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The Morrison Formation is a distinctive sequence of
Upper Jurassic The Late Jurassic is the third epoch of the Jurassic Period, and it spans the geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic strata.Owen 1987. In European lithostratigraphy, the name ...
sedimentary rock Sedimentary rocks are types of rock (geology), rock formed by the cementation (geology), cementation of sediments—i.e. particles made of minerals (geological detritus) or organic matter (biological detritus)—that have been accumulated or de ...
found in the western United States which has been the most fertile source of
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
s in North America. It is composed of
mudstone Mudstone, a type of mudrock, is a fine-grained sedimentary rock whose original constituents were clays or muds. Mudstone is distinguished from ''shale'' by its lack of fissility.Blatt, H., and R.J. Tracy, 1996, ''Petrology.'' New York, New York, ...
,
sandstone Sandstone is a Clastic rock#Sedimentary clastic rocks, clastic sedimentary rock composed mainly of grain size, sand-sized (0.0625 to 2 mm) silicate mineral, silicate grains, Cementation (geology), cemented together by another mineral. Sand ...
,
siltstone Siltstone, also known as aleurolite, is a clastic sedimentary rock that is composed mostly of silt. It is a form of mudrock with a low clay mineral content, which can be distinguished from shale by its lack of fissility. Although its permeabil ...
, and
limestone Limestone is a type of carbonate rock, carbonate sedimentary rock which is the main source of the material Lime (material), lime. It is composed mostly of the minerals calcite and aragonite, which are different Polymorphism (materials science) ...
and is light gray, greenish gray, or red. Most of the fossils occur in the green siltstone beds and lower sandstones, relics of the rivers and floodplains of the
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
period. It is centered in
Wyoming Wyoming ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States, Western United States. It borders Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho t ...
and
Colorado Colorado is a U.S. state, state in the Western United States. It is one of the Mountain states, sharing the Four Corners region with Arizona, New Mexico, and Utah. It is also bordered by Wyoming to the north, Nebraska to the northeast, Kansas ...
, with outcrops in
Montana Montana ( ) is a landlocked U.S. state, state in the Mountain states, Mountain West subregion of the Western United States. It is bordered by Idaho to the west, North Dakota to the east, South Dakota to the southeast, Wyoming to the south, an ...
,
North Dakota North Dakota ( ) is a U.S. state in the Upper Midwest, named after the indigenous Dakota people, Dakota and Sioux peoples. It is bordered by the Canadian provinces of Saskatchewan and Manitoba to the north and by the U.S. states of Minneso ...
,
South Dakota South Dakota (; Sioux language, Sioux: , ) is a U.S. state, state in the West North Central states, North Central region of the United States. It is also part of the Great Plains. South Dakota is named after the Dakota people, Dakota Sioux ...
,
Nebraska Nebraska ( ) is a landlocked U.S. state, state in the Midwestern United States, Midwestern region of the United States. It borders South Dakota to the north; Iowa to the east and Missouri to the southeast, both across the Missouri River; Ka ...
,
Kansas Kansas ( ) is a landlocked U.S. state, state in the Midwestern United States, Midwestern region of the United States. It borders Nebraska to the north; Missouri to the east; Oklahoma to the south; and Colorado to the west. Kansas is named a ...
, the panhandles of
Oklahoma Oklahoma ( ; Choctaw language, Choctaw: , ) is a landlocked U.S. state, state in the South Central United States, South Central region of the United States. It borders Texas to the south and west, Kansas to the north, Missouri to the northea ...
and
Texas Texas ( , ; or ) is the most populous U.S. state, state in the South Central United States, South Central region of the United States. It borders Louisiana to the east, Arkansas to the northeast, Oklahoma to the north, New Mexico to the we ...
,
New Mexico New Mexico is a state in the Southwestern United States, Southwestern region of the United States. It is one of the Mountain States of the southern Rocky Mountains, sharing the Four Corners region with Utah, Colorado, and Arizona. It also ...
,
Arizona Arizona is a U.S. state, state in the Southwestern United States, Southwestern region of the United States, sharing the Four Corners region of the western United States with Colorado, New Mexico, and Utah. It also borders Nevada to the nort ...
,
Utah Utah is a landlocked state in the Mountain states, Mountain West subregion of the Western United States. It is one of the Four Corners states, sharing a border with Arizona, Colorado, and New Mexico. It also borders Wyoming to the northea ...
, and
Idaho Idaho ( ) is a landlocked U.S. state, state in the Pacific Northwest and Mountain states, Mountain West subregions of the Western United States. It borders Montana and Wyoming to the east, Nevada and Utah to the south, and Washington (state), ...
. Equivalent rocks under different names are found in Canada. It covers an area of 1.5 million square kilometers (600,000 square miles), although only a tiny fraction is exposed and accessible to
geologist A geologist is a scientist who studies the structure, composition, and History of Earth, history of Earth. Geologists incorporate techniques from physics, chemistry, biology, mathematics, and geography to perform research in the Field research, ...
s and
paleontologists Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
. Over 75% is still buried under the prairie to the east, and much of its western paleogeographic extent was eroded during
exhumation Burial, also known as interment or inhumation, is a method of final disposition whereby a dead body is placed into the ground, sometimes with objects. This is usually accomplished by excavating a pit or trench, placing the deceased and object ...
of the
Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in great-circle distance, straight-line distance from the northernmost part of Western Can ...
. It was named after
Morrison, Colorado Morrison is a home rule municipality in Jefferson County, Colorado, United States. The population was 396 at the 2020 census. History This small foothills settlement is named after George Morrison (April 16, 1822 – June 11, 1895), a builde ...
, where some of the first fossils in the formation were discovered by
Arthur Lakes Arthur Lakes (December 21, 1844—November 21, 1917) was an American geologist, artist, writer, teacher and Episcopalian minister. He captured much of his geological and palaeontological field work in sketches and watercolours. Lakes is credite ...
in 1877. That same year, it became the center of the
Bone Wars The Bone Wars, also known as the Great Dinosaur Rush, was a period of intense and ruthlessly competitive fossil hunting and discovery during the Gilded Age of American history, marked by a heated rivalry between Edward Drinker Cope (of the Aca ...
, a fossil-collecting rivalry between early paleontologists
Othniel Charles Marsh Othniel Charles Marsh (October 29, 1831 – March 18, 1899) was an American professor of paleontology. A prolific fossil collector, Marsh was one of the preeminent paleontologists of the nineteenth century. Among his legacies are the discovery or ...
and
Edward Drinker Cope Edward Drinker Cope (July 28, 1840 – April 12, 1897) was an American zoologist, paleontology, paleontologist, comparative anatomy, comparative anatomist, herpetology, herpetologist, and ichthyology, ichthyologist. Born to a wealthy Quaker fam ...
. In Colorado, New Mexico, and Utah, the Morrison Formation was a major source of
uranium Uranium is a chemical element; it has chemical symbol, symbol U and atomic number 92. It is a silvery-grey metal in the actinide series of the periodic table. A uranium atom has 92 protons and 92 electrons, of which 6 are valence electrons. Ura ...
ore.


Geologic history

According to
radiometric dating Radiometric dating, radioactive dating or radioisotope dating is a technique which is used to Chronological dating, date materials such as Rock (geology), rocks or carbon, in which trace radioactive impurity, impurities were selectively incorporat ...
, the Morrison Formation dates from 156.3 ± 2 million years old ( Ma) at its base, to 146.8 ± 1 million years old at the top, which places it in the earliest
Kimmeridgian In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 154.8 ±0.8 Ma and 149.2 ±0.7 Ma (million years ago). The Kimmeridgian follows the Oxfordian ...
, and early
Tithonian In the geological timescale, the Tithonian is the latest age (geology), age of the Late Jurassic Epoch and the uppermost stage (stratigraphy), stage of the Upper Jurassic Series. It spans the time between 149.2 ±0.7 annum, Ma and 143.1 ±0.6 (mi ...
stages Stage, stages, or staging may refer to: Arts and media Acting * Stage (theatre), a space for the performance of theatrical productions * Theatre, a branch of the performing arts, often referred to as "the stage" * ''The Stage'', a weekly Brit ...
of the late Jurassic. This is similar in age to the
Solnhofen Limestone The Solnhofen Limestone or Solnhofen Plattenkalk, formally known as the Altmühltal Formation, is a Jurassic Konservat-Lagerstätte that preserves a rare assemblage of fossilized organisms, including highly detailed imprints of soft bodied organi ...
Formation in Germany and the
Tendaguru Formation The Tendaguru Formation, or Tendaguru Beds are a highly List of fossiliferous stratigraphic units in Tanzania, fossiliferous Lithostratigraphy, formation and Lagerstätte located in the Lindi Region of southeastern Tanzania. The formation represe ...
in
Tanzania Tanzania, officially the United Republic of Tanzania, is a country in East Africa within the African Great Lakes region. It is bordered by Uganda to the northwest; Kenya to the northeast; the Indian Ocean to the east; Mozambique and Malawi to t ...
. The age and much of the fauna is similar to the
Lourinhã Formation The Lourinhã Formation () is a fossil-rich geological formation in western Portugal, named for the municipality of Lourinhã. The formation is mostly Late Jurassic in age (Kimmeridgian/Tithonian), with the top of the formation extending into the ...
in Portugal. Throughout the western United States, it variously overlies the Middle Jurassic Summerville,
Sundance A Sun Dance is a Native American ceremony. Sun dance or Sundance may also refer to: Places ;Canada * Sundance, Calgary, Alberta, a neighbourhood * Sundance, Manitoba, a ghost town ;United States * Sundance, New Mexico, a census-designated pl ...
, Bell Ranch, Wanakah, and Stump Formations. At the time, the supercontinent of
Laurasia Laurasia () was the more northern of two large landmasses that formed part of the Pangaea supercontinent from around ( Mya), the other being Gondwana. It separated from Gondwana (beginning in the late Triassic period) during the breakup of Pa ...
had recently split into the continents of North America and Eurasia, although they were still connected by land bridges. North America moved north and was passing through the subtropical regions. The Morrison Basin, which stretched from New Mexico in the south to Alberta and Saskatchewan in the north, was formed during the
Nevadan orogeny The Nevadan orogeny occurred along the western margin of North America during the Late Jurassic to Early Cretaceous approximately 155 Ma to 145 Ma. Throughout the duration of this orogeny there were at least two different kinds of orogenic process ...
, a precursor event to later orogenic episodes that created the Rocky Mountains started pushing up to the west. The deposits from their east-facing
drainage basin A drainage basin is an area of land in which all flowing surface water converges to a single point, such as a river mouth, or flows into another body of water, such as a lake or ocean. A basin is separated from adjacent basins by a perimeter, ...
s, carried by streams and
river A river is a natural stream of fresh water that flows on land or inside Subterranean river, caves towards another body of water at a lower elevation, such as an ocean, lake, or another river. A river may run dry before reaching the end of ...
s from the Elko Highlands (along the borders of present-day Nevada and Utah) and deposited in
swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
y lowlands, lakes, river channels and
floodplain A floodplain or flood plain or bottomlands is an area of land adjacent to a river. Floodplains stretch from the banks of a river channel to the base of the enclosing valley, and experience flooding during periods of high Discharge (hydrolog ...
s, became the Morrison Formation. In the north, the
Sundance Sea The Sundance Sea was an epeiric sea that existed in North America during the mid-to-late Jurassic Period of the Mesozoic Era. It was an arm of what is now the Arctic Ocean, and extended through what is now western Canada into the central western ...
, an extension of the
Arctic Ocean The Arctic Ocean is the smallest and shallowest of the world's five oceanic divisions. It spans an area of approximately and is the coldest of the world's oceans. The International Hydrographic Organization (IHO) recognizes it as an ocean, ...
, stretched through Canada down to the United States.
Coal Coal is a combustible black or brownish-black sedimentary rock, formed as rock strata called coal seams. Coal is mostly carbon with variable amounts of other Chemical element, elements, chiefly hydrogen, sulfur, oxygen, and nitrogen. Coal i ...
is found in the Morrison Formation of Montana, which means that the northern part of the formation, along the shores of the sea, was wet and swampy, with more vegetation. Aeolian, or wind-deposited sandstones, are found in the southwestern part, which indicates it was much more arid—a
desert A desert is a landscape where little precipitation occurs and, consequently, living conditions create unique biomes and ecosystems. The lack of vegetation exposes the unprotected surface of the ground to denudation. About one-third of the la ...
, with sand
dune A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, flat ...
s.


Stratigraphy

The Morrison Formation is subdivided into several members, the occurrence of which are varied across the geographic extent of the Morrison. Members are (in alphabetical order): * Bluff Sandstone Member (AZ, CO, NM, UT): Well-sorted, light brown to white sandstone with large grains and components of chert. Interpreted as being deposited in an aeolian setting, at the edge of a dune field. *Brushy Basin Member (AZ, CO, NM, UT): conglomerate interbedded with mudstone; up to fifty percent by volume is made up of altered vitric ash, which originated as felsic ash falls. Deposition likely occurred in a fluvial-lacustrine environment, with the lacustrine component tending towards playas. *Fiftymile Member (UT): Mainly present in the Kaiparowits basin, consisting of interbedded sandstone and mudstone, with minimal conglomerate. Locally, it is the uppermost member and has contact with the
Dakota Formation The Dakota is a sedimentary rock, sedimentary geologic unit name of Formation (stratigraphy), formation and Group (stratigraphy), group rank in Midwestern North America. The Dakota units are generally composed of sandstones, mudstones, clays, and ...
. *Jackpile Sandstone Member (NM): primarily a whitish crossbedded subarkose sandstone with a clay matrix. It is interbedded with variegated, pale-green to red, bentonitic mudstone lenses. However, recent
detrital zircon geochronology Detrital zircon geochronology is the science of geochronology, analyzing the age of zircons deposited within a specific sedimentary rock, sedimentary unit by examining their inherent radioisotopes, most commonly the uranium–lead dating, uranium ...
results have suggested that the Jackpile Sandstone Member is part of the
Burro Canyon Formation The Burro Canyon Formation is an Early Cretaceous Period sedimentary geologic formation, found in western Colorado, the Chama Basin and eastern San Juan Basin of northern New Mexico, and in eastern Utah, US. Description The lithology of the ...
. *Ralston Creek Member (CO): formerly a considered separate formation and recently reclassified as the basal member of the Morrison in eastern Colorado. It appears analogous to the Tidwell and Salt Wash Members. This reclassification is supported by more detailed examination of the contacts and radiometric dating. The Ralston Creek contains conglomerate, sandstone, gypsum-mudstone, and gypsum-sandstone-mudstone facies; it is undetermined if the gypsum is of marine or lacustrine origin. *Recapture Member (AZ, CO, NM, UT): forms the bottom of the Morrison across most of its range, overlying the
Entrada Entrada is a Spanish or Portuguese word meaning ''entry'' and may refer to: *Entrada Sandstone, a geological formation spread across Wyoming, Colorado, New Mexico, Arizona and Utah *Entradas, a town in Castro Verde, Portugal *La Entrada, a town in ...
and Wanakah Formations. Consists of clayey sandstone and
claystone Mudrocks are a class of fine-grained siliciclastic sedimentary rocks. The varying types of mudrocks include siltstone, claystone, mudstone and shale. Most of the particles of which the stone is composed are less than and are too small to ...
, representing a fluvial setting, interbedded with purely aeolian sandstone facies; in places, it also contains a large (up to nineteen percent) of
orthoclase Orthoclase, or orthoclase feldspar ( endmember formula K Al Si3 O8), is an important tectosilicate mineral which forms igneous rock. The name is from the Ancient Greek for "straight fracture", because its two cleavage planes are at right angles ...
feldspar inclusions. *Salt Wash Member (CO, UT): composed of fluvial sandstone, with occasional conglomeratic tendencies. *Tidwell Member (AZ, CO, NM, UT): in the northern part of the
Colorado Plateau The Colorado Plateau is a physiographic and desert region of the Intermontane Plateaus, roughly centered on the Four Corners region of the Southwestern United States. This plateau covers an area of 336,700 km2 (130,000 mi2) within w ...
, it is the basal member of the Morrison. Mainly composed of siltstone, shale, and sandstone, and occasionally incorporates limestone
clasts Clastic rocks are composed of fragments, or clasts, of pre-existing minerals and rock. A clast is a fragment of geological detritus,Essentials of Geology, 3rd Ed, Stephen Marshak, p. G-3 chunks, and smaller grains of rock broken off other rocks b ...
, along with thin beds of limestone. Depositional environments range from mudflats to fluvial, to evaporate and
lacustrine A lake is often a naturally occurring, relatively large and fixed body of water on or near the Earth's surface. It is localized in a basin or interconnected basins surrounded by dry land. Lakes lie completely on land and are separate from t ...
. The Morrison as a whole resembles the Tidwell. *Unkpapa Sandstone Member (SD): occurs primarily in western South Dakota as a well-sorted, fine-grained sandstone, consisting primarily of quartz, with some feldspar inclusions. Locally overlain by the
Lakota Formation The Lakota Formation is a sequence of rocks of early Cretaceous (Berriasian to Barremian) age from Western North America. Located in South Dakota, the name of the formation is derived from the Lakota people, Lakota Native Americans in the United ...
or the main body of the Morrison, and overlies the Redwater Shale Member of the
Sundance Formation The Sundance Formation is a western North American sequence of Middle Jurassic to Upper Jurassic age Dating from the Bathonian to the Oxfordian, around 168-157 Ma, It is up to 100 metres thick and consists of marine shale, sandy shale, sandstone ...
. Occasionally referred to as a separate formation, chiefly within the
Black Hills The Black Hills is an isolated mountain range rising from the Great Plains of North America in western South Dakota and extending into Wyoming, United States. Black Elk Peak, which rises to , is the range's highest summit. The name of the range ...
region. *Westwater Canyon Member (AZ, CO, NM, UT): consists of sandstone interbedded with mudstone lenses and the occasional conglomerate component. Deposited in a braided-stream environment, high in organic matter. The term "Poison Canyon Sandstone" is informally applied to the upper sandstone sections of the member. The Westwater Canyon Member is the main source of uranium ore in the Morrison, especially in the
San Juan Basin The San Juan Basin is a geologic structural basin located near the Four Corners region of the Southwestern United States. The basin covers 7,500 square miles and resides in northwestern New Mexico, southwestern Colorado, and parts of Utah a ...
. *Windy Hill Member (CO, SD, UT, WY): Formerly included as the upper member of the Sundance Formation, as, like the rest of the Sundance, it was deposited in marine settings; however, it is separated by an
unconformity An unconformity is a buried erosional or non-depositional surface separating two rock masses or strata of different ages, indicating that sediment deposition was not continuous. In general, the older layer was exposed to erosion for an interval ...
and interfingers with the Morrison, meriting the nomenclature shift. Composed of limey, fossiliferous sandstone, generally interpreted to be deposited in a marine setting. Other informal or disused designations of the Morrison include the Stockett Bed in Montana, an unofficial sub-unit which contains bituminous coal; the outdated terms Casamero, Chavez, and Prewitt Sandstone for the Brushy Basin, Recapture, and Westwater Canyon, respectively; and the Bullington Member, which has been discarded entirely.


Fossil content

Though many of the Morrison Formation fossils are fragmentary, they are sufficient to provide a good picture of the flora and fauna in the Morrison Basin during the Kimmeridgian. Overall, the
climate Climate is the long-term weather pattern in a region, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteoro ...
was dry, similar to a
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...
but, since there were no
angiosperm Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed within a fruit ...
s (
grass Poaceae ( ), also called Gramineae ( ), is a large and nearly ubiquitous family (biology), family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos, the grasses of natural grassland and spe ...
es,
flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
s, and some
trees In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only p ...
), the flora was quite different.
Conifers Conifers () are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All e ...
, the dominant plants of the time, were to be found with
ginkgo ''Ginkgo'' is a genus of non-flowering seed plants, assigned to the gymnosperms. The scientific name is also used as the English common name. The order to which the genus belongs, Ginkgoales, first appeared in the Permian, , and ''Ginkgo'' is n ...
s,
cycad Cycads are seed plants that typically have a stout and woody (ligneous) trunk (botany), trunk with a crown (botany), crown of large, hard, stiff, evergreen and (usually) pinnate leaves. The species are dioecious, that is, individual plants o ...
s,
tree fern Tree ferns are arborescent (tree-like) ferns that grow with a trunk (botany), trunk elevating the fronds above ground level, making them trees. Many extant tree ferns are members of the order Cyatheales, to which belong the families Cyatheaceae ( ...
s, and horsetail rushes. Much of the fossilized vegetation was
riparian A riparian zone or riparian area is the interface between land and a river or stream. In some regions, the terms riparian woodland, riparian forest, riparian buffer zone, riparian corridor, and riparian strip are used to characterize a ripar ...
, living along the river flood plains. Along the rivers, there were
fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...
,
frog A frog is any member of a diverse and largely semiaquatic group of short-bodied, tailless amphibian vertebrates composing the order (biology), order Anura (coming from the Ancient Greek , literally 'without tail'). Frog species with rough ski ...
s,
salamander Salamanders are a group of amphibians typically characterized by their lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. All t ...
s,
lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...
s,
crocodile Crocodiles (family (biology), family Crocodylidae) or true crocodiles are large, semiaquatic reptiles that live throughout the tropics in Africa, Asia, the Americas and Australia. The term "crocodile" is sometimes used more loosely to include ...
s,
turtle Turtles are reptiles of the order (biology), order Testudines, characterized by a special turtle shell, shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Crypt ...
s,
pterosaur Pterosaurs are an extinct clade of flying reptiles in the order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosaurs are the earli ...
s,
crayfish Crayfish are freshwater crustaceans belonging to the infraorder Astacidea, which also contains lobsters. Taxonomically, they are members of the superfamilies Astacoidea and Parastacoidea. They breathe through feather-like gills. Some spe ...
,
clam Clam is a common name for several kinds of bivalve mollusc. The word is often applied only to those that are deemed edible and live as infauna, spending most of their lives halfway buried in the sand of the sea floor or riverbeds. Clams h ...
s, and mammaliforms. The
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
s were most likely riparian, as well. Hundreds of dinosaur fossils have been discovered, such as ''
Allosaurus ''Allosaurus'' ( ) is an extinct genus of theropod dinosaur that lived 155 to 145 million years ago during the Late Jurassic period ( Kimmeridgian to late Tithonian ages). The first fossil remains that could definitively be ascribed to th ...
'', ''
Ceratosaurus ''Ceratosaurus'' (from Greek 'horn' and 'lizard') is a genus of carnivorous theropod dinosaur that lived in the Late Jurassic period (Kimmeridgian to Tithonian ages). The genus was first described in 1884 by American paleontologist Othni ...
'', ''
Torvosaurus ''Torvosaurus'' () is a genus of large Megalosaurinae, megalosaurine Theropoda, theropod dinosaur that lived approximately 165 to 148 million years ago during the Callovian to Tithonian ages of the late Middle Jurassic, Middle and Late Jurassi ...
'', ''
Camptosaurus ''Camptosaurus'' ( ) is a genus of plant-eating, beaked ornithischian dinosaurs of the Late Jurassic Period (geology), period of western North America and possibly also Europe. The name means 'flexible lizard' (Ancient Greek, Greek (') meaning ...
'', ''
Ornitholestes ''Ornitholestes'' (meaning "bird robber") is a small theropod dinosaur of the late Jurassic ( Brushy Basin Member of the Morrison Formation, middle Kimmeridgian age, about 154 million years agoTurner, C.E. and Peterson, F., (1999). "Biostratigrap ...
'', several
stegosaurs Stegosauria is a group of Herbivore, herbivorous ornithischian dinosaurs that lived during the Jurassic and early Cretaceous Period (geology), periods. Stegosaurian fossils have been found mostly in the Northern Hemisphere (North America, Europe a ...
comprising at least two species of ''
Stegosaurus ''Stegosaurus'' (; ) is a genus of herbivorous, four-legged, armored dinosaur from the Late Jurassic, characterized by the distinctive kite-shaped upright plates along their backs and spikes on their tails. Fossils of the genus have been fo ...
'' and the slightly older ''
Hesperosaurus ''Hesperosaurus'' (meaning "western lizard", from Classical Greek (') "western" and (') "lizard") is a herbivorous stegosaurian dinosaur from the Kimmeridgian age of the Jurassic period, approximately 156 million years ago. Fossils of ''Hesper ...
'', and the early
ankylosaurs Ankylosauria is a group of herbivorous dinosaurs of the clade Ornithischia. It includes the great majority of dinosaurs with Armour (zoology), armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short ...
, ''
Mymoorapelta ''Mymoorapelta'' (Meaning "Shield from the Mygatt-Moore Quarry" after a combination of the names of the discoverers of the Mygatt-Moore Quarry that fossils were originally collected from, and the Greek word πέλτα, meaning "shield") is a Nod ...
'' and ''
Gargoyleosaurus ''Gargoyleosaurus'' (meaning "gargoyle lizard") is one of the earliest ankylosaurs known from reasonably complete fossil remains. The holotype was discovered in 1995 at the Bone Cabin Quarry West locality, in Albany County, Wyoming in exposures ...
'', most notably a very broad range of
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
s (the giants of the
Mesozoic The Mesozoic Era is the Era (geology), era of Earth's Geologic time scale, geological history, lasting from about , comprising the Triassic, Jurassic and Cretaceous Period (geology), Periods. It is characterized by the dominance of archosaurian r ...
era). Since at least some of these species are known to have nested in the area (''Camptosaurus''
embryo An embryo ( ) is the initial stage of development for a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male sp ...
es have been discovered), there are indications that it was a good
environment Environment most often refers to: __NOTOC__ * Natural environment, referring respectively to all living and non-living things occurring naturally and the physical and biological factors along with their chemical interactions that affect an organism ...
for dinosaurs and not just home to migratory, seasonal populations. However, the large body mass of the sauropods has been interpreted as an adaptation to migration in times of drought. Sauropods that have been discovered include ''
Diplodocus ''Diplodocus'' (, , or ) is an extinct genus of diplodocid sauropod dinosaurs known from the Late Jurassic of North America. The first fossils of ''Diplodocus'' were discovered in 1877 by S. W. Williston. The generic name, coined by Othnie ...
'' (most famously, the first nearly complete specimen of ''D. carnegii'', which is now exhibited at the
Carnegie Museum of Natural History The Carnegie Museum of Natural History (abbreviated as CMNH) is a natural history museum in the Oakland (Pittsburgh), Oakland neighborhood of Pittsburgh, Pennsylvania. It was founded by List of people from the Pittsburgh metropolitan area, Pit ...
, in Pittsburgh, Pennsylvania), ''
Camarasaurus ''Camarasaurus'' ( ) is a genus of sauropod dinosaur that lived in North America during the Late Jurassic period. Its fossil remains have been found in the Morrison Formation, dating to the Kimmeridgian and Tithonian ages of the Jurassic, betwe ...
'' (the most commonly found sauropod), ''
Brachiosaurus ''Brachiosaurus'' () is a genus of sauropod dinosaur that lived in North America during the Late Jurassic, about . It was first Species description, described by American paleontologist Elmer S. Riggs in 1903 in paleontology, 1903 from fossi ...
'', ''
Apatosaurus ''Apatosaurus'' (; meaning "deceptive lizard") is a genus of herbivorous sauropod dinosaur that lived in North America during the Late Jurassic period. Othniel Charles Marsh described and named the first-known species, ''A. ajax'', in 1877, a ...
'', ''
Brontosaurus ''Brontosaurus'' (; meaning "thunder lizard" from the Ancient Greek, Greek words , "thunder" and , "lizard") is a genus of herbivorous sauropod dinosaur that lived in present-day United States during the Late Jurassic period. It was describe ...
'', ''
Barosaurus ''Barosaurus'' ( ) is a genus of giant, long-tailed, long-necked, herbivore, plant-eating Sauropoda, sauropod dinosaur closely related to the more familiar ''Diplodocus''. Remains have been found in the Morrison Formation from the Jurassic, Up ...
'', the uncommon ''
Haplocanthosaurus ''Haplocanthosaurus'' (meaning "simple spined lizard") is a genus of diplodocoid sauropod dinosaur. Two species, ''H. delfsi'' and ''H. priscus'', are known from incomplete fossil skeletons. They lived during the Late Jurassic (Kimmeridgian sta ...
'' and ''
Supersaurus ''Supersaurus'' (meaning "super lizard") is a genus of diplodocid sauropod dinosaur that lived in North America during the Late Jurassic period. The type species, ''S. vivianae'', was first discovered by Vivian Jones of Delta, Colorado, in the ...
''. The very diversity of the sauropods has raised some questions about how they could all co-exist. While their body shapes are very similar (long neck, long tail, huge elephant-like body), they are assumed to have had very different feeding strategies, in order for all to have existed in the same time frame and similar environment.


Sites and quarries

Locations where significant Morrison Formation fossil discoveries have been made include:


Colorado

*
Garden Park, Colorado Garden Park is a paleontological site in Fremont County, Colorado, known for its Jurassic dinosaurs and the role the specimens played in the infamous Bone Wars of the late 19th century. Located north of Cañon City, the name originates from the ...
: One of the three major sites excavated by the paleontologists
Othniel Charles Marsh Othniel Charles Marsh (October 29, 1831 – March 18, 1899) was an American professor of paleontology. A prolific fossil collector, Marsh was one of the preeminent paleontologists of the nineteenth century. Among his legacies are the discovery or ...
and
Edward Drinker Cope Edward Drinker Cope (July 28, 1840 – April 12, 1897) was an American zoologist, paleontology, paleontologist, comparative anatomy, comparative anatomist, herpetology, herpetologist, and ichthyology, ichthyologist. Born to a wealthy Quaker fam ...
during the Bone Wars in 1877, though most of the specimens were too incomplete to classify (''nomina dubia'') during the 1877-78 field seasons. The first nearly complete skeletons of ''
Stegosaurus ''Stegosaurus'' (; ) is a genus of herbivorous, four-legged, armored dinosaur from the Late Jurassic, characterized by the distinctive kite-shaped upright plates along their backs and spikes on their tails. Fossils of the genus have been fo ...
,
Ceratosaurus ''Ceratosaurus'' (from Greek 'horn' and 'lizard') is a genus of carnivorous theropod dinosaur that lived in the Late Jurassic period (Kimmeridgian to Tithonian ages). The genus was first described in 1884 by American paleontologist Othni ...
,'' and ''
Allosaurus ''Allosaurus'' ( ) is an extinct genus of theropod dinosaur that lived 155 to 145 million years ago during the Late Jurassic period ( Kimmeridgian to late Tithonian ages). The first fossil remains that could definitively be ascribed to th ...
'' were discovered at the site, including the type specimens of the former two and the proposed neotype of ''Allosaurus fragilis,'' in the 1883-1886 Yale field seasons. In 1992, a specimen of ''Stegosaurus stenops'' was discovered with its
armor Armour (Commonwealth English) or armor (American English; see American and British English spelling differences#-our, -or, spelling differences) is a covering used to protect an object, individual, or vehicle from physical injury or damage, e ...
still in place, which confirmed that the dinosaur had two rows of plates on its back. *
Dry Mesa Quarry The Dry Mesa Dinosaur Quarry is situated in southwestern Colorado, United States, near the town of Delta Delta commonly refers to: * Delta (letter) (Δ or δ), the fourth letter of the Greek alphabet * D (NATO phonetic alphabet: "Delta"), the fo ...
, Colorado: A wide variety of fauna, as well as the most diverse set of dinosaurs from any Morrison Formation quarry. The first dig was in 1972, by researchers from
Brigham Young University Brigham Young University (BYU) is a Private education, private research university in Provo, Utah, United States. It was founded in 1875 by religious leader Brigham Young and is the flagship university of the Church Educational System sponsore ...
. Unique specimens include the longest dinosaur known, ''
Supersaurus ''Supersaurus'' (meaning "super lizard") is a genus of diplodocid sauropod dinosaur that lived in North America during the Late Jurassic period. The type species, ''S. vivianae'', was first discovered by Vivian Jones of Delta, Colorado, in the ...
'', the chimeric ''
Ultrasauros ''Supersaurus'' (meaning "super lizard") is a genus of diplodocid sauropod dinosaur that lived in North America during the Late Jurassic period. The type species, ''S. vivianae'', was first discovered by Vivian Jones of Delta, Colorado, in the ...
'', and the largest
carnivore A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they ar ...
on the continent, ''
Torvosaurus ''Torvosaurus'' () is a genus of large Megalosaurinae, megalosaurine Theropoda, theropod dinosaur that lived approximately 165 to 148 million years ago during the Callovian to Tithonian ages of the late Middle Jurassic, Middle and Late Jurassi ...
''. * Fruita Paleontological Resource Area: Badlands sites located south of
Fruita Fruita () is a home rule municipality located in western Mesa County, Colorado, United States. The city population was 13,395 at the 2020 United States census. Fruita is a part of the Grand Junction, Colorado Metropolitan Statistical Area and ...
, were actively worked by George Callison from California State University and the Los Angeles County Museum of Natural History. Numerous specimens of mammals, lizards, and crocodiles were found. Most recently, '' Fruitafossor windscheffelia'' and the new dinosaur ''
Fruitadens ''Fruitadens'' is a genus of heterodontosaurid dinosaur. The name means "Fruita teeth", in reference to Fruita, Colorado (USA), where its fossils were first found. It is known from partial skulls and skeletons from at least four individuals of ...
'' were described from the area. *
Purgatoire River track site The Purgatoire River track site, also called the Picketwire Canyonlands tracksite, is one of the largest dinosaur Fossil trackway, tracksites in North America. The site is located on public land of the Comanche National Grassland, along the ...
, Otero County.


Utah

*
Cleveland-Lloyd Dinosaur Quarry Jurassic National Monument, at the site of the Cleveland-Lloyd Dinosaur Quarry, well known for containing the densest concentration of Jurassic dinosaur fossils ever found, is a paleontological site located near Cleveland, Utah, in the San Rafa ...
, Utah: First excavated by geologists from the University of Utah in the late 1920s.
William Lee Stokes William Lee Stokes (March 27, 1915, Black Hawk, Carbon County, Utah - December 12, 1994) was a geologist and paleontologist who is best known for his work at Cleveland-Lloyd Dinosaur Quarry in Emery County, Utah. William Stokes lived to be 79 ...
led an expedition from Princeton in 1939. During the Jurassic, the quarry was likely an ephemeral pond, where dinosaurs gathered and died due to severe drought. Their bodies were reworked by seasonal flooding events, which also added other partial carcasses from elsewhere. ''
Allosaurus fragilis ''Allosaurus'' ( ) is an extinct genus of theropoda, theropod dinosaur that lived 155 to 145 million years ago during the Late Jurassic Geologic time scale, period (Kimmeridgian to late Tithonian Geologic time scale, ages). The first fossil ...
'' is by far the most common dinosaur at this site, making it a model organism for studies of paleobiology in basal theropods. The rare theropods ''
Stokesosaurus ''Stokesosaurus'' (meaning "Stokes' lizard") is a genus of small (around in length), carnivorous early tyrannosauroid theropod dinosaurs from the late Jurassic period of Utah, United States. History From 1960 onwards Utah geologist William ...
'' and ''
Marshosaurus ''Marshosaurus'' is a genus of medium-sized carnivorous Theropoda, theropod dinosaur, belonging to the family Piatnitzkysauridae, from the Late Jurassic Morrison Formation of Utah and possibly Colorado. Discovery and naming During the 1960s, o ...
'' specimens were also first discovered here. *
Dinosaur National Monument Dinosaur National Monument is an American national monument located on the southeast flank of the Uinta Mountains on the border between Colorado and Utah at the confluence of the Green River (Colorado River tributary), Green and Yampa River, Y ...
, Utah: First excavated by Earl Douglas working for the Carnegie Museum in 1909 with the purpose of finding sauropods from the Morrison Formation for public display. Monument also has fossilized dinosaurs from the
Cedar Mountain Formation The Cedar Mountain Formation is the name given to a distinctive sedimentary geologic formation in eastern Utah, spanning most of the early and mid-Cretaceous. The formation was named for Cedar Mountain (Utah), Cedar Mountain in northern Emery Cou ...
. *
Hanksville-Burpee Quarry Hanksville-Burpee Quarry is a paleontological excavation site approximately wide by long near Hanksville, Utah, US, where scientists have found a large mix of remains of sauropods, trees, freshwater clams and other species dating between 145 mi ...
, Hanksville


Wyoming

*
Bone Cabin Quarry Bone Cabin Quarry is a dinosaur quarry that lay approximately northwest of Laramie, Wyoming, near historic Como Bluff. During the summer of 1897 Walter Granger, a paleontologist from the American Museum of Natural History, came upon a hillside l ...
, Wyoming *
Como Bluff Como Bluff is a long ridge extending east–west, located between the towns of Rock River and Medicine Bow, Wyoming. The ridge is an anticline, formed as a result of compressional geological folding. Three geological formations, the Sundance, ...
, Wyoming: One of the most renowned fossil sites in North America. It was first worked by Cope and particularly Marsh in 1877 and has been the source of many different sauropods and non-dinosaur species. The
Cloverly Formation The Cloverly Formation is a formation (geology), geological formation of Early Cretaceous, Early and Late Cretaceous age (Valanginian to Cenomanian stage) that is present in parts of Montana, Wyoming, Colorado and Utah in the western United State ...
from the Cretaceous and some
Triassic The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ...
strata are also exposed at this location. * The
Wyoming Dinosaur Center The Wyoming Dinosaur Center is located in Thermopolis, Wyoming and is one of the few dinosaur museums in the world to have excavation sites within driving distance. The museum displays the Thermopolis Specimen of ''Archaeopteryx'', which is one of ...
,
Thermopolis Thermopolis is the county seat and most populous town in Hot Springs County, Wyoming, United States. As of the 2020 U.S. Census, the town population was 2,725. Thermopolis, Greek for "hot city", is home to numerous natural hot springs, in which ...
* Ten Sleep, including Dana Quarry from where at least 12 sauropods and theropods are recovered.


Economic geology

The Morrison Formation contains
uranium Uranium is a chemical element; it has chemical symbol, symbol U and atomic number 92. It is a silvery-grey metal in the actinide series of the periodic table. A uranium atom has 92 protons and 92 electrons, of which 6 are valence electrons. Ura ...
deposits, including the Jackpile uranium body discovered near
Grants, New Mexico Grants is a city in Cibola County, New Mexico, United States. It is located about west of Albuquerque. The population was 9,163 at the 2020 Census. It is the county seat of Cibola County. Grants is located along the Trails of the Ancients B ...
in 1951. The ore deposits in the rich Grants mineral belt are concentrated in sandstone beds of the Westwater Canyon Member and the Jackpile Member. Mines in this belt produced of U3O8 between 1948 and 2002. The uranium was precipitated by plant debris and
humate Humate was until 1996 a German trance group, then just the German record producer Gerret Frerichs, who has released singles with record labels such as MFS and Superstition. His work has been featured on many compilation albums including the '' G ...
that acted as reducing agents.


See also

* Chugwater Formation *
Lists of dinosaur-bearing stratigraphic units This list of dinosaur-bearing rock formations is a list of geologic formations in which dinosaur fossils have been documented. * List of stratigraphic units with dinosaur body fossils * List of stratigraphic units with dinosaur trace fossils ** ...
*
List of fossil sites This list of fossil sites is a worldwide list of localities known well for the presence of fossils. Some entries in this list are notable for a single, unique find, while others are notable for the large number of fossils found there. Many of ...
''(with link directory)'' * List of Paleobiota of the Morrison Formation *
Uranium mining and the Navajo people The relationship between uranium mining and the Navajo people began in 1944 in northeastern Arizona, northwestern New Mexico, and southeastern Utah. In the 1950s, the Navajo Nation was situated directly in the uranium mining belt that experience ...


Citations


Further reading

* Foster, J. 2007. Jurassic West: The Dinosaurs of the Morrison Formation and Their World. Indiana University Press. 389pp. * Foster, J.R. 2003. Paleoecological Analysis of the Vertebrate Fauna of the Morrison Formation (Upper Jurassic), Rocky Mountain Region, U.S.A. Albuquerque, New Mexico: New Mexico Museum of Natural History and Science. Bulletin 23. * Jenkins, J.T. and J.L. Jenkins. 1993. Colorado's Dinosaurs. Denver, Colorado: Colorado Geologic Survey. Special Publication 35. * Mateus, O. 2006. Late Jurassic dinosaurs from the Morrison Formation, the Lourinhã and Alcobaça Formations (Portugal), and the Tendaguru Beds (Tanzania): a comparison. in Foster, J.R. and Lucas, S. G. R.M., eds., 2006, Paleontology and Geology of the Upper Jurassic Morrison Formation. New Mexico Museum of Natural History and Science Bulletin 36: 223-231.


External links


Morrison Natural History Museum
home page. *

', Columbia University lecture on the Morrison Formation.

from the National Park Service.
Spatial distribution of Morrison in Macrostrat

Paleobiology Database: Dalton Well Dinosaur Track Site (Morrison Formation): Late/Upper Jurassic, Utah

Paleobiology Database: Black Ridge Dinosaur Track Site (Morrison Formation): Late/Upper Jurassic, Colorado




* ttps://www.britannica.com/place/Morrison-Formation Britannica.com: Morrison Formation geology {{Chronostratigraphy of Colorado, Mesozoic state=expanded Geologic formations of the United States Jurassic System of North America Late Jurassic North America Jurassic United States Kimmeridgian Stage Tithonian Stage Mudstone formations of the United States Siltstone formations of the United States Limestone formations of the United States Sandstone formations of the United States Fluvial deposits Shallow marine deposits Jurassic fossil record Fossiliferous stratigraphic units of North America Jurassic paleontological sites of North America Jurassic Arizona Jurassic Colorado Jurassic Idaho Jurassic Kansas Jurassic Montana Jurassic Nebraska Jurassic formations of New Mexico Jurassic North Dakota Jurassic geology of Oklahoma Jurassic geology of South Dakota Jurassic Texas Jurassic geology of Utah Jurassic geology of Wyoming Colorado Plateau Dinosaur National Monument Geology of the Rocky Mountains