Bolt's Farm
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Bolt's Farm is a
palaeontological Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
site in the
Cradle of Humankind The Cradle of Humankind is a paleoanthropological site that is located about northwest of Johannesburg, South Africa, in the Gauteng province. Declared a World Heritage Site by UNESCO in 1999, the site is home to the largest known concentrat ...
World Heritage Site,
Gauteng Gauteng ( , ; Sotho-Tswana languages, Sotho-Tswana for 'place of gold'; or ) is one of the nine provinces of South Africa. Situated on the Highveld, Gauteng is the smallest province by land area in South Africa. Although Gauteng accounts f ...
province, South Africa. With more than 30 fossil deposits dating back 4.5 Ma, it is one of the oldest sites currently discovered in the Cradle of Humankind. It consists of multiple cavities, pits, and quarries, where caves have eroded away, exposing their
fossiliferous A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved i ...
interiors. Although this site has not yet yielded the
hominid The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); '' Gorilla'' (the ...
fossils for which the Cradle of Humankind is known, Bolt's Farm is still an important source of fossils from various species of
Early Pliocene Early may refer to: Places in the United States * Early, Iowa, a city * Early, Texas, a city * Early Branch, a stream in Missouri * Early County, Georgia * Fort Early, Georgia, an early 19th century fort Music * Early B, stage name of Jamaican d ...
and
Plio-Pleistocene The Plio-Pleistocene is an informally described geological pseudo-period, which begins about 5 million years ago (Mya) and, drawing forward, combines the time ranges of the formally defined Pliocene and Pleistocene epochs—marking from about 5&n ...
fauna, including
primates Primates is an order of mammals, which is further divided into the strepsirrhines, which include lemurs, galagos, and lorisids; and the haplorhines, which include tarsiers and simians ( monkeys and apes). Primates arose 74–63  ...
and
big cats The term "big cat" is typically used to refer to any of the five living members of the genus ''Panthera'', namely the tiger, lion, jaguar, leopard, and snow leopard. All cats descend from the ''Felidae'' family, sharing similar musculature, c ...
.


Geography

Bolt's Farm is located about southwest of the
Sterkfontein Sterkfontein (Afrikaans for ''Strong Spring'') is a set of limestone caves of special interest in paleoanthropology located in Gauteng province, about northwest of Johannesburg, South Africa in the Muldersdrift area close to the town of K ...
archaeological site and is located in the southwestern corner of the
Cradle of Humankind The Cradle of Humankind is a paleoanthropological site that is located about northwest of Johannesburg, South Africa, in the Gauteng province. Declared a World Heritage Site by UNESCO in 1999, the site is home to the largest known concentrat ...
World Heritage Site. The site is carved out of the Malmani Subgroup of the Transvaal Supergroup, which formed during the
Palaeoproterozoic The Paleoproterozoic Era (also spelled Palaeoproterozoic) is the first of the three sub-divisions ( eras) of the Proterozoic eon, and also the longest era of the Earth's geological history, spanning from (2.5–1.6  Ga). It is further subd ...
era around 2.6–2.4 billion years ago under what was then an
inland sea An inland sea (also known as an epeiric sea or an epicontinental sea) is a continental body of water which is very large in area and is either completely surrounded by dry land (landlocked), or connected to an ocean by a river, strait or " arm of ...
. The pockmarked appearance of the area is the result of heavy ancient cave erosion down to the floors and walls of the passages, resulting in dozens of dolomite solution cavities and palaeokarst pits that have preserved fossils in
breccia Breccia ( , ; ) is a rock composed of large angular broken fragments of minerals or Rock (geology), rocks cementation (geology), cemented together by a fine-grained matrix (geology), matrix. The word has its origins in the Italian language ...
,
speleothem A speleothem (; ) is a geological formation made by mineral deposits that accumulate over time in natural caves. Speleothems most commonly form in calcareous caves due to carbonate dissolution reactions. They can take a variety of forms, depen ...
,
sandstone Sandstone is a Clastic rock#Sedimentary clastic rocks, clastic sedimentary rock composed mainly of grain size, sand-sized (0.0625 to 2 mm) silicate mineral, silicate grains, Cementation (geology), cemented together by another mineral. Sand ...
, and
siltstone Siltstone, also known as aleurolite, is a clastic sedimentary rock that is composed mostly of silt. It is a form of mudrock with a low clay mineral content, which can be distinguished from shale by its lack of fissility. Although its permeabil ...
. These deposits occur over a stretch of hillside. It is unknown if these caves were once interconnected and contemporary with one another, or if they existed separately. The oldest of these deposits yet known is Waypoint 160, which was discovered in 1996 and has been dated back to the
Early Pliocene Early may refer to: Places in the United States * Early, Iowa, a city * Early, Texas, a city * Early Branch, a stream in Missouri * Early County, Georgia * Fort Early, Georgia, an early 19th century fort Music * Early B, stage name of Jamaican d ...
, approximately 4.5–4 Ma. Almost all other sites have been dated back only to the
Plio-Pleistocene The Plio-Pleistocene is an informally described geological pseudo-period, which begins about 5 million years ago (Mya) and, drawing forward, combines the time ranges of the formally defined Pliocene and Pleistocene epochs—marking from about 5&n ...
; dates vary from one pit to the next, although most fall in the range of 2–1 Ma. One site, New Cave, may hold fossils that are significantly more recent; remains there have been dated to the
Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as the Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division ...
into the
Holocene The Holocene () is the current geologic time scale, geological epoch, beginning approximately 11,700 years ago. It follows the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene to ...
. Palaeontological studies of the fossils recovered at the site seem to suggest that, at the time the site was active, the region was drier and the area was more open than it is today. The species recovered from Bolt's Farm and the greater Sterkfontein Valley area may have inhabited a range of local habitat types, ranging from covered forest to open grasslands. Browser and grazer species, as well as ground and tree-dwelling primates, are represented in the fossil record. At least one location on the site dated to 2–1.5 Ma, Pit 23, has been theorised to have been a "death trap" where ancient fauna inadvertently fell to their deaths and their remains were preserved over time.


Discovery and history of excavations

The area on the northern edge of a farm owned by Billy Bolt, for whom the site is named, was first used as a
speleothem A speleothem (; ) is a geological formation made by mineral deposits that accumulate over time in natural caves. Speleothems most commonly form in calcareous caves due to carbonate dissolution reactions. They can take a variety of forms, depen ...
quarry during the late 19th and early 20th centuries. Miners harvested this speleothem for use in
lime Lime most commonly refers to: * Lime (fruit), a green citrus fruit * Lime (material), inorganic materials containing calcium, usually calcium oxide or calcium hydroxide * Lime (color), a color between yellow and green Lime may also refer to: Bo ...
production. This process unearthed fossil deposits, which were first sampled and examined by palaeontologist
Robert Broom Robert Broom Fellow of the Royal Society, FRS FRSE (30 November 1866 6 April 1951) was a British- South African medical doctor and palaeontologist. He qualified as a medical practitioner in 1895 and received his DSc in 1905 from the University ...
during a 1936 expedition to Transvaal. Broom named many of the fossil
taxa In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
recovered from the site. In 1947 and 1948, a team of researchers from the
University of California Museum of Paleontology The University of California Museum of Paleontology (UCMP) is a paleontology museum located on the campus of the University of California, Berkeley. The museum is within the Valley Life Sciences Building (VLSB), designed by George W. Kelham ...
(UCMP) led by palaeontologists
Frank Peabody Frank Elmer Peabody (28 August 1914 - 27 June 1958), was an American palaeontologist noted for his research on fossil trackways and reptile and amphibian skeletal structure. He attended high school and junior college in the San Francisco Bay Are ...
and
Charles Lewis Camp Charles Lewis Camp (March 12, 1893 – August 14, 1975) was an American Palaeontology, palaeontologist and Zoology, zoologist, working from the University of California, Berkeley. He took part in excavations at the 'Placerias Quarry', in 1930 and ...
, called the
University of California The University of California (UC) is a public university, public Land-grant university, land-grant research university, research university system in the U.S. state of California. Headquartered in Oakland, California, Oakland, the system is co ...
Africa Expedition, began exploring palaeontological sites across eastern and southern Africa. A team of the southern branch of the expedition led by Peabody was responsible for the first excavation at Bolt's Farm. The mining had created rubble chunks containing fossiliferous
breccia Breccia ( , ; ) is a rock composed of large angular broken fragments of minerals or Rock (geology), rocks cementation (geology), cemented together by a fine-grained matrix (geology), matrix. The word has its origins in the Italian language ...
; early investigations retrieved material from this rubble, instead of mining it ''
in situ is a Latin phrase meaning 'in place' or 'on site', derived from ' ('in') and ' ( ablative of ''situs'', ). The term typically refers to the examination or occurrence of a process within its original context, without relocation. The term is use ...
''. Additionally, the UCMP excavations were centred on the main quarry. Multiple smaller excavations have been completed in the decades since. Historically, Bolt's Farm has been ignored for sites where
hominid The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); '' Gorilla'' (the ...
fossils have already been discovered, which are of much more interest to archaeologists; thus, the study of the site has lagged behind that of its neighbors like
Sterkfontein Sterkfontein (Afrikaans for ''Strong Spring'') is a set of limestone caves of special interest in paleoanthropology located in Gauteng province, about northwest of Johannesburg, South Africa in the Muldersdrift area close to the town of K ...
. Until the 1990s, the precise locations of multiple fossiliferous pits within the larger site were poorly recorded and thus unable to be located by subsequent archaeologists. Renewed surveying began in the 1990s has attempted to rediscover the deposits first described in—but barely explored since—the 1940s. In the process, these surveys have unsurprisingly uncovered new sites. In 1991, Basil Cooke published the first updated map of the site since the 1940s excavations; however, attempts to line up Peabody's original map and site descriptions with sites identified in more recent surveys remain a source of confusion for researchers. However, Cooke's survey marked the beginning of a period of renewed interest in Bolt's Farm. A survey done by French researchers from 1996 to 1999 revealed new fossil species and sites, including Waypoint 160, which is thought to be the oldest known site in the Cradle of Humankind. A survey done by the Cradle of Humankind's Human Origins and Past Environments Research Unit (HRU) team between 2008 and 2011 identified 23 old sites and eight new ones. Further surveys carried out in the late 2010s have used
GIS A geographic information system (GIS) consists of integrated computer hardware and software that store, manage, analyze, edit, output, and visualize geographic data. Much of this often happens within a spatial database; however, this is not ...
and aerial drone surveying to more accurately identify these sites. Some of the specimens collected during the 1940s excavations have been lost; however, many of those surviving are stored at the
University of California Museum of Paleontology The University of California Museum of Paleontology (UCMP) is a paleontology museum located on the campus of the University of California, Berkeley. The museum is within the Valley Life Sciences Building (VLSB), designed by George W. Kelham ...
at the
University of California, Berkeley The University of California, Berkeley (UC Berkeley, Berkeley, Cal, or California), is a Public university, public Land-grant university, land-grant research university in Berkeley, California, United States. Founded in 1868 and named after t ...
. Beginning in the late 1950s, loans and
repatriation Repatriation is the return of a thing or person to its or their country of origin, respectively. The term may refer to non-human entities, such as converting a foreign currency into the currency of one's own country, as well as the return of mi ...
efforts have handed over many of the fossils to the Evolutionary Studies Institute at the
University of the Witwatersrand The University of the Witwatersrand, Johannesburg (), commonly known as Wits University or Wits, is a multi-campus Public university, public research university situated in the northern areas of central Johannesburg, South Africa. The universit ...
in
Johannesburg Johannesburg ( , , ; Zulu language, Zulu and Xhosa language, Xhosa: eGoli ) (colloquially known as Jozi, Joburg, Jo'burg or "The City of Gold") is the most populous city in South Africa. With 5,538,596 people in the City of Johannesburg alon ...
. The
Ditsong National Museum of Natural History The Ditsong National Museum of Natural History, formerly the Transvaal Museum, is a natural history museum situated in Pretoria, South Africa. It is located on Paul Kruger Street, between Visagie and Minnaar Streets, opposite the Pretoria City H ...
in
Pretoria Pretoria ( ; ) is the Capital of South Africa, administrative capital of South Africa, serving as the seat of the Executive (government), executive branch of government, and as the host to all foreign embassies to the country. Pretoria strad ...
is also in possession of Bolt's Farm specimens, where many are kept on semi-permanent display to the public.


Palaeontology

Fossils recovered from the Bolt's Farm pits represent a wide array of
macrofauna Fauna (: faunae or faunas) is all of the animal life present in a particular region or time. The corresponding terms for plants and fungi are ''flora'' and ''funga'', respectively. Flora, fauna, funga and other forms of life are collectively r ...
. Animals active in the Bolt's Farm area during the
Plio-Pleistocene The Plio-Pleistocene is an informally described geological pseudo-period, which begins about 5 million years ago (Mya) and, drawing forward, combines the time ranges of the formally defined Pliocene and Pleistocene epochs—marking from about 5&n ...
included
big cat The term "big cat" is typically used to refer to any of the five living members of the genus ''Panthera'', namely the tiger, lion, jaguar, leopard, and snow leopard. All cats descend from the ''Felidae'' family, sharing similar musculature, c ...
s, non-
hominid The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); '' Gorilla'' (the ...
primate Primates is an order (biology), order of mammals, which is further divided into the Strepsirrhini, strepsirrhines, which include lemurs, galagos, and Lorisidae, lorisids; and the Haplorhini, haplorhines, which include Tarsiiformes, tarsiers a ...
s, snakes, horses,
mustelid The Mustelidae (; from Latin , weasel) are a diverse family of carnivoran mammals, including weasels, badgers, otters, polecats, martens, grisons, and wolverines. Otherwise known as mustelids (), they form the largest family in the suborde ...
s,
bovids The Bovidae comprise the family (biology), biological family of cloven-hoofed, ruminant mammals that includes Bos, cattle, bison, Bubalina, buffalo, antelopes (including Caprinae, goat-antelopes), Ovis, sheep and Capra (genus), goats. A member o ...
, pigs,
antelope The term antelope refers to numerous extant or recently extinct species of the ruminant artiodactyl family Bovidae that are indigenous to most of Africa, India, the Middle East, Central Asia, and a small area of Eastern Europe. Antelopes do ...
,
rodent Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...
s,
jackal Jackals are Canidae, canids native to Africa and Eurasia. While the word has historically been used for many canines of the subtribe Canina (subtribe), canina, in modern use it most commonly refers to three species: the closely related black-b ...
s, reptiles, and
birds of prey Birds of prey or predatory birds, also known as (although not the same as) raptors, are hypercarnivorous bird species that actively predation, hunt and feed on other vertebrates (mainly mammals, reptiles and smaller birds). In addition to speed ...
. Signs of various
microfauna Microfauna ( and ) are microscopic animals and organisms that exhibit animal-like qualities and have body sizes that are usually <0.1 mm. Microfauna are represented in the animal kingdom (e.g.
have also been uncovered but not studied in-depth. The deposits have also yielded some of the oldest primate fossils in the Cradle of Humankind; they belong to ''
Parapapio ''Parapapio'' is a genus of prehistoric baboons closely resembling the forest-dwelling mangabeys. ''Parapapio'' is distinguished from other '' Papio'' by the lack of an anteorbital drop, thin browridges, absence of maxillary fossae or a sagittal ...
'' (specifically, ''Pp. broomi'' and ''Pp. whitei''), extinct species of baboons that have also been located at
Sterkfontein Sterkfontein (Afrikaans for ''Strong Spring'') is a set of limestone caves of special interest in paleoanthropology located in Gauteng province, about northwest of Johannesburg, South Africa in the Muldersdrift area close to the town of K ...
,
Taung Taung is a small town situated in the North West Province of South Africa. The name means ''place of the lion'' and was named after Tau, the King of the Barolong people. ''Tau'' is the Tswana word for lion. Taung skull fossil site In 1924, a sk ...
, and
Makapansgat Makapansgat () (or Makapan Valley World Heritage Site) is an archaeological location within the Makapansgat and Zwartkrans Valleys, northeast of Mokopane in Limpopo province, South Africa. It is an important palaeontological site, with the loca ...
. The ''Pp. whitei'' specimens from Bolt's Farm are the most complete remains of this species that have been discovered. Other primate remains include ''
Theropithecus ''Theropithecus'' is a genus of primates in the family Cercopithecidae. It contains a single living species, the gelada (''Theropithecus gelada''), native to the Ethiopian Highlands. Additional species are known from fossils, including: * '' The ...
'', ''
Papio robinsoni Baboons are primates comprising the genus ''Papio'', one of the 23 genera of Old World monkeys, in the family Cercopithecidae. There are six species of baboon: the hamadryas baboon, the Guinea baboon, the olive baboon, the yellow baboon, the ...
'' and '' Cercopithecoides williamsi''. Because hominid fossils contemporary to the depositional period have been recovered in similar conditions in east African archaeological sites, several researchers have put forth the suggestion that hominid fossils could be found at Bolt's Farm as well. Other species represented in the Bolt's Farm fossil deposits are ''
Antidorcas recki ''Antidorcas recki'' is an extinct species of gazelle, related to the extant springbok, from the Pliocene and Pleistocene of southern and eastern Africa. Taxonomy ''Antidorcas recki'' was named in 1932 by Schwarz, and assigned to '' Adenota'' ...
'', '' Ictonyx bolti'', '' Metridiochoerus andrewsi'', '' Euryotomys bolti'', '' Elephantulus antiquus'', '' Boltimys broomi'', and '' Proteles cristatus''. Bolt's Farm has yielded some of the best specimens of
Plio-Pleistocene The Plio-Pleistocene is an informally described geological pseudo-period, which begins about 5 million years ago (Mya) and, drawing forward, combines the time ranges of the formally defined Pliocene and Pleistocene epochs—marking from about 5&n ...
big cats in South Africa, including a complete skull and jaw of the saber-toothed cat '' Dinofelis barlowi''. The first fossil remains of a snake belonging to the family ''
Elapidae Elapidae (, commonly known as elapids , from , variant of "sea-fish") is a family of snakes characterized by their permanently erect fangs at the front of the mouth. Most elapids are venomous, with the exception of the genus '' Emydocephalus ...
'' recovered from southern Africa were collected from Bolt's Farm in 2009. The 2016 discovery at Bolt's Farm of the first fossil '' Agama'' lizard recovered from the Cradle of Humankind was announced in 2020.


References


External links


Bolt's Farm on the official Cradle of Humankind website
{{Greater Johannesburg, natural Pliocene paleontological sites of Africa Pleistocene paleontological sites of Africa Mogale City Local Municipality Landforms of Gauteng Archaeological sites of Southern Africa Cradle of Humankind