Brachiaria Deflexa
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''Urochloa'', commonly known as signalgrass,''Urochloa''.
USDA PLANTS.
is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of plants in the
grass family Poaceae ( ), also called Gramineae ( ), is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos, the grasses of natural grassland and species cultivated in ...
, native to
tropical The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where the Sun can never be directly overhead. This is because of Earth's ax ...
and
subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones immediately to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Ge ...
regions of
Eurasia Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...
, Africa, Australia, the
Americas The Americas, sometimes collectively called America, are a landmass comprising the totality of North America and South America.''Webster's New World College Dictionary'', 2010 by Wiley Publishing, Inc., Cleveland, Ohio. When viewed as a sing ...
, and various islands.''Urochloa''.
Grass Manual. Flora of North America.
Watson, L. and M. J. Dallwitz

The Grass Genera of the World. DELTA – DEscription Language for TAxonomy.
Several species of the genus ''Urochloa'' are cultivated as
forage Forage is a plant material (mainly plant leaves and stems) eaten by grazing livestock. Historically, the term ''forage'' has meant only plants eaten by the animals directly as pasture, crop residue, or immature cereal crops, but it is also used m ...
and some species of were probably first introduced unintentionally to the Americas in the colonial period, from slave ships. ''Urochloa eminii'' was introduced to Brazil in 1952. ''Urochloa'' is the most widely used tropical grass in Central and South America, with about 40 million hectares planted in Brazil alone.


Biology

This genus was described in 1812. It is similar to the genus ''
Panicum ''Panicum'' (panicgrass) is a large genus of about 250 species of Poaceae, grasses native throughout the tropical regions of the world, with a few species extending into the northern temperate zone. They are often large, Annual plant, annual or P ...
'', and some authors believe ''Panicum'' is ancestral to it. A
phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
analysis concluded that species of the former genus ''Brachiaria'' formed a
monophyletic group In biology, a clade (), also known as a monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach to taxonomy ...
with those of ''Urochloa'', along with the genera ''
Eriochloa ''Eriochloa'' is a widespread genus of plants in the grass family, commonly called cupgrass. They are found across much of Africa, Asia, Australia, and the Americas, plus a few places in European Russia. ; Species * ''Eriochloa acuminata'' &ndas ...
'' and ''
Melinis ''Melinis'' is a genus of African and Arabian plants in the Poaceae, grass family. The generic name is derived from the Greek ''meline'' meaning "millet". Species 23 species are currently accepted: ; formerly included numerous species now reg ...
'', and that further molecular and morphological work is needed to establish clear relationships. ''Urochloa'' species are annual or perennial grasses, most lacking
rhizome In botany and dendrology, a rhizome ( ) is a modified subterranean plant stem that sends out roots and Shoot (botany), shoots from its Node (botany), nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from ...
s. The
inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
is a branching
panicle In botany, a panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a p ...
, and the plant reaches about a meter in height.Watson, L. and M. J. Dallwitz. (2008)
''Brachiaria''.
The Grass Genera of the World. Retrieved 7 November 2011.
The plants are bisexual and the flowers are fleshy, with 3
anthers The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filamen ...
.Clayton, W. D., et al. (2002 onwards)
''Brachiaria''.
GrassBase - The Online World Grass Flora.
Some species have a prominent vein in the center of the leaf. ''Brachiaria'' are C4 species and can tolerate drier conditions and more light exposure than some other plants.


Ecology and conservation

''Urochloa'' species can grow in many environments, from swamps to shady forest to semidesert, but generally do best in
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...
s and other open tropical ecosystems such as in East Africa. In Angola, ''Urochloa brizantha'' grows on termite mounds and in the
ecotone An ecotone is a transitional area between two plant communities, where these meet and integrate. Examples include areas between grassland and forest, estuaries and lagoon, freshwater and sea water etc. An ecotone may be narrow or wide, and it ma ...
between grassland and woodland habitat.Estes, R. D. and R. K. Estes. (1974). The biology and conservation of the giant sable antelope, ''Hippotragus niger variani'' Thomas, 1916. ''Proceedings of the Academy of Natural Sciences of Philadelphia'' 126, 73-104. In the
Kora National Reserve Kora may refer to: Places India * Kora, Bardhaman, West Bengal * Kora, Bharuch, Gujarat * Korha, Katihar, also known as Kora, in Bihar * Kora, Kendrapara, Odisha * Kora, Wardha, Maharastra * Kora, Tumakuru, Karnataka * Toyaguda, Adilabad, T ...
in Kenya, ''Urochloa'' species dominate the ground layer along with those of the genus ''
Aristida ''Aristida'' is a very nearly Cosmopolitan distribution, cosmopolitan genus of plants in the Poaceae, grass family. ''Aristida'' is distinguished by having three Awn (Botany), awns (bristles) on each lemma (botany), lemma of each floret. The gen ...
''. In India, the native ''Urochloa'' ''ramosa'' is an important food source for the
Eurasian collared dove The Eurasian collared dove (''Streptopelia decaocto''), often simply just collared dove, is a dove species native to Europe, Asia, and northern Africa. It has also been introduced to Japan, North and Central America, and the islands in the Cari ...
and ''Urochloa'' species are forage for other local herbivores. In North America, the native ''Urochloa platyphylla (''broadleaf signalgrass), grows after heavy rains and then reproduces prodigiously and quickly, sometimes becoming a
weed A weed is a plant considered undesirable in a particular situation, growing where it conflicts with human preferences, needs, or goals.Harlan, J. R., & deWet, J. M. (1965). Some thoughts about weeds. ''Economic botany'', ''19''(1), 16-24. Pla ...
. Wide expanses of the tropics, especially the
Neotropics The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In biogeog ...
, have been converted to pastures of ''Urochloa'' species to support
livestock Livestock are the Domestication, domesticated animals that are raised in an Agriculture, agricultural setting to provide labour and produce diversified products for consumption such as meat, Egg as food, eggs, milk, fur, leather, and wool. The t ...
. In Brazil, 80 million hectares of native habitat have been planted with African grasses, mostly ''Urochloa''.
Introduced species An introduced species, alien species, exotic species, adventive species, immigrant species, foreign species, non-indigenous species, or non-native species is a species living outside its native distributional range, but which has arrived ther ...
such as ''Urochloa'' grasses can degrade habitat and compete with native species. In
Northern Australia The unofficial geographic term Northern Australia includes those parts of Queensland and Western Australia north of latitude 26th parallel south, 26° and all of the Northern Territory. Those local government areas of Western Australia and Q ...
, the exotic ''Urochloa eminii'' competes with the native tree '' Alphitonia petriei'' by inhibiting the growth of seedlings, slowing the conversion of abandoned pastureland to natural forest. In the
Paragominas Paragominas is a municipality in the state of Pará in the Northern region of Brazil. Paragominas mine, one of the largest bauxite mines in the world, is approximately 70 km away. According to an estimate dated 1 July 2020, the city has a p ...
area of Brazil, however, native forest outcompetes cultivated stands of ''Urochloa ''and other exotic forage grasses, and ranchers struggle to maintain pasture cover. Native species may also utilize exotic ''Urochloa ''as a food resource, such as the
rock cavy The rock cavy or ''mocó'' (''Kerodon rupestris'') is a cavy species endemic to eastern Brazil which has also been introduced to the Atlantic island of Fernando de Noronha. The rock cavy is called ''mocó'' in Brazilian Portuguese, and ''koriko ...
, a native rodent of the
caatinga Caatinga () is a type of semi-arid tropical vegetation, and an ecoregion characterized by this vegetation in interior northeastern Brazil. The name "Caatinga" comes from the Tupi word '' ka'atinga'', meaning "white forest" or "white vegetat ...
.


Cultivation

''Urochloa ''is the single most important genus of forage grass for pastures in the tropics. ''Urochloa ''cultivars can grow in infertile and acidic soils. Brazil is the leading user and producer of ''Urochloa ''seeds in the Americas. Mexico has put effort into improving its trade in ''Urochloa ''cultivars, and the grass is thought to have made a positive impact on its milk and beef industries.Holmann, F., et al. (2004)
Impact of the adoption of ''Brachiaria'' grasses: Central America and Mexico.
''Livestock Research for Rural Development'' 16, Art. #98.
Central America Central America is a subregion of North America. Its political boundaries are defined as bordering Mexico to the north, Colombia to the southeast, the Caribbean to the east, and the Pacific Ocean to the southwest. Central America is usually ...
n countries have also increased seed sales and area planted in the grass. The annual growth rate of seed sales in 2009 was 32% in Mexico, 62% in Honduras, 45% in Nicaragua, 39% in Costa Rica, and 54% in Panama. The area planted with ''Urochloa ''during this period was about 6.5% of the total surface of permanent grasses in Mexico, 12.5% in Honduras, 1% in Nicaragua, 18.7% in Costa Rica, and 0.1% in Panama. Agricultural pests of ''Urochloa ''include
spittlebugs Cercopidae are the largest family of Cercopoidea, a xylem-feeding insect group, commonly called froghoppers. They belong to the hemipteran suborder Auchenorrhyncha. A 2023 phylogenetic study of the family suggested the elevation of the New Worl ...
,
leafcutter ant Leafcutter ants are fungus-growing ants that share the behaviour of cutting leaves which they carry back to their nests to farm fungus. Next to humans, leafcutter ants form some of the largest and most complex animal societies on Earth. In a few ...
s, and
mound-building termites Mound-building termites are a group of termite species that live in mounds which are made of a combination of soil, termite saliva and dung. These termites live in Africa, Australia and South America. The mounds sometimes have a diameter of ...
. Other insect pests include: *shoot flies ''
Atherigona oryzae ''Atherigona oryzae'', the rice shoot fly, is a species of fly in the family Muscidae Muscidae are a family of flies found in the superfamily Muscoidea. Muscidae, some of which are commonly known as house flies or stable flies due to their ...
'', ''
Atherigona pulla ''Atherigona pulla'', the proso millet shoot fly, is a species of fly in the family Muscidae. The larvae feed on the central growing shoots of crops such as proso millet and little millet. It is found in South Asia.Ravulapenta Sathish, M Manjuna ...
'', and ''
Atherigona punctata ''Atherigona punctata'', the Coimbatore wheat stem fly, is a species of fly in the family Muscidae. In South India, it is a pest of the wheat plant, ''Triticum aestivum Common wheat (''Triticum aestivum''), also known as bread wheat, is a cu ...
'' *caseworm '' Paraponyx stagnalis'' *red hairy caterpillars '' Amsacta albistriga'' and '' Amsacta moorei''


Species

As of 2024, more than a hundred species have been accepted in the genus ''Urochloa'': *''
Urochloa adspersa ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa advena ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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'' *''
Urochloa albicoma ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa ambigens ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa argentea ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa arida ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa arizonica ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa arrecta ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa atrisola ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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'' *''
Urochloa bovonei ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa brachyura'' - eastern and southern Africa *''
Urochloa breviglumis ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa brevispicata ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa brizantha'' - tropical and southern Africa *''
Urochloa burmanica ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa caboverdiana ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa chusqueoides ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa ciliatissima'' *''
Urochloa clavipila ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa comata ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa comorense ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa deflexa ''Urochloa deflexa'', commonly known as Guinea millet, is an Annual plant, annual millet grass belonging to the grass family (Poaceae). It is native to many regions such as Africa, India, and Pakistan in both tropical and subtropical regions. It ...
'' *'' Urochloa dictyoneura'' *''
Urochloa distachyoides ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa distachyos ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa dura ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa echinolaenoides'' - Democratic Republic of the Congo, Tanzania, Malawi, Zambia *''
Urochloa eminii ''Urochloa eminii'', commonly known as Congo grass, is a species of forage crop in the family Poaceae Poaceae ( ), also called Gramineae ( ), is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as g ...
'' *''
Urochloa falcifera ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa foliosa ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa fusca ''Urochloa fusca'', the browntop signalgrass, is a wild grass species with a native range extending from Paraguay in South America to the southern United States (Alabama, Arizona, Florida, Georgia, New Mexico, Texas, and Oklahoma), and it is now ...
'' (browntop signalgrass) - southern USA to Argentina *''
Urochloa fusiformis ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa gilesii ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa glomerata ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa glumaris'' - SE Asia,
Indian subcontinent The Indian subcontinent is a physiographic region of Asia below the Himalayas which projects into the Indian Ocean between the Bay of Bengal to the east and the Arabian Sea to the west. It is now divided between Bangladesh, India, and Pakista ...
*''
Urochloa grossa ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa holosericea ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa humbertiana ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa jaliscana ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa jubata ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa kurzii ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa lachnantha ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa lata ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa leersioides ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa lindiensis ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa longiflora ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa lorentziana ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa marlothii ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa megastachya ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa meziana ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa mollis ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa multiculma ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa mutica ''Urochloa mutica'', commonly known as para grass, buffalo grass, Mauritius signal grass, pasto pare, malojilla, gramalote, parana, Carib grass, and Scotch grass, is a species of grass.Stone, Katharine R. 2010''Urochloa mutica''.In: Fire Effects ...
'' *''
Urochloa nana ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa nigropedata ''Urochloa nigropedata'', commonly known as spotted brachiaria, is a perennial grass belonging to the grass family (Poaceae). It is native to Southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed upon ...
'' *''
Urochloa notochthona ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa oblita ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa occidentalis ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa oligobrachiata ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa oligotricha'' - Africa *'' Urochloa olivacea'' - western Mexico *''
Urochloa ophryodes ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa orthostachys ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa ovalis ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa panicoides ''Urochloa panicoides'' is a fodder grass originating in Southern Africa. Description This annual grass has decumbent or upright stems up to a meter long. It may root where its lower nodes contact the substrate. The leaves have linear or lance- ...
'' (panic liverseed grass) - Africa, southern Asia *'' Urochloa pauciflora'' - western Mexico *''
Urochloa paucispicata ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa piligera ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa plantaginea ''Urochloa plantaginea'' or ''Brachiaria plantaginea'' is an annual grass. Its leaf blades are thin and broadly linear. It has rooting at the lower nodes. The ascending ends and flowering branches are normally 8-12 in (20-30 cm) high; the plant a ...
'' *''
Urochloa platynota ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa platyphylla ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa platyrrhachis'' - Zambia, Democratic Republic of the Congo *''
Urochloa polyphylla ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa polystachya ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa praetervisa ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa psammophila ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa pseudodichotoma ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa pubigera ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa ramosa'' *''
Urochloa reptans ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa reticulata ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa rudis'' - Somalia, Kenya, Tanzania *''
Urochloa rugulosa ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa sclerochlaena'' - Ethiopia, Kenya *''
Urochloa semiundulata ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa serrata ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa serrifolia ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa setigera'' - Indian subcontinent, southern China, SE. Asia *''
Urochloa stigmatisata ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa subulifolia ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa tanimbarensis ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa texana ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa trichopodioides'' - Democratic Republic of the Congo, Tanzania, Kenya, Ethiopia *'' Urochloa trichopus'' - Africa,
Arabian Peninsula The Arabian Peninsula (, , or , , ) or Arabia, is a peninsula in West Asia, situated north-east of Africa on the Arabian plate. At , comparable in size to India, the Arabian Peninsula is the largest peninsula in the world. Geographically, the ...
*''
Urochloa turbinata ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa umbellata ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa umbratilis ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa uzondoiensis ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa villosa ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa whiteana ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa wittei ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Urochloa xantholeuca ''Urochloa'', commonly known as signalgrass,''Urochloa''.
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Kew World Checklist of Selected Plant Families
/ref> such as ''
Alloteropsis ''Alloteropsis'' (from the Greek ''allotrios'' ("strange") and ''opsis'' ("appearance")) is a genus of Old World plants in the grass family.
'', ''
Ixophorus ''Ixophorus'' is a genus of Latin American plants in the grass family. The only recognized species is ''Ixophorus unisetus''.Hiser, K. M''Ixophorus'' Schltdl. Grass Manual. Flora of North America.Kellogg, E. A., et al. (2004)Taxonomy, phylogeny, ...
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Oplismenus ''Oplismenus'' is a small genus of annual or perennial grasses, commonly known as basketgrass, found throughout the tropics, subtropics, and in some cases, temperate regions of the Americas, Africa, Asia, and Australia. The systematics of the ge ...
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Panicum ''Panicum'' (panicgrass) is a large genus of about 250 species of Poaceae, grasses native throughout the tropical regions of the world, with a few species extending into the northern temperate zone. They are often large, Annual plant, annual or P ...
'', and '' Rupichloa''.


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{{Authority control Panicoideae Poaceae genera Grasses of Africa Grasses of Asia Grasses of Europe Grasses of North America Grasses of Oceania Grasses of South America Taxa named by Palisot de Beauvois Plants described in 1812 Cereals