The brown roofed turtle (''Pangshura smithii'') is a

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

of turtle in the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Geoemydidae The Geoemydidae (formerly known as Bataguridae) are one of the largest and most diverse families in the order Testudines (turtles), with about 70 species A species () is often defined as the largest group of organisms in which any two in ...

. The species is

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

South Asia South Asia is the southern Subregion#Asia, subregion of Asia that is defined in both geographical and Ethnicity, ethnic-Culture, cultural terms. South Asia, with a population of 2.04 billion, contains a quarter (25%) of the world's populatio ...

. Two subspecies are recognized.

Etymology

The specific name, ''smithii'', is in honor of Scottish

zoologist Zoology ( , ) is the scientific study of animals. Its studies include the structure, embryology, classification, habits, and distribution of all animals, both living and extinct, and how they interact with their ecosystems. Zoology is one ...

Andrew Smith.

Description

The

carapace A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the unde ...

of ''P. smithii'' is much depressed and feebly keeled. The nuchal shield is small, trapezoidal, and broadest posteriorly. The first vertebral has sinuous lateral borders and is usually a little narrower in front than behind. The second vertebral is shortest, broader than long, and usually with straight or slightly convex posterior border. The third vertebral is considerably longer than broad, subquadrangular, and its posterior border is straight or slightly convex. The fourth vertebral is longest, tapering anteriorly and forming a narrow suture with the third. The fifth vertebral is much broader than the others. The large

plastron The turtle shell is a shield for the ventral and dorsal parts of turtles (the Order (biology), order Testudines), completely enclosing all the turtle's vital organs and in some cases even the head. It is constructed of modified bony elements such ...

is feebly angulated laterally. The front lobe is rounded. The hind lobe is angularly notched and as long as or a little shorter than the width of the bridge. The longest median suture is that between the abdominals, which about equals the length of the front lobe. The gulars are usually shorter than the suture between the humerals, their suture with the latter shields forming a right angle. The inguinal is large, and the axillary is smaller. The head is moderate. The snout is short, obtuse, and feebly prominent. The jaws have denticulated edges. The upper jaw is not notched mesially. The alveolar surface of the upper jaw is broad, and the median ridge is nearer the inner than the outer border. There are bony choanae between the orbits. The width of the lower jaw at the symphysis is less than the diameter of the orbit. The fore limbs have large transverse scales. The carapace is pale olive-brown above, and the dorsal keel is usually blackish. The plastral shields and the lower surface of the marginals are dark brown, bordered with yellow. The straight-line carapace length is . Brown roofed turtles exhibit ZZ/ZW sex determination, in contrast to the

temperature-dependent sex determination Temperature-dependent sex determination (TSD) is a type of environmental sex determination in which the temperatures experienced during embryonic/larval development determine the sex of the offspring. It is observed in reptiles and teleost fish, ...

of most turtles.

Subspecies and geographic ranges

Two subspecies are recognized, including the

nominotypical subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...

. *''P. s. smithii'' , is found in the

Indus The Indus ( ) is a transboundary river of Asia and a trans- Himalayan river of South and Central Asia. The river rises in mountain springs northeast of Mount Kailash in the Western Tibet region of China, flows northwest through the dis ...

Ganges The Ganges ( ; in India: Ganga, ; in Bangladesh: Padma, ). "The Ganges Basin, known in India as the Ganga and in Bangladesh as the Padma, is an international which goes through India, Bangladesh, Nepal and China." is a trans-boundary rive ...

, and

Brahmaputra The Brahmaputra is a trans-boundary river which flows through Southwestern China, Northeastern India, and Bangladesh. It is known as Brahmaputra or Luit in Assamese, Yarlung Tsangpo in Tibetan, the Siang/Dihang River in Arunachali, and ...

River drainages in

Bangladesh Bangladesh, officially the People's Republic of Bangladesh, is a country in South Asia. It is the List of countries and dependencies by population, eighth-most populous country in the world and among the List of countries and dependencies by ...

India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...

, and

Pakistan Pakistan, officially the Islamic Republic of Pakistan, is a country in South Asia. It is the List of countries and dependencies by population, fifth-most populous country, with a population of over 241.5 million, having the Islam by country# ...

. *''P. s. pallidipes'' , is found in the northern tributaries of the Ganges in India and

Nepal Nepal, officially the Federal Democratic Republic of Nepal, is a landlocked country in South Asia. It is mainly situated in the Himalayas, but also includes parts of the Indo-Gangetic Plain. It borders the Tibet Autonomous Region of China Ch ...

Habitat

The preferred natural

habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...

s of ''P. smithii'' are freshwater

swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...

s and rivers. These creatures also prefer to bask in the sun, twice a day. They often prefer muddy areas. www.reptile-database.org.

References

External links