonbar, from=Q2429513
Salvelinus
Fish of Japan
Fish of North America
Fish of the Western United States
Provincial symbols of Alberta
Fish described in 1859
ESA threatened species