''Brittagnathus'' is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of four-limbed

vertebrate Vertebrates () are animals with a vertebral column (backbone or spine), and a cranium, or skull. The vertebral column surrounds and protects the spinal cord, while the cranium protects the brain. The vertebrates make up the subphylum Vertebra ...

tetrapod A tetrapod (; from Ancient Greek :wiktionary:τετρα-#Ancient Greek, τετρα- ''(tetra-)'' 'four' and :wiktionary:πούς#Ancient Greek, πούς ''(poús)'' 'foot') is any four-Limb (anatomy), limbed vertebrate animal of the clade Tetr ...

") from the

Late Devonian The Devonian ( ) is a geologic period and system of the Paleozoic era during the Phanerozoic eon, spanning 60.3 million years from the end of the preceding Silurian period at million years ago ( Ma), to the beginning of the succeeding ...

Greenland Greenland is an autonomous territory in the Danish Realm, Kingdom of Denmark. It is by far the largest geographically of three constituent parts of the kingdom; the other two are metropolitan Denmark and the Faroe Islands. Citizens of Greenlan ...

. It contains a single species, ''Brittagnathus minutus'', which is based on a complete lower jaw recovered from an ''

Acanthostega ''Acanthostega'', from Ancient Greek ἄκανθα (''ákantha''), meaning "spine", and στέγη (''stégē''), meaning "roof", is an extinct genus of stem tetrapoda, stem-tetrapod, among the first vertebrates, vertebrate animals to have recogn ...

bonebed A bone bed is any geological stratum or deposit that contains bones of whatever kind. Inevitably, such deposits are sedimentary in nature. Not a formal term, it tends to be used more to describe especially dense collections such as Lagerstätte. ...

in the Britta Dal Formation. It is the fourth named genus of "tetrapod" (more precisely a

stem-tetrapod The Stem Tetrapoda are a cladistically defined group, consisting of all animals more closely related to extant four-legged vertebrates than to their closest extant relatives (the lungfish), but excluding the crown group Tetrapoda. They are thus ...

or stegocephalian) from the Late Devonian of Greenland, after ''

Ichthyostega ''Ichthyostega'', from Ancient Greek ἰχθύς (''ikthús''), meaning "fish", and στέγη (''stégē''), meaning "roof", is an Extinction, extinct genus of limbed tetrapodomorpha, tetrapodomorphs from the Devonian, Late Devonian of what is ...

'', ''Acanthostega'', and '' Ymeria''. The jaw is only 4.5 cm (1.8 in.) long, with the total length estimated at 25 cm, making ''Brittagnathus'' the smallest known Devonian "tetrapod". A

phylogenetic analysis In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical data ...

places it among

Carboniferous The Carboniferous ( ) is a Geologic time scale, geologic period and System (stratigraphy), system of the Paleozoic era (geology), era that spans 60 million years, from the end of the Devonian Period Ma (million years ago) to the beginning of the ...

stegocephalians, particularly the small whatcheeriid ''

Pederpes ''Pederpes'' ("Peter's Foot") is an extinct genus of early Carboniferous stegocephalian, dating from 348 to 347.6 Ma in the Tournaisian age (lower Mississippian). ''Pederpes'' contains one species, ''P. finneyae'', 1 m long. This most basal C ...

'', rather than the contemporary Devonian stem-tetrapods. This provides support for an origin for Carboniferous-type tetrapods as early as the Devonian.

References

Stegocephali Monotypic prehistoric lobe-finned fish genera Late Devonian sarcopterygians Devonian sarcopterygians of North America Devonian Greenland Fossils of Greenland Fossil taxa described in 2020 Taxa named by Jenny Clack {{Paleo-tetrapodomorph-stub