Bulimulinae

Bulimulidae is a taxonomic family of medium-sized to large, air-breathing, tropical and sub-tropical land snails, terrestrial pulmonate gastropod molluscs in the superfamily Orthalicoidea.MolluscaBase eds. (2020). MolluscaBase. Bulimulidae Tryon, 1867. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=549409 on 2020-06-27

Distribution

Distribution of species in the family Bulimulidae includes Ecuador (9 genera) and other South American countries. Some species also occur in North America.

Fossil record

The family's oldest fossil record dates from the late Cretaceous of Brazil (Itaboraí Basin).

Anatomy

Members of this family have a haploid chromosome number between 26 and 30 (according to the values in this table).Barker G. M.: Gastropods on Land: ''Phylogeny, Diversity and Adaptive Morphology''. in Barker G. M. (ed.):
The biology of terrestrial molluscs
'. CABI Publishing, Oxon, UK, 2001, . 1-146, cited pages: 139 and 142. Cited as Bulimulidae in the source.

Shells of species within Bostrycinae have a smooth protoconch and the genital organs feature a relatively long penis sheath (ca. 1/4–1/6 total phallus length).

Taxonomy

Previously, the members of the Orthalicidae were also included in this family, as the subfamily Orthalicinae, and the taxa listed here were placed in their own subfamily, the Bulimulinae.

2005 taxonomy

Bulimulinae was placed in the family Orthalicidae according to the taxonomy of the Gastropoda (Bouchet & Rocroi, 2005).

The subfamily Bulimulinae included 3 tribes:
* Tribe Bulimulini Tryon, 1867 - synonyms: Bulimidae Guilding, 1828 (inv.); Berendtiinae P. Fischer & Crosse, 1872; Bothiembryontidae Iredale, 1937
* Tribe Odontostomini Pilsbry & Vanatta, 1898 - synonym: Tomogeridae Jousseaume, 1877
* Tribe Simpulopsini Schileyko, 1999

2010 taxonomy

Breure et al. (2010) moved '' Bothriembryon'' to Placostylidae (since 2012 as Bothriembryontidae), elevated Bulimulinae to Bulimulidae and Odontostomini to Odontostomidae.

2012 taxonomy

Breure & Romero (2012) confirmed previous results from 2010, additionally they elevated Simpulopsini to Simpulopsidae. There are three subfamilies within Bulimulidae:

* subfamily Bulimulinae Tryon, 1867
* subfamily Peltellinae Gray, 1855
* subfamily Bostrycinae Breure, 2012

Genera

Genera in the family Bulimulidae include:
;subfamily Bulimulinae Tryon, 1887: (synonym: Berendtiinae P. Fischer & Crosse, 1872)
* '' Anctus'' E. von Martens, 1860
* '' Auris'' Spix, 1827
* '' Berendtia'' Crosse & P. Fischer, 1869
* '' Bocourtia'' Rochebrune, 1882
* '' Bulimulus'' Leach, 1814 - type genus of the subfamily Bulimulinae
* '' Cochlorina'' Jan, 1830
* '' Graptostracus'' Pilsbry, 1939
* ''Kora'' Simone, 2012
* '' Lopesianus'' Weyrauch, 1958
* '' Naesiotus'' Albers, 1850
* ''Neopetraeus'' E. von Martens, 1885
* '' Newboldius'' Pilsbry, 1932
* '' Otostomus'' H. Beck, 1837
* '' Oxychona'' Mörch, 1852
* '' Protoglyptus'' Pilsbry, 1897
* '' Pseudoxychona'' Pilsbry, 1930
* '' Rabdotus'' Albers, 1850
* '' Scutalus'' Albers, 1850
* '' Spartocentrum'' Dall, 1895
* '' Sphaeroconcha'' Breure, 1978
* '' Stenostylus'' Pilsbry, 1898
* '' Suniellus'' Breure, 1978
;subfamily Peltellinae
* '' Drymaeus'' Albers, 1850
* '' Peltella'' Gray, 1855 - type genus of the subfamily Peltellinae

;subfamily Bostrycinae
* ''Bostryx'' Troschel, 1847 sensu lato- type genus of the subfamily Bostrycinae

;Unassigned within Bulimulidae
* '' Bulimus'' Bruguière, 1789 (temporary name)
* ''Itaborahia'' Maury, 1935
* † '' Oreoconus'' D. W. Taylor in McKenna et al., 1962
* † '' Palaeobulimulus'' Parodiz, 1949
* '' Stapafurdius'' Simone, 2021
* † ''Tocobaga'' Auffenberg, Slapcinsky & Portell, 2015

Synonyms

* ''Adzharia'' P. Hesse, 1933: synonym of '' Bulimulus'' Leach, 1814 (junior synonym)
* ''Antidrymaeus'' Germain, 1907: synonym of ''Drymaeus (Mesembrinus)'' Albers, 1850 represented as '' Drymaeus'' Albers, 1850
* ''Atahualpa'' Strebel, 1910: synonym of '' Thaumastus'' E. von Martens, 1860
* ''Ataxellus'' Dall, 1912: synonym of ''Bostryx (Geopyrgus)'' Pilsbry, 1896 represented as''Bostryx'' Troschel, 1847
* ''Ataxus'' Albers, 1850: synonym of ''Bostryx'' Troschel, 1847
* ''Bilamelliferus'' Weyrauch, 1958: synonym of ''Bostryx'' Troschel, 1847 (junior synonymy)
* ''Cochlogena'' Férussac, 1821: synonym of '' Bulimulus'' Leach, 1814 (junior synonym)
* ''Diaphanomormus'' Weyrauch, 1964: synonym of ''Drymaeus (Mesembrinus)'' Albers, 1850 epresented as '' Drymaeus'' Albers, 1850
* ''Elatibostryx'' Weyrauch, 1958: synonym of ''Bostryx'' Troschel, 1847
* ''Floreziellus'' Weyrauch, 1967: synonym of ''Bostryx'' Troschel, 1847 (junior synonymy)
* ''Geoceras'' Pilsbry, 1896: synonym of ''Bostryx (Geopyrgus)'' Pilsbry, 1896 represented as ''Bostryx'' Troschel, 1847
* ''Globulinus'' Crosse & P. Fischer, 1875: synonym of '' Rabdotus'' Albers, 1850
* ''Goniognathmus'' Crosse & P. Fischer, 1875: synonym of '' Drymaeus'' Albers, 1850 (junior synonym)
* ''Hamadryas'' Albers, 1850: synonym of '' Drymaeus'' Albers, 1850
* ''Hannarabdotus'' Emerson & Jacobson, 1964: synonym of '' Rabdotus'' Albers, 1850
* ''Kionoptyx'' Haas, 1966: synonym of ''Bostryx'' Troschel, 1847 (junior synonymy)
* ''Kuschelenia'' Hylton Scott, 1951: synonym of ''Bocourtia (Kuschelenia)'' Hylton Scott, 1951 represented as '' Bocourtia'' Rochebrune, 1882
* ''Leptodrymaeus'' Pilsbry, 1946: synonym of ''Drymaeus (Mesembrinus)'' Albers, 1850 represented as '' Drymaeus'' Albers, 1850
* ''Leptomerus'' Albers, 1850: synonym of '' Bulimulus'' Leach, 1814
* ''Leptomormus'' Weyrauch, 1958: synonym of ''Drymaeus (Mesembrinus)'' Albers, 1850 represented as '' Drymaeus'' Albers, 1850
* ''Liostracus'' ic synonym of ''Leiostracus'' Albers, 1850 (misspelling of original genus, ''Leiostracus'' Albers, 1850)
* ''Loboa'' Ihering, 1917: synonym of '' Bulimulus'' Leach, 1814
* ''Mormus'' E. von Martens, 1860: synonym of '' Drymaeus'' Albers, 1850
* ''Multifasciatus'' Weyrauch, 1958: synonym of ''Bostryx (Geopyrgus)'' Pilsbry, 1896 represented as ''Bostryx'' Troschel, 1847
* ''Naesiotellus'' Weyrauch, 1967: synonym of ''Bostryx'' Troschel, 1847 (junior synonym)
* ''Navicula'' Spix, 1827: synonym of '' Cochlorina'' Jan, 1830
* ''Nesiotes'' E. von Martens, 1860: synonym of '' Naesiotus'' Albers, 1850
* ''Obstrussus'' Parodiz, 1946: synonym of '' Naesiotus'' Albers, 1850
* ''Olinodia'' Dall, 1920: synonym of ''Naesiotus (Reclasta)'' Dall, 1920 represented as '' Naesiotus'' Albers, 1850
* ''Orphaicus'' Schaufuss, 1869: synonym of '' Thaumastus'' E. von Martens, 1860
* ''Orphnus'' Albers, 1850: synonym of '' Thaumastus'' E. von Martens, 1860
* ''Orthotomium'' Crosse & P. Fischer, 1875: synonym of '' Rabdotus'' Albers, 1850
* ''Pachyotus'' H. Beck, 1837: synonym of ''Auris'' Spix, 1827
* ''Pachytholus'' Strebel, 1909: synonym of '' Thaumastus'' E. von Martens, 1860
* ''Pampasinus'' Weyrauch, 1958: synonym of ''Bostryx (Platybostryx)'' Pilsbry, 1896: synonym of''Bostryx'' Troschel, 1847
* ''Peronaeus'' Albers, 1850: synonym of ''Bostryx'' Troschel, 1847
* ''Phenacotaxus'' Dall, 1912: synonym of ''Bostryx (Geopyrgus)'' Pilsbry, 1896 represented as ''Bostryx'' Troschel, 1847
* ''Plecochilus'' Agassiz, 1846: synonym of '' Plekocheilus'' Guilding, 1827 (unjustifed emendation)
* ''Pseudoperonaeus'' Weyrauch, 1958: synonym of ''Bostryx (Geopyrgus)'' Pilsbry, 1896 represented as ''Bostryx'' Troschel, 1847
* ''Pseudorhodea'' Dall, 1895: synonym of ''Rabdotus (Plicolumna)'' J.G. Cooper, 1895 represented as '' Rabdotus'' Albers, 1850
* ''Pyrgus'' Albers, 1850: synonym of ''Bostryx (Geopyrgus)'' Pilsbry, 1896 represented as ''Bostryx'' Troschel, 1847
* ''Rhabdotus'' ic synonym of '' Rabdotus'' Albers, 1850 (incorrect subsequent spelling)
* ''Rhaphiellus'' L. Pfeiffer, 1856: synonym of '' Naesiotus'' Albers, 1850
* ''Rimatula'' Parodiz, 1946: synonym of '' Protoglyptus'' Pilsbry, 1897
* ''Saeronia'' Dall, 1920: synonym of '' Naesiotus'' Albers, 1850
* ''Semiclausaria'' L. Pfeiffer, 1856: synonym of '' Drymaeus'' Albers, 1850 (junior synonym)
* ''Siphalomphix'' Rafinesque, 1833: synonym of '' Bulimulus'' Leach, 1814
* ''Sonorina'' Pilsbry, 1896: synonym of ''Rabdotus (Leptobyrsus)'' Crosse & P. Fischer, 1875 represented as '' Rabdotus'' Albers, 1850
* ''Spiroscutalus'' Pilsbry, 1932: synonym of '' Scutalus'' Albers, 1850
* ''Stenostoma'' Spix, 1827: synonym of '' Anctus'' E. von Martens, 1860
* ''Tatutor'' Jousseaume, 1887: synonym of '' Thaumastus'' E. von Martens, 1860
* ''Teneritia'' J. Mabille, 1898: synonym of '' Spartocentrum'' Dall, 1895
* ''Tholus'' Strebel, 1909: synonym of '' Thaumastus'' E. von Martens, 1860
* '' Xenothauma'' Fulton 1896: Powell, A. W. B., 1979. ''New Zealand Mollusca'', William Collins Publishers Ltd, Auckland, New Zealand. . synonym of '' Scutalus'' Albers, 1850
* ''Zaplagius'' Pilsbry, 1896: synonym of '' Cochlorina'' Jan, 1830

References

This article incorporates CC-BY-3.0 text from the reference

* Bouchet P., Rocroi J.P., Hausdorf B., Kaim A., Kano Y., Nützel A., Parkhaev P., Schrödl M. & Strong E.E. (2017). Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia. 61(1-2): 1-526

Further reading

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External links

{{taxonbar, from=Q2927957

Taxa named by George Washington Tryon
Gastropod families