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The blue-headed macaw or Coulon's macaw (''Primolius couloni'') is a
macaw Macaws are a group of New World parrots that are long-tailed and often colorful. They are popular in aviculture or as companion parrots, although there are conservation concerns about several species in the wild. Biology Of the many differe ...
native to eastern Peru, northwestern
Bolivia , image_flag = Bandera de Bolivia (Estado).svg , flag_alt = Horizontal tricolor (red, yellow, and green from top to bottom) with the coat of arms of Bolivia in the center , flag_alt2 = 7 × 7 square p ...
(mainly in Pando), and far western Brazil (in
Acre The acre is a unit of land area used in the imperial Imperial is that which relates to an empire, emperor, or imperialism. Imperial or The Imperial may also refer to: Places United States * Imperial, California * Imperial, Missouri * Imp ...
). It has a total length of about 41 cm (16 in), making it a member of the group of smaller macaws sometimes known as the mini-macaws, which includes any species of macaw with a total length of 50 cm (20 in) or less. As in all macaws, its tail is long and pointed and the bill is large and heavy. The
specific epithet In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bot ...
''couloni'' is in honor of Swiss naturalist
Paul Louis Coulon Paul may refer to: *Paul (given name), a given name (includes a list of people with that name) *Paul (surname), a list of people People Christianity *Paul the Apostle (AD c.5–c.64/65), also known as Saul of Tarsus or Saint Paul, early Chris ...
(1804-1894).


Description

The blue-headed macaw is 41 cm (16 in) long. It has mainly green plumage (often tinged olive, esp. below) with the head, flight feathers and primary coverts blue. The uppertail has a maroon base, a narrow green center and a blue tip. The undertail and underwing are greenish-yellow similar to that of several other small macaws (e.g. red-bellied and golden-collared macaw). The medium-sized bill is pale greyish-horn with a black base (extent varies, but upper mandible in adults typically appears mainly pale). The iris is whitish with a narrow, often barely visible, maroon eye-ring. Unlike most other macaws, the facial skin and lores are dark greyish. The legs are dull pinkish.
Juvenile Juvenile may refer to: *Juvenile status, or minor (law), prior to adulthood *Juvenile (organism) *Juvenile (rapper) (born 1975), American rapper * ''Juvenile'' (2000 film), Japanese film * ''Juvenile'' (2017 film) *Juvenile (greyhounds), a greyho ...
s resemble adults, but with the entire bill black, greyer legs, darker iris and the facial skin and lores white.


Taxonomy

In recent years it has often been placed in the genus ''Propyrrhura''; however, as per
ICZN The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its publisher, the I ...
rules, the species is now considered to be in the genus '' Primolius''. Earlier, it has also been placed in the genus ''
Ara ARA may refer to: Media and the arts * American-Romanian Academy of Arts and Sciences * '' Artistička Radna Akcija'', compilation album released in former Yugoslavia * Associate of the Royal Academy, denoting membership in the British Royal Acad ...
'', which, at present, is only used for some of the larger macaws.


Habitat

It is found in the south-west
Amazon Amazon most often refers to: * Amazons, a tribe of female warriors in Greek mythology * Amazon rainforest, a rainforest covering most of the Amazon basin * Amazon River, in South America * Amazon (company), an American multinational technolog ...
and adjacent east
Andean The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S l ...
foothills Foothills or piedmont are geographically defined as gradual increases in elevation at the base of a mountain range, higher hill range or an upland area. They are a transition zone between plains and low relief hills and the adjacent topographi ...
. It prefers openings in humid forest, e.g. along rivers or clearings, but has also been observed in
Moriche Palm ''Mauritia flexuosa'', known as the moriche palm, ''ité'' palm, ''ita'', ''buriti'', ''muriti'', ''miriti'' (Brazil), ''canangucho'' (Colombia), ''acho'' (Ecuador), or ''aguaje'' (Peru), is a palm tree. It grows in and near swamps and other wet ...
swamps and outskirts of towns. It occurs from the
lowland Upland and lowland are conditional descriptions of a plain based on elevation above sea level. In studies of the ecology of freshwater rivers, habitats are classified as upland or lowland. Definitions Upland and lowland are portions of p ...
s up to an altitude of 1550 m (5100 ft). As several other
parrot Parrots, also known as psittacines (), are birds of the roughly 398 species in 92 genera comprising the order Psittaciformes (), found mostly in tropical and subtropical regions. The order is subdivided into three superfamilies: the Psittaco ...
s, the blue-headed macaw is known to visit clay-licks.


Behavior

Little is known about its reproductive behavior in the wild, but a possible nest has been recorded in a bamboo cavity and another in a tree cavity. In captivity, the reproductive cycle appears to be roughly annual with a clutch size of 2-4 being the norm. In the wild it is typically seen in groups of 2-4 individuals, with occasional records of groups up to c. 60 individuals. Its call is higher-pitched and softer than that of most other macaws.


Status

Until recently it was considered fairly common, but a review in 2006 by
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding ...
suggested it was rare with a decreasing total population of 1000-2500 individuals. It has therefore been uplisted to
endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inv ...
in the 2007
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
. Parts of the range of this species remain poorly known, but Tobias & Brightsmith (2007) has suggested that previous estimates were too low, with actual number of 9200-46000 mature individuals more likely. It has therefore been suggested that
vulnerable Vulnerable may refer to: General * Vulnerability * Vulnerability (computing) * Vulnerable adult * Vulnerable species Music Albums * ''Vulnerable'' (Marvin Gaye album), 1997 * ''Vulnerable'' (Tricky album), 2003 * ''Vulnerable'' (The Used album) ...
might be a more appropriate category for this species. Much of the forest within its range remains intact, but habitat loss could be a threat, at least locally. It occurs in several protected areas, e.g.
Tambopata-Candamo Tambopata National Reserve ( es, Reserva Nacional Tambopata) is a Peruvian nature reserve located in the southeastern region of Madre de Dios. It was established on September 4, 2000, by decree of President Alberto Fujimori. The reserve protects se ...
and
Manu Manu may refer to: Geography *Manú Province, a province of Peru, in the Madre de Dios Region ** Manú National Park, Peru **Manú River, in southeastern Peru * Manu River (Tripura), which originates in India and flows into Bangladesh *Manu Temp ...
. The capture of individuals for the wild bird trade potentially presents a serious problem. It is rare in captivity and consequently prices are high (US$12,500 but the past few years have declined as low as $4,000 - 5,000 US in some EU countries). International trade in this species was virtually unknown in 1993, where
CITES CITES (shorter name for the Convention on International Trade in Endangered Species of Wild Fauna and Flora, also known as the Washington Convention) is a multilateral treaty to protect endangered plants and animals from the threats of interna ...
only registered three legally traded individuals, but this had risen to 55 individuals in 2000. As of August 2007, the
International Species Information System Species360 (formerly International Species Information System or ISIS) founded in 1974, is an international non-profit organization that maintains an online database of wild animals under human care. , the organization serves more than 1,000 zoo ...
listed only 26 individuals in
zoo A zoo (short for zoological garden; also called an animal park or menagerie) is a facility in which animals are kept within enclosures for public exhibition and often bred for conservation purposes. The term ''zoological garden'' refers to zoo ...
s outside its native countries, and
Loro Parque Loro Parque (Spanish for "parrot park") or 'Loro Park' is a 135,000 m² (13.5 ha) zoo on the outskirts of Puerto de la Cruz on Tenerife, Spain where it houses an extensive and diverse reserve of animal and plant species. The park was conceived ...
(not included on ISIS) has 35+ individuals. CITES reported that approx. 50 illegally held blue-headed macaws were seized throughout the World in 1993–2000, and an investigation in
Germany Germany, officially the Federal Republic of Germany (FRG),, is a country in Central Europe. It is the most populous member state of the European Union. Germany lies between the Baltic and North Sea to the north and the Alps to the sou ...
in 2001 resulted in approx. 30 individuals being seized.


References

* Juniper & Parr (1998). ''A Guide to the Parrots of the World''. Pica Press, East Sussex.
To reassign the genus of three macaws
South American Classification Committee. * Tobias, J., and D. J. Brightsmith (2007). ''Distribution, ecology and conservation status of the Blue-headed Macaw Primolius couloni.'' Biological Conservation. 139: 126–138.


External links



(for Peru) with RangeMap
Photo-High Res-(Close-up)

ArticleArticle-2
pbase—"Macaws and Conures"
Blue-headed macaw on Avibase
{{Taxonbar, from=Q934442 blue-headed macaw blue-headed macaw Birds of the Peruvian Amazon blue-headed macaw blue-headed macaw