Brown Argus
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''Aricia agestis'', the brown argus, is a
butterfly Butterflies are winged insects from the lepidopteran superfamily Papilionoidea, characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The oldest butterfly fossi ...
in the family
Lycaenidae Lycaenidae is the second-largest family (biology), family of butterflies (behind Nymphalidae, brush-footed butterflies), with over 6,000 species worldwide, whose members are also called gossamer-winged butterflies. They constitute about 30% of ...
. It is found throughout the
Palearctic realm The Palearctic or Palaearctic is a biogeographic realm of the Earth, the largest of eight. Confined almost entirely to the Eastern Hemisphere, it stretches across Europe and Asia, north of the foothills of the Himalayas, and North Africa. The ...
, north to northern
Jutland Jutland (; , ''Jyske Halvø'' or ''Cimbriske Halvø''; , ''Kimbrische Halbinsel'' or ''Jütische Halbinsel'') is a peninsula of Northern Europe that forms the continental portion of Denmark and part of northern Germany (Schleswig-Holstein). It ...
(
Denmark Denmark is a Nordic countries, Nordic country in Northern Europe. It is the metropole and most populous constituent of the Kingdom of Denmark,, . also known as the Danish Realm, a constitutionally unitary state that includes the Autonomous a ...
) and east to Siberia and the
Tian Shan The Tian Shan, also known as the Tengri Tagh or Tengir-Too, meaning the "Mountains of God/Heaven", is a large system of mountain ranges in Central Asia. The highest peak is Jengish Chokusu at high and located in Kyrgyzstan. Its lowest point is ...
.


Subspecies

*''A. a. agestis'' southern and central Europe *''A. a. calida'' Chavignerie Sicily, Italy, Asia Minor *''A. a. azerbaidzhana'' Obraztsov, 1935 Transcaucasia, Caucaus Major *''A. a. nazira'' (Moore, 1865) Darvaz, western Pamirs, north-western Himalayas Brown argus (Aricia agestis) female.jpg, female ''A. a. agestis'', UK Brown argus (Aricia agestis calida) Italy.jpg, female ''A. a. calida'', Italy Aricia agestis, Brown Argus, Minera, North Wales, May 2017 (34680423115).jpg , larva


Description in Seitz

''L. astrarche'' Bgstr. (= ''medon'' Hufn., ''agestis'' Schiff., ''idas'' Gerh. (?), ''nazira'' Moore) (79 k). Above similar to the preceding, L. anteros ">Aricia_anteros.html" ;"title="Aricia anteros">L. anteros deep dark brown, with strongly marked discocellular spot, especially in the male. Typical specimens have this spot black and bear red spots at the outer margin. Underside with numerous ocelli, the hindwing with a pale smear from the apex of the cell to the centre of the outer margin. From Scandinavia to the Sahara and northern India, and from the Canary Isles to the island of Askold in the Pacific, ascending in the mountains up to 10,000 ft. (Doherty). — Besides accidental aberrations in the usual directions of variation, a number of seasonal and geographical forms have been established. If the spring-brood of the Central European form is regarded as name-typical ''astrarche'', ab. ''aestiva'' Stgr. is the name for the second brood, which occurs regularly in the south and more rarely in the north and which connects ''astrarche'' with ''calida'' Bell. (79 k, 80 a) [''A. a. calida'' (Bellier, 1862) . The latter has a much darker, almost coffee-brown, under-side, with a broader band of red spots. Egg pale green, flattened, with the top concave and the surface minutely reticulate (Tutt, Harrison, Gillmer). Larva light green, with a purple dorsal stripe and a similar stripe along the sides, between them oblique pale smears; in spring and summer (in the south throughout the year) on various plants, such as ''Helianthemum'', ''Centaurea'', ''Erodium'', etc. Pupa pale yellowish green, sometimes with a brown dorsal stripe, on or near the ground. The butterflies are on the wing in May and again from July onward, in the south the whole summer until the autumn, in several broods. They fly everywhere, in fields, on field-paths, on rocky hills, even in gardens, always close to the ground and belong to the very commonest butterflies. During the hours of flight they settle with spread wings on blades of grass, blossoms, or clods of earth, while they sleep with closed wings generally hidden under umbels and the inflorescences of grasses.


Appearance and biology (Great Britain)

:''Note that information on this species applies to Great Britain and some details may not be consistent with the species in other parts of its range.'' Although one of the "blues", both sexes are brown on the uppersides with a band of orange spots at the border of each wing. They can be mistaken for other female blues but the brown argus never has any blue scales at the base of the wings like other female blues often do. It could also be mistaken for the northern brown argus (''Aricia artaxerxes'') were it not for the fact that their ranges do not overlap in the UK, unlike on continental Europe. The underside has the typical "blue" pattern of a greyish/brownish ground colour with black spots outlined in white and a row of orange spots along the border. The pattern of the black spots is the best way to distinguish this species from female
common Common may refer to: As an Irish surname, it is anglicised from Irish Gaelic surname Ó Comáin. Places * Common, a townland in County Tyrone, Northern Ireland * Boston Common, a central public park in Boston, Massachusetts * Cambridge Com ...
, chalkhill and
Adonis In Greek mythology, Adonis (; ) was the mortal lover of the goddesses Aphrodite and Persephone. He was considered to be the ideal of male beauty in classical antiquity. The myth goes that Adonis was gored by a wild boar during a hunting trip ...
blues as they lack the black spot found near the base of the forewing which is present on these three species. This species has seen an expansion in its range in recent years and is widely distributed across south-east England and most of
the Midlands The Midlands is the central region of England, to the south of Northern England, to the north of southern England, to the east of Wales, and to the west of the North Sea. The Midlands comprises the ceremonial counties of Derbyshire, Herefords ...
with colonies occurring in
Wales Wales ( ) is a Countries of the United Kingdom, country that is part of the United Kingdom. It is bordered by the Irish Sea to the north and west, England to the England–Wales border, east, the Bristol Channel to the south, and the Celtic ...
and as far north as
Yorkshire Yorkshire ( ) is an area of Northern England which was History of Yorkshire, historically a county. Despite no longer being used for administration, Yorkshire retains a strong regional identity. The county was named after its county town, the ...
. These northern sites have seen a lot of confusion in recent years with genetic studies looking at various colonies to separate the two ''Aricia'' species. Until a few years ago, these northern colonies were thought to be the northern brown argus and more colonies may yet be found to be misidentified. Like other blues it is common on the chalk downlands of southern England but will also use other habitats such as woodland clearings, coastal grasslands and heathland. It is not closely related to the Scotch argus.


Life cycle and food plants

:''Note that information on this species applies to Great Britain and some details may not be consistent with the species in other parts of its range.'' Common rock-rose ('' Helianthemum nummularium'') is the favoured food plant on
calcareous soil Calcareous () is an adjective meaning "mostly or partly composed of calcium carbonate", in other words, containing lime or being chalky. The term is used in a wide variety of scientific disciplines. In zoology ''Calcareous'' is used as an adje ...
s. In other habitats dove's-foot cranesbill ('' Geranium molle'') and common stork's-bill ('' Erodium cicutarium'') are used and possibly other ''
Geranium ''Geranium'' is a genus of 422 species of annual, biennial, and perennial plants that are commonly known as geraniums or cranesbills. They are found throughout the temperate regions of the world and the mountains of the tropics, with the gre ...
'' species as well. Eggs are laid singly on the underside of leaves. The typically
slug Slug, or land slug, is a common name for any apparently shell-less Terrestrial mollusc, terrestrial gastropod mollusc. The word ''slug'' is also often used as part of the common name of any gastropod mollusc that has no shell, a very reduced ...
-like lycid larvae are green with a pale line along each side and always attended by ants. They hibernate as fully-grown larvae and pupate the following spring. There are two broods a year in the southern colonies with adults on the wing in May and June and again in late July till mid-September but further north they are single brooded and fly in June and July.


See also

*
List of butterflies of Great Britain This is a list of butterfly, butterflies of Great Britain, including extinct, naturalised species and those of dubious origin. The list comprises butterfly species listed in ''The Moths and Butterflies of Great Britain and Ireland'' by Emmet ''et a ...


References

{{Taxonbar, from=Q1294181 agestis Butterflies of Europe Butterflies described in 1775