Brotia Distribution
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''Brotia'' is a genus of
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
n
freshwater snail Freshwater snails are gastropod mollusks which live in fresh water. There are many different families. They are found throughout the world in various habitats, ranging from ephemeral pools to the largest lakes, and from small seeps and springs ...
s, gastropod molluscs in the
taxonomic Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical classification) is a scheme of classification, especially a hierarchical classification, in which things are organized into groups or types. A ...
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Pachychilidae Pachychilidae, common name pachychilids, is a taxonomic family of freshwater snails, gastropod molluscs in the clade Sorbeoconcha.Gofas, S. (2014). Pachychilidae. Accessed through: World Register of Marine Species at http://www.marinespecies. ...
.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioidea: Pachychilidae)". ''
Malacologia ''Malacologia'' is a peer-reviewed scientific journal in the field of malacology, the study of mollusks. The journal publishes articles in the fields of molluscan systematics, ecology, population ecology, genetics, molecular genetics, evolution, a ...
'' 48
159
251.
The generic name ''Brotia'' is apparently in honor of a Swiss
malacologist Malacology is the branch of invertebrate zoology that deals with the study of the Mollusca (mollusks or molluscs), the second-largest phylum of animals in terms of described species after the arthropods. Mollusks include snails and slugs, clams, ...
Auguste Louis Brot Auguste Louis Brot (September 18, 1821 ''Sevue Suisse de zoologies'', vol. 4, 1896-189"Auguste Louss Brot"/ref>- August 30, 1896) was a Swiss malacologist (conchologist). ''The American Naturalist'', March 1897p.270/ref> After several years of st ...
(1821-1896).


Distribution

Species of ''Brotia'' occur in
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does in ...
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s of
Southeast Asia Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
, ranging from
Northern India North India is a loosely defined region consisting of the northern part of India. The dominant geographical features of North India are the Indo-Gangetic Plain and the Himalayas, which demarcate the region from the Tibetan Plateau and Central ...
in the west through to Sumatra in the east. This range includes
India India, officially the Republic of India ( Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the ...
,
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
,
Bangladesh Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the List of countries and dependencies by population, eighth-most populous country in the world, with a population exceeding 165 million pe ...
,
Thailand Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is b ...
, Laos,
Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making it ...
, China (1 species),
Cambodia Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailan ...
,
Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...
and
Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania between the Indian and Pacific oceans. It consists of over 17,000 islands, including Sumatra, Java, Sulawesi, and parts of Borneo and New Guine ...
( Sumatra and
Borneo Borneo (; id, Kalimantan) is the List of islands by area, third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java Isl ...
only).


Biology

Species are found predominantly in fast flowing, well oxygenated rivers, occasionally also in lakes. They are
gonochoristic In biology, gonochorism is a sexual system where there are only two sexes and each individual organism is either male or female. The term gonochorism is usually applied in animal species, the vast majority of which are gonochoric. Gonochorism c ...
, and
viviparous Among animals, viviparity is development of the embryo inside the body of the parent. This is opposed to oviparity which is a reproductive mode in which females lay developing eggs that complete their development and hatch externally from the ...
, retaining developing eggs and youngs in special brood pouch.


Species

Species within the genus ''Brotia'' include: * † ''
Brotia alpina ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (F. Sandberger, 1871) * ''
Brotia annamita ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' Köhler, Holford M., Do & Ho, 2009Köhler F., Holford, Do V. T. & Ho T. H. (2009). "Exploring a largely unknown fauna: On the diversity of pachychilid freshwater gastropods in Vietnam (Caenogastropoda: Cerithioidea)". ''
Molluscan Research Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is es ...
'' 29(3): 121-146
abstract
* ''
Brotia angulifera ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' Brot, 1872 - incertae sedis * ''
Brotia armata ''Brotia armata'' is a species of freshwater snail with an operculum, an aquatic gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastrop ...
'' (Brandt, 1874) * '' Brotia assamensis'' (Nevill, 1885) - incertae sedis * ''
Brotia beaumetzi ''Sulcospira tonkiniana'' is a species of freshwater snail with an operculum, an aquatic gastropod mollusk in the family Pachychilidae. Distribution This species occurs in Vietnam.Köhler F. & Dames C. (2009). "Phylogeny and systematics of ...
'' (Brot, 1887) - incertae sedis * ''
Brotia binodosa ''Brotia binodosa'' is a species of freshwater snail with an operculum. It is an aquatic gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called ...
'' (Blanford, 1903) * † ''
Brotia bittneri ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Oppenheim, 1895) * ''
Brotia boenana ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Brot, 1881) * ''
Brotia borneensis ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Schepman, 1896) - incertae sedis * † ''
Brotia castellaunensis ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Boussac, 1911) * ''
Brotia citrina ''Brotia citrina'' is a species of freshwater snail with an operculum, an aquatic gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastr ...
'' (Brot, 1868) * ''
Brotia clavaeformis ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Brot, 1874) * ''
Brotia costula ''Brotia costula'' is a species of gastropod belonging to the family Pachychilidae Pachychilidae, common name pachychilids, is a taxonomic family of freshwater snails, gastropod molluscs in the clade Sorbeoconcha.Gofas, S. (2014). Pachychilida ...
'' (Rafinesque, 1833) * ''
Brotia cylindrus ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Brot, 1886) - incertae sedis * ''
Brotia dautzenbergiana ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Morlet, 1884) * ''
Brotia episcopalis ''Brotia episcopalis'' is a species of freshwater snail with an operculum, an aquatic gastropod mollusk in the family Pachychilidae. Taxonomy In the 20th century, ''B. episcopalis'' was frequently synonymized with ''Brotia costula ''B ...
'' (H. Lea & I. Lea, 1851) * ''
Brotia godwini ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Brot, 1875) * '' Brotia henriettae'' (Griffith & Pidgeon, 1834) * ''
Brotia herculea ''Brotia herculea'', common name the giant tower cap snail, is a species of freshwater snail with an operculum, an aquatic gastropod mollusc in the family Pachychilidae. Description ''B. herculea'' has a taenioglossan radula. Its shell is ...
'' (Gould, 1846) * ''
Brotia hoabinhensis ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' Köhler, Holford, Do & Ho, 2009 * ''
Brotia huberi ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' Thach, 2021 * ''
Brotia indragirica ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (von Martens, 1900) * ''
Brotia insolita ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Brot, 1868) * ''
Brotia iravadica ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Blanford, 1869) * ''
Brotia jullieni ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Deshayes, 1874) * ''
Brotia kelantanesis ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Preston, 1907) * ''
Brotia laodelectata ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' Köhler, 2008 * † ''
Brotia lhazeensis ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' Yü, 1982 * † ''
Brotia lombersensis ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Noulet, 1867) * † ''
Brotia majevitzae ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Oppenheim, 1901) * ''
Brotia manningi ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' Brandt, 1968 * ''
Brotia mariae ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' Köhler, 2008 * ''
Brotia microsculpta ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' Brandt, 1968 * ''
Brotia pageli ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioidea: ...
'' (Thiele, 1908) * ''
Brotia pagodula ''Brotia pagodula'' is a species of freshwater snail with an operculum, an aquatic gastropod mollusk in the family Pachychilidae. Taxonomy ''Brotia pagodula'' is the type species of the genus ''Brotia''.Köhler F., Holford, Do V. T. & Ho T. ...
'' (Gould, 1847) - type species * † ''
Brotia palaeocostula ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' Gurung, Takayasu & Matsuoka, 1997 * ''
Brotia paludiformis ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Solem, 1966) * ''
Brotia peninsularis ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Brandt, 1974) * ''
Brotia persculpta ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Ehrmann, 1922) * ''
Brotia praetermissa ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' Köhler & Glaubrecht, 2002 * '' Brotia pseudoasperata'' Brandt, 1968 * '' Brotia pseudosulcospira'' (Brandt, 1968) * ''
Brotia siamensis ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Brot, 1886) * ''
Brotia solemiana ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Brandt, 1968) * '' Brotia sooloensis'' (Reeve, 1859) - incertae sedis * ''
Brotia spinata ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Godwin-Austen, 1872) - incertae sedis * ''
Brotia subgloriosa ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Brandt, 1968) * ''
Brotia sumatrensis ''Brotia sumatrensis'' is a species of freshwater snail with an operculum, an aquatic gastropod mollusk in the family Pachychilidae. Previously frequently synonymized with ''Brotia costula'', biological systematics was able to molecularly ...
'' (Brot, 1875) * ''
Brotia torquata ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioidea ...
'' (von dem Busch, 1842) * ''
Brotia verbecki ''Brotia verbecki'' is a species of freshwater snail with an operculum, an aquatic gastropod mollusk in the family Pachychilidae. Distribution This species occurs in: * Sumatra Sumatra is one of the Sunda Islands of western Indo ...
'' (Brot, 1886) * ''
Brotia wykoffi ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Brandt, 1974) * ''
Brotia yunnanensis ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioid ...
'' Köhler, Du & Yang, 2010Köhler F., Du L.-N. & Yang J.-X. (2010). "A new species of ''Brotia'' from Yunnan, China (Caenogastropoda, Pachychilidae)". ''
Zoosystematics and Evolution ''Zoosystematics and Evolution'' is a peer-reviewed open access scientific journal covering zoological systematics and evolution. It was established in 1898 as ''Mitteilungen aus der Zoologischen Sammlung des Museums für Naturkunde in Berlin'' () ...
'' 86(2): 295-300. .
* ''
Brotia zonata ''Brotia'' is a genus of Southeast Asian freshwater snails, gastropod molluscs in the taxonomic family Pachychilidae.Köhler F. & Glaubrecht M. (2006). "A systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' (Cerithioide ...
'' (Benson, 1836) - incertae sedis Synonyms: * ''Brotia asperata'' (Lamarck, 1822): synonym of ''
Jagora asperata ''Jagora asperata'' is a species of freshwater snail with an operculum, an aquatic gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastr ...
'' (Lamarck, 1822) * ''Brotia baccata'' (Gould, 1847): synonym of '' Brotia henriettae'' (Gray, 1834) * ''Brotia delavayana'' Heude, 1888 : synonym of ''Sulcospira delavayana'' (Heude, 1888) * † ''Brotia escheri aquitanica'' (Noulet, 1846): synonym of † ''Tinnyea aquitanica'' (Noulet, 1846) * † ''Brotia escheri auingeri'' (Handmann, 1882): synonym of † ''Tinnyea lauraea'' (Matheron, 1842) * ''Brotia hainanensis'' (Brot, 1872): synonym of ''Sulcospira hainanensis'' (Brot, 1874) * ''Brotia hamonvillei'' : synonym of ''Sulcospira tonkiniana'' (Morelet, 1886) * ''Brotia infracostata'' (Mousson, 1848): synonym of ''Sulcospira infracostata'' (Mousson, 1848) * ''Brotia proteus'' (Bavay & Dautzenberg, 1908): synonym of ''Sulcospira dakrongensis'' Köhler, Holford, Do & Ho, 2009 * ''Brotia swinhoei'' (H. Adams, 1870): synonym of ''Sulcospira swinhoei'' (H. Adams, 1870) * ''Brotia testudinaria'' (von dem Busch, 1842):Köhler F. & Glaubrecht M. (2001). "Toward a systematic revision of the Southeast Asian freshwater gastropod ''Brotia'' H. – Adams, 1866 (Cerithioidea: Pachychilidae): an account of species from around the South China Sea". ''Journal of Molluscan Studies'' 67(3): 281-318. synonym of ''Sulcospira testudinaria'' (von dem Busch, 1842) * ''Brotia tourannensis'': synonym of ''Sulcospira tourannensis'' (Souleyet, 1852) * ''Brotia variabilis'' (Benson, 1836): synonym of ''
Brotia costula ''Brotia costula'' is a species of gastropod belonging to the family Pachychilidae Pachychilidae, common name pachychilids, is a taxonomic family of freshwater snails, gastropod molluscs in the clade Sorbeoconcha.Gofas, S. (2014). Pachychilida ...
'' (Rafinesque, 1833) Variability of shells of ''Brotia'' (images in the first row are in the same ratio): File:Brotia armata shell.png, ''
Brotia armata ''Brotia armata'' is a species of freshwater snail with an operculum, an aquatic gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastrop ...
'' File:Brotia binodosa shell.png, ''
Brotia binodosa ''Brotia binodosa'' is a species of freshwater snail with an operculum. It is an aquatic gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called ...
'' File:Brotia herculea shell.png, ''
Brotia herculea ''Brotia herculea'', common name the giant tower cap snail, is a species of freshwater snail with an operculum, an aquatic gastropod mollusc in the family Pachychilidae. Description ''B. herculea'' has a taenioglossan radula. Its shell is ...
''


References


External links

* Köhler F. & Deein G. (2010). "Hybridisation as potential source of incongruence in the morphological and mitochondrial diversity of a Thai freshwater gastropod (Pachychilidae, ''Brotia'' H. Adarns, 1866)". ''
Zoosystematics and Evolution ''Zoosystematics and Evolution'' is a peer-reviewed open access scientific journal covering zoological systematics and evolution. It was established in 1898 as ''Mitteilungen aus der Zoologischen Sammlung des Museums für Naturkunde in Berlin'' () ...
'' 86(2): 301-314. . {{Taxonbar, from=Q990852 Gastropod genera Brotia, Brotia Freshwater snails