''Bothragonus'' is a

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of marine

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...

es belonging to the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Agonidae Agonidae is a family of small, bottom-dwelling, cold-water marine fish. Common names for members of this family include poachers, Irish lords, sea ravens, alligatorfishes, starsnouts, hooknoses, and rockheads. They are notable for having elongate ...

, the poachers and related fishes. It is the only genus in

monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...

Bothragoninae. These fishes are found in the northern

Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five Borders of the oceans, oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean, or, depending on the definition, to Antarctica in the south, and is ...

Taxonomy

''Bothragonus'' was first proposed as a genus by the American

zoologist Zoology ( , ) is the scientific study of animals. Its studies include the structure, embryology, classification, habits, and distribution of all animals, both living and extinct, and how they interact with their ecosystems. Zoology is one ...

Theodore Gill Theodore Nicholas Gill (March 21, 1837 – September 25, 1914) was an American ichthyologist, mammalogist, malacologist, and librarian. Career Born and educated in New York City under private tutors, Gill early showed interest in natural hist ...

in 1883 with ''Hypsagonus swanii'', which had been described in 1876 by

Franz Steindachner Franz Steindachner (11 November 1834 in Vienna – 10 December 1919 in Vienna) was an Austrian zoologist, ichthyologist, and herpetologist. He published over 200 papers on fishes and over 50 papers on reptiles and amphibians. Steindachner des ...

from

Port Townsend Port Townsend is a city on the Quimper Peninsula in Jefferson County, Washington, United States. The population was 10,148 at the 2020 United States Census. It is the county seat and only incorporated city of Jefferson County. In addition ...

Puget Sound Puget Sound ( ; ) is a complex estuary, estuarine system of interconnected Marine habitat, marine waterways and basins located on the northwest coast of the U.S. state of Washington (state), Washington. As a part of the Salish Sea, the sound ...

Washington Washington most commonly refers to: * George Washington (1732–1799), the first president of the United States * Washington (state), a state in the Pacific Northwest of the United States * Washington, D.C., the capital of the United States ** A ...

, as its

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

. In 1971 the Soviet ichthyologist Georgii Ustinovich Lindberg proposed that the genus Bothragonus be placed in the new subfamily of the Agonidae, the Bothragoninae, although this was not universally accepted. The 5th edition of ''

Fishes of the World ''Fishes of the World'' is a standard reference for the systematics of fishes. It was first written in 1976 by the American ichthyologist Joseph S. Nelson (1937–2011). Now in its fifth edition (2016), the work is a comprehensive overview of t ...

'' recognises the validity of the Bothragoninae as a subfamily of the Agonidae.

Species

There are currently two recognized species in this genus: * ''

Bothragonus occidentalis ''Bothragonus occidentalis'' is a fish in the family Agonidae.''Bothragonus oc ...

Lindberg Lindberg is a municipality in the district of Regen in Bavaria in Germany in the immediate neighbourhood of the larger town Zwiesel. Location Lindberg lies in the Danube Forest (''Donau-Wald'') region in the middle of the Bavarian Forest on ...

, 1935 * '' Bothragonus swanii'' (

Steindachner Franz Steindachner (11 November 1834 in Vienna – 10 December 1919 in Vienna) was an Austrian zoologist, ichthyologist, and herpetologist. He published over 200 papers on fishes and over 50 papers on reptiles and amphibians. Steindachner descri ...

, 1876) (Rockhead)

Etymology

''Bothragonus'' is a combination of ''bothra'', meaning "cavity", a reference to the deep pit on the nape, and '' Agonus'' the

type genus In biological taxonomy, the type genus (''genus typica'') is the genus which defines a biological family and the root of the family name. Zoological nomenclature According to the International Code of Zoological Nomenclature, "The name-bearin ...

of the Agonidae. 2

Characteritics

''Bothragonus'' is characterised by the head and the front part of the body having a rhomboid cross section with the rear part of the body being laterally compressed. The first

dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...

and the

anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...

are poorly developed. There is a row of bony plates located between the upper and main

lateral line The lateral line, also called the lateral line organ (LLO), is a system of sensory organs found in fish, used to detect movement, vibration, and pressure gradients in the surrounding water. The sensory ability is achieved via modified epithelia ...

s and another row is located between the two lower lateral lines. There is an enlarged, spineless plate on the snout and the lower side of the head is armoured with a number of sharply spined plates. The sensory canal on the occipital is broken at its middle. The urohyal has no pelvic keel, there is a postpelvic spine and two epurals. These fishes have a maximum published

total length Fish measurement is the measuring of individual fish and various parts of their anatomies, for data used in many areas of ichthyology, including taxonomy and fishery biology. Overall length Standard length (SL) is the length of a fish measured f ...

of .

Distribution and habitat

''Bothragonus'' are found in the eastern and western North Pacific Ocean. They are found in shallow waters of less than depth and may be found in tidal pools.

References

{{Taxonbar, from=Q149138, from2=Q61882768 Agonidae Taxa named by Theodore Gill