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The great skua (''Stercorarius skua''), sometimes known by the name bonxie in Britain, is a large
seabird Seabirds (also known as marine birds) are birds that are adaptation, adapted to life within the marine ecosystem, marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent ...
in the skua family Stercorariidae. It is roughly the size of a
herring gull Herring gull is a common name for several birds in the genus ''Larus'', all formerly treated as a single species. Three species are still combined in some taxonomies: * American herring gull (''Larus smithsonianus'') - North America * European h ...
. It mainly eats fish caught at the sea surface or taken from other birds.


Taxonomy

The great skua was described from the
Faroe Islands The Faroe Islands ( ) (alt. the Faroes) are an archipelago in the North Atlantic Ocean and an autonomous territory of the Danish Realm, Kingdom of Denmark. Located between Iceland, Norway, and the United Kingdom, the islands have a populat ...
and
Iceland Iceland is a Nordic countries, Nordic island country between the Atlantic Ocean, North Atlantic and Arctic Oceans, on the Mid-Atlantic Ridge between North America and Europe. It is culturally and politically linked with Europe and is the regi ...
by the Danish zoologist
Morten Thrane Brünnich Morten Thrane Brünnich (30 September 1737 – 19 September 1827) was a Danish zoologist and mineralogist. Biography Brünnich was born in Copenhagen, the son of a portrait painter. He studied oriental languages and theology, but soon became i ...
in 1764 under the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Catharacta skua''. It is now placed in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
'' Stercorarius'' that was introduced by the French zoologist
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosophy, natural philosopher. Brisson was born on 30 April 1723 at Fontenay-le-Comte in the Vendée department of western France. Note that page 14 ...
in 1760. The English name and species name "skua" is believed to originate from the Faroese ''skúvur'' or ''skúgvur'' and is the only known bird name to originate from the
Faroes The Faroe Islands ( ) (alt. the Faroes) are an archipelago in the North Atlantic Ocean and an autonomous territory of the Danish Realm, Kingdom of Denmark. Located between Iceland, Norway, and the United Kingdom, the islands have a populat ...
that has come into regular use elsewhere. In Britain, it is sometimes known by the name bonxie, a
Shetland Shetland (until 1975 spelled Zetland), also called the Shetland Islands, is an archipelago in Scotland lying between Orkney, the Faroe Islands, and Norway, marking the northernmost region of the United Kingdom. The islands lie about to the ...
name of Norse origin. The genus name ''Stercorarius'' is
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
and means "of dung"; the food disgorged by other birds when pursued by skuas was once thought to be excrement. The species is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
: no
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognised.


Description

Great skuas measure long and have a
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the opposite wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingsp ...
. One study found that 112 males weighed an average of and that 125 females weighed an average of . Adults are a streaked greyish brown, with a black cap, while juveniles are a warmer brown and unstreaked below. They have a short, blunt tail, and a powerful flight. The great skua's call is a harsh ''hah-hah-hah-hah''; quacking and croaking noises have also been heard. Distinguishing this skua from the other North Atlantic skuas (
parasitic jaeger The parasitic jaeger (North America) or Arctic skua (Europe) (''Stercorarius parasiticus''), is a seabird in the skua family Stercorariidae. It is a migratory species breeding in Northern Scandinavia, Scotland, Iceland, Greenland, Northern Canada ...
,
pomarine jaeger The pomarine jaeger (''Stercorarius pomarinus''), pomarine skua, or pomatorhine skua, is a seabird in the skua family Stercorariidae. It is a migrant, wintering at sea in the tropical oceans. Taxonomy The Pomarine Jaeger is most closely related ...
and
long-tailed jaeger The long-tailed skua or long-tailed jaeger (''Stercorarius longicaudus'') is a seabird in the skua family Stercorariidae. Etymology The word "jaeger" is derived from the German word ''Jäger'', meaning "hunter". The English word "skua" comes fr ...
) is relatively straightforward. The
herring gull Herring gull is a common name for several birds in the genus ''Larus'', all formerly treated as a single species. Three species are still combined in some taxonomies: * American herring gull (''Larus smithsonianus'') - North America * European h ...
size, massive barrel chest and white wing flashes of this bird are distinctive even at a distance. It is sometimes said to give the impression of a
common buzzard The common buzzard (''Buteo buteo'') is a medium-to-large bird of prey which has a large range. It is a member of the genus '' Buteo'' in the family Accipitridae. The species lives in most of Europe and extends its breeding range across much of ...
. Identification of this skua is only complicated when it is necessary to distinguish it from the closely related large southern-hemisphere skuas. Despite its name, the great skua is marginally smaller on average than the other 3 large southern-hemisphere skuas, although not by enough to distinguish them by size in the field. Some authorities still regard the great skua as conspecific with some of these southern skuas, and as a group they have sometimes been separated in the genus ''Catharacta'', although currently this taxonomy is not commonly followed.


Origins

Genetic studies have found surprising similarities between the great skua and the pomarine skua, despite their dissimilar appearance. Many ornithologists now believe either that the great skua originated as a hybrid between the pomarine skua and one of the southern-hemisphere species, presumably as a result of vagrancy or migration across the equator by the southern species, or that the pomarine skua evolved from hybridization of the great skua and one of the small Arctic species (see
pomarine jaeger The pomarine jaeger (''Stercorarius pomarinus''), pomarine skua, or pomatorhine skua, is a seabird in the skua family Stercorariidae. It is a migrant, wintering at sea in the tropical oceans. Taxonomy The Pomarine Jaeger is most closely related ...
for details).


Breeding

The great skua breeds in
Iceland Iceland is a Nordic countries, Nordic island country between the Atlantic Ocean, North Atlantic and Arctic Oceans, on the Mid-Atlantic Ridge between North America and Europe. It is culturally and politically linked with Europe and is the regi ...
,
Norway Norway, officially the Kingdom of Norway, is a Nordic countries, Nordic country located on the Scandinavian Peninsula in Northern Europe. The remote Arctic island of Jan Mayen and the archipelago of Svalbard also form part of the Kingdom of ...
, the
Faroe Islands The Faroe Islands ( ) (alt. the Faroes) are an archipelago in the North Atlantic Ocean and an autonomous territory of the Danish Realm, Kingdom of Denmark. Located between Iceland, Norway, and the United Kingdom, the islands have a populat ...
, and on
Scottish Scottish usually refers to something of, from, or related to Scotland, including: *Scottish Gaelic, a Celtic Goidelic language of the Indo-European language family native to Scotland *Scottish English *Scottish national identity, the Scottish ide ...
islands, with some individuals breeding on mainland Scotland and in the northwest of
Ireland Ireland (, ; ; Ulster Scots dialect, Ulster-Scots: ) is an island in the North Atlantic Ocean, in Northwestern Europe. Geopolitically, the island is divided between the Republic of Ireland (officially Names of the Irish state, named Irelan ...
. They breed on coastal moorland and rocky islands, usually laying two spotted olive-brown eggs in grass-lined nests. Like other skuas, they will fly at the head of a human or other intruder approaching their nest. Although it cannot inflict serious damage, such an experience with a bird of this size is frightening. They are a migrant species, wintering at sea in the
Atlantic Ocean The Atlantic Ocean is the second largest of the world's five borders of the oceans, oceanic divisions, with an area of about . It covers approximately 17% of Earth#Surface, Earth's surface and about 24% of its water surface area. During the ...
and regularly reaching
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
n waters. They are vagrant to Mediterranean countries (e.g. Turkey).


Diet

They eat mainly fish, birds,
egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the ...
s, carrion, offal,
rodent Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...
s, rabbits, and occasionally berries. They have been known to prey on lambs, and even pony foals. Probably their most prolific food source is by-catch abandoned by fishing vessels. They will often obtain fish by robbing
gull Gulls, or colloquially seagulls, are seabirds of the subfamily Larinae. They are most closely related to terns and skimmers, distantly related to auks, and even more distantly related to waders. Until the 21st century, most gulls were placed ...
s,
tern Terns are seabirds in the family Laridae, subfamily Sterninae, that have a worldwide distribution and are normally found near the sea, rivers, or wetlands. Terns are treated in eleven genera in a subgroup of the family Laridae, which also ...
s and even
northern gannet The northern gannet (''Morus bassanus'') is a seabird, the largest species of the gannet family, Sulidae. It is native to the coasts of the Atlantic Ocean, breeding in Western Europe and Northeastern North America. It is the largest seabird in t ...
s of their catches. They will also directly attack and kill other seabirds, up to the size of herring gulls. Like most other skua
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
, it continues this piratical behaviour throughout the year, showing less agility and more brute force than the smaller skuas when it harasses its victims. A common technique is to fly up to a gannet in mid-air and grab it by the wing, so that it stalls and falls into the sea, where the great skua then physically attacks it until it surrenders its catch. Due to its size, aggressive nature and fierce defence of its nest, the great skua has little to fear from other predators. While fledglings can fall prey to
rat Rats are various medium-sized, long-tailed rodents. Species of rats are found throughout the order Rodentia, but stereotypical rats are found in the genus ''Rattus''. Other rat genera include '' Neotoma'' (pack rats), '' Bandicota'' (bandicoo ...
s,
cat The cat (''Felis catus''), also referred to as the domestic cat or house cat, is a small domesticated carnivorous mammal. It is the only domesticated species of the family Felidae. Advances in archaeology and genetics have shown that the ...
s or the
Arctic fox The Arctic fox (''Vulpes lagopus''), also known as the white fox, polar fox, or snow fox, is a small species of fox native to the Arctic regions of the Northern Hemisphere and common throughout the Tundra#Arctic tundra, Arctic tundra biome. I ...
, healthy adults are threatened only by greater raptors such as the
golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known bird of pr ...
, the
white-tailed eagle The white-tailed eagle (''Haliaeetus albicilla''), sometimes known as the 'sea eagle', is a large bird of prey, widely distributed across temperate Eurasia. Like all eagles, it is a member of the family Accipitridae (or accipitrids) which also ...
, and more rarely, at sea by the
orca The orca (''Orcinus orca''), or killer whale, is a toothed whale and the largest member of the oceanic dolphin family. The only extant species in the genus '' Orcinus'', it is recognizable by its black-and-white-patterned body. A cosmopol ...
.


Predation account

An aerial
apex predator An apex predator, also known as a top predator or superpredator, is a predator at the top of a food chain, without natural predators of its own. Apex predators are usually defined in terms of trophic dynamics, meaning that they occupy the hig ...
, the great skua is an also an aggressive
kleptoparasite Kleptoparasitism (originally spelt clepto-parasitism, meaning "parasitism by theft") is a form of feeding in which one animal deliberately takes food from another. The strategy is Evolutionarily stable strategy, evolutionarily stable when stealin ...
, deliberately harassing birds as large as gannets to steal a free meal. It also readily kills and eats smaller birds such as
puffins Puffins are any of three species of small alcids (auks) in the bird genus ''Fratercula''. These are pelagic seabirds that feed primarily by diving in the water. They breed in large colonies on coastal cliffs or offshore islands, nesting in crev ...
. Great skuas show little to no fear of humans – anybody getting close to the nest will be repeatedly dive-bombed by the territorial adults. Unusual behaviour by St Kilda's skuas was recorded in 2007 during research into recent falls in the
Leach's storm petrel Leach's storm petrel or Leach's petrel (''Hydrobates leucorhous'') is a small seabird of the tubenose order. It is named after the British zoologist William Elford Leach. The scientific name is derived from Ancient Greek. ''Hydrobates'' is from ...
population. Using night vision gear, ecologists observed the skuas preying on the petrels at night, a remarkable strategy for a seabird.


Gallery

File:GreatSkuaInWater.jpg, Great skua in water File:Skua_Runde.jpg, At the Norwegian bird island
Runde Runde is an island in Herøy Municipality (Møre og Romsdal), Herøy Municipality in Møre og Romsdal county, Norway. The island has a population of 113 people (as of 2015), and it is connected by the Runde Bridge to the island of Remøya to th ...
File:Skua_Runde2.jpg, At Runde File:Great skua (Stercorarius skua), Hallig Hooge.jpg, Great skua at North Sea island Hallig Hooge / Germany File:Große Raubmöwe (Stercorarius skua) auf Handa Island.JPG, At Handa Island / Scotland Stercorarius skua MHNT.ZOO.2010.11.139.8.jpg , Museum specimen


References

*


External links

* at the Handa Island Skua Project * * * * * * * * {{Authority control
great skua The great skua (''Stercorarius skua''), sometimes known by the name bonxie in Britain, is a large seabird in the skua family Stercorariidae. It is roughly the size of a herring gull. It mainly eats fish caught at the sea surface or taken fro ...
Birds of the Arctic Birds of Iceland Birds of Scandinavia Birds of the Atlantic Ocean
great skua The great skua (''Stercorarius skua''), sometimes known by the name bonxie in Britain, is a large seabird in the skua family Stercorariidae. It is roughly the size of a herring gull. It mainly eats fish caught at the sea surface or taken fro ...
Taxa named by Morten Thrane Brünnich Taxobox binomials not recognized by IUCN Apex predators