Brent D. Mishler (born 1953) is an American
botanist
Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...
who from was the director of the
University and Jepson Herbaria at the
University of California, Berkeley
The University of California, Berkeley (UC Berkeley, Berkeley, Cal, or California), is a Public university, public Land-grant university, land-grant research university in Berkeley, California, United States. Founded in 1868 and named after t ...
, from 1993 to 2023. He was also a professor in the Department of Integrative Biology, where he taught phylogenetics, plant diversity, and island biology. Following his retirement in 2024 he is now Distinguished Professor Emeritus and Curator of Bryophytes, and continues an active research program.
Early life and education
Mishler attended Bonita High School in La Verne, California, and worked as a Ranger-Naturalist for the Los Angeles County Nature Center Unit (Parks and Recreation Department). He received his B.S. and M.S. from
California State Polytechnic University, Pomona
California State Polytechnic University Pomona (Cal Poly Pomona) is a Public university, public Institute of Technology (United States)#Polytechnic universities, polytechnic research university in Pomona, California, United States. It is the l ...
, in 1975 and 1978 respectively, then got his Ph.D. from
Harvard University
Harvard University is a Private university, private Ivy League research university in Cambridge, Massachusetts, United States. Founded in 1636 and named for its first benefactor, the History of the Puritans in North America, Puritan clergyma ...
in 1984, working with Carroll E. Wood,
Peter F. Stevens and
Norton G. Miller He did his dissertation work on the systematics and evolution of the moss genus ''
Syntrichia''.
Career
Research
Mishler was on the faculty of the Botany Department at
Duke University
Duke University is a Private university, private research university in Durham, North Carolina, United States. Founded by Methodists and Quakers in the present-day city of Trinity, North Carolina, Trinity in 1838, the school moved to Durham in 1 ...
from 1984. He moved to the Department of Integrative Biology at
University of California, Berkeley
The University of California, Berkeley (UC Berkeley, Berkeley, Cal, or California), is a Public university, public Land-grant university, land-grant research university in Berkeley, California, United States. Founded in 1868 and named after t ...
in 1993.He has maintained two main areas of research: empirical studies of ecology, phylogeny, systematics, and development of mosses, and investigations into the theoretical basis of systematic and evolutionary biology. He has published over 200
scientific papers.

Most recently, Mishler has helped to develop spatial phylogenetic tools for studying
biodiversity
Biodiversity is the variability of life, life on Earth. It can be measured on various levels. There is for example genetic variability, species diversity, ecosystem diversity and Phylogenetics, phylogenetic diversity. Diversity is not distribut ...
and
endemism
Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
, using large-scale phylogenies and collection data in a geographic and statistical framework.
Species concept
Mishler has had a particular interest in
concepts of species over his 40-year career, in relation to theories of biological classification in general. He realized early on that the evolutionary processes operating in different branches of the
tree of life
The tree of life is a fundamental archetype in many of the world's mythology, mythological, religion, religious, and philosophy, philosophical traditions. It is closely related to the concept of the sacred tree.Giovino, Mariana (2007). ''The ...
are distinct enough that different criteria need to be applied to decide when lineages are diverged enough to be called species. He later recognized that deciding which among the many nested levels in the vast tree of life should be ranked as a species is arbitrary and misleading, and began to advocate a rankless approach as was already being advocated for higher taxonomic levels by advocates of phylogenetic nomenclature in the
PhyloCode
The ''International Code of Phylogenetic Nomenclature'', known as the ''PhyloCode'' for short, is a formal set of rules governing phylogenetic nomenclature. Its current version is specifically designed to regulate the naming of clades, leaving the ...
naming system. Mishler argues that the species rank can and should be done away with, to be replaced by a multi-level approach to systematics, ecology, evolution, and conservation.
His two recent books
[Swartz, B. and B.D. Mishler (eds.). 2022 ''Speciesism in Biology and Culture: How Human Exceptionalism is Pushing Planetary Boundaries''. Springer Nature. ] explore this radical view of "species", and what it means for science and society to move to a rankless, phylogenetic view of biodiversity.
Quentin Wheeler and Rudolf Meier note that the Mishler version of the phylogenetic species concept differs from that of Wheeler and
Norman Platnick
Norman Ira Platnick (December 30, 1951 – April 8, 2020) was an American biological systematist and arachnologist. At the time of his death, he was a professor emeritus of the Richard Gilder Graduate School and Peter J. Solomon Family Curator E ...
, and that biologists like
Ernst Mayr
Ernst Walter Mayr ( ; ; 5 July 1904 – 3 February 2005) was a German-American evolutionary biologist. He was also a renowned Taxonomy (biology), taxonomist, tropical explorer, ornithologist, Philosophy of biology, philosopher of biology, and ...
disagree with both versions.
Reception
Werner Kunz, noting that
Charles Darwin
Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English Natural history#Before 1900, naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all speci ...
"did not actually believe in the existence of species", describes Mishler as having an "extreme attitude ... related to denying any existence of biological species", shared with the botanist Konrad Bachmann, that the idea of a biological species is "worn out, hopelessly vague, or even evidently wrong".
The evolutionary biologist
James Mallet writes that some taxonomists, including Mishler, "even argue that named
Linnean ranks, including species, are no longer useful in taxonomy at all".
The philosopher Marco Nathan distinguishes two kinds of pluralism, heterogeneity and theory-dependence.
He defines heterogeneity as meaning that species are defined differently according to their attributes, e.g. Mayr's "interbreeding natural populations ... reproductively isolated" applies only to organisms that reproduce sexually, so asexual groups like bacteria need a different definition of species. Nathan defines theory-dependence as meaning that taxa are assigned according to the organisms being discussed and "the explanatory target at hand", so there is no unique or "natural" method for defining a species or taxa at any other level. Nathan calls the "common" labelling of heterogeneity as pluralism by biologists such as Mishler "dangerous" as it overlooks the distinction of the two kinds.
References
Primary
''These sources are used for basic information about the subject's life and work.''
Secondary
{{DEFAULTSORT:Mishler, Brent
American botanists
University of California, Berkeley faculty
1953 births
Living people
California State Polytechnic University, Pomona alumni
Harvard Graduate School of Arts and Sciences alumni
Duke University faculty