Bolboceratidae
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Bolboceratidae is a
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
of
beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
. It was historically treated as a
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
of the earth-boring dung beetles, or "dor beetles" (family Geotrupidae), but has been considered a separate family by many authors since 1995.Bolboceratidae at TOL
/ref> Some recent classifications have persisted in treating bolboceratids as a subfamily (e.g.) but these classifications are contradicted by recent phylogenetic studies of relationships indicating that bolboceratids are not closely related to geotrupids (e.g., that bolboceratids are more closely related to Pleocomidae and
Passalidae Passalidae is a family of beetles known variously as "bessbugs", "bess beetles", "betsy beetles", "betsy bugs", or "horned passalus beetles". Nearly all of the 500-odd species are tropical; species found in North America are notable for their siz ...
).


Genera

* '' Athyreus'' * '' Australobolbus'' * '' Blackbolbus'' * '' Blackburnium'' * '' Bolbaffer'' * '' Bolbaffroides'' * '' Bolbelasmus'' * '' Bolbobaineus'' * '' Bolbocaffer'' * '' Bolbocerastes'' * '' Bolboceratex'' * '' Bolboceratops'' * '' Bolbocerodema'' * '' Bolboceroides'' * '' Bolbocerosoma'' * '' Bolbochromus'' * '' Bolbogonium'' * '' Bolbohamatum'' * '' Bolboleaus'' * '' Bolborhachium'' * '' Bolborhinum'' * '' Bolborhombus'' * '' Bolbothyreus'' * '' Bolbotrypes'' * '' Bradycinetulus'' * '' Cretobolbus'' * '' Elephastomus'' * '' Eubolbitus'' * '' Eucanthus'' * '' Gilletinus'' * '' Halffterobolbus'' * '' Meridiobolbus'' * '' Mesoathyreus'' * '' Mimobolbus'' * '' Namibiobolbus'' * '' Namibiotrupes'' * '' Neoathyreus'' * '' Odonteus'' * '' Parabolbapium'' * '' Parathyreus'' * '' Pereirabolbus'' * '' Prototrupes'' * '' Pseudoathyreus'' * '' Socotrabolbus'' * '' Somalobolbus'' * '' Senaspidius'' * '' Zefevazia''


Fossil genera

* †'' Amberathyreus'' Bai and Zhang 2017
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. Th ...
, Myanmar,
Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
* †'' Cretobolbus'' Nikolajev 1996,
Zaza Formation The Zaza Formation is a geological formation located in Buryatia (Russia). It dates to the Lower Cretaceous. The age of the formation is disputed, and is considered likely to be Valanginian-Hauterivian, or Aptian in age. It comprises sandstones, ...
, Russia,
Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...


References


External links


Encyclopedia of Life entry
Beetle families {{Scarabaeoidea-stub