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The Eurasian blackcap (''Sylvia atricapilla'') is a bird usually known simply as the blackcap. It is a common and widespread
typical warbler The typical warblers are small birds belonging to the genus ''Sylvia'' in the "Old World warbler" (or sylviid warbler) family Sylviidae.Del Hoyo, J.; Elliot, A., & Christie, D. (editors). (2006). ''Handbook of the Birds of the World''. Volume 1 ...
. It has mainly olive-grey upperparts and pale grey underparts, and differences across the five
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are small. Both sexes have a neat coloured cap to the head, black in the male and reddish-brown in the female. The male's typical song is a rich musical warbling, often ending in a loud high-pitched crescendo, but a simpler song is given in some isolated areas, such as valleys in the
Alps The Alps () are some of the highest and most extensive mountain ranges in Europe, stretching approximately across eight Alpine countries (from west to east): Monaco, France, Switzerland, Italy, Liechtenstein, Germany, Austria and Slovenia. ...
. The blackcap's closest relative is the
garden warbler The garden warbler (''Sylvia borin'') is a common and widespread small bird that breeds in most of Europe and in the Palearctic to western Siberia. It is a plain, long-winged and long-tailed typical warbler with brown upperparts and dull white ...
, which looks quite different but has a similar song. The blackcap breeds in much of Europe, western Asia and northwestern Africa, and its preferred habitat is mature
deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...
woodland. The male holds a territory when breeding, which is defended against garden warblers as well as other blackcaps. The nest is a neat cup, built low in
bramble ''Rubus'' is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, most commonly known as brambles. Fruits of various species are known as raspberries, blackberries, dewberries, and bristleberries. I ...
s or scrub, and the
clutch A clutch is a mechanical device that allows an output shaft to be disconnected from a rotating input shaft. The clutch's input shaft is typically attached to a motor, while the clutch's output shaft is connected to the mechanism that does th ...
is typically 4–6 mainly buff eggs, which hatch in about 11 days. The chicks
fledge Fledging is the stage in a flying animal's life between egg, hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnera ...
in 11–12 days, but are cared for by both adults for some time after leaving the nest. The blackcap is a partial migrant; birds from the colder areas of its range winter in scrub or trees in northwestern Europe, around the Mediterranean and in tropical Africa. Some birds from Germany and western continental Europe have adapted to spending the winter in gardens in Great Britain and Ireland. Insects are the main food in the breeding season, but, for the rest of the year, blackcaps survive primarily on small fruit. Garden birds also eat bread, fat and
peanut The peanut (''Arachis hypogaea''), also known as the groundnut, goober (US), goober pea, pindar (US) or monkey nut (UK), is a legume crop grown mainly for its edible seeds. It is widely grown in the tropics and subtropics by small and large ...
s in winter. Despite extensive hunting in Mediterranean countries and the natural hazards of predation and disease, the blackcap has been extending its range for several decades, and is classified by the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the stat ...
as
least concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
. Its rich and varied song has led to it being described as the "mock nightingale" and it has featured in literature, films and music. In Messiaen's opera ''
Saint François d'Assise ''Saint François d'Assise : Scènes Franciscaines'' (English: Franciscan Scenes of Saint Francis of Assisi), or simply ''Saint François d'Assise'', is an opera in three acts and eight scenes by French composer Olivier Messiaen, who was also ...
'', the
saint In Christianity, Christian belief, a saint is a person who is recognized as having an exceptional degree of sanctification in Christianity, holiness, imitation of God, likeness, or closeness to God in Christianity, God. However, the use of the ...
is represented by themes based on the blackcap's song.


Taxonomy

The genus ''Sylvia'', the
typical warbler The typical warblers are small birds belonging to the genus ''Sylvia'' in the "Old World warbler" (or sylviid warbler) family Sylviidae.Del Hoyo, J.; Elliot, A., & Christie, D. (editors). (2006). ''Handbook of the Birds of the World''. Volume 1 ...
s, forms part of a large family of Old World warblers, the
Sylviidae Sylviidae is a family of passerine birds that includes the typical warblers ''Sylvia'', and closely related genus '' Curruca'', formerly included in ''Sylvia''. They are found in Eurasia and Africa, with the greatest diversity in the Mediterrane ...
. The blackcap and its nearest relative, the
garden warbler The garden warbler (''Sylvia borin'') is a common and widespread small bird that breeds in most of Europe and in the Palearctic to western Siberia. It is a plain, long-winged and long-tailed typical warbler with brown upperparts and dull white ...
, are an ancient
species pair In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
which diverged very early from the rest of the genus at between 12 and 16 million years ago. In the course of time, these two species have become sufficiently distinctive that they have been placed in separate subgenera, with the blackcap in subgenus ''Sylvia'' and the garden warbler in ''Epilais''.Shirihai ''et al'' (2001) pp. 25–27. These
sister species In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
have a breeding range which extends farther northeast than all other ''Sylvia'' species except the
lesser whitethroat The lesser whitethroat (''Curruca curruca'') is a common and widespread typical warbler which breeds in temperate Europe, except the southwest, and in the western and central Palearctic. This small passerine bird is strongly migratory, winterin ...
and
common whitethroat The common whitethroat or greater whitethroat (''Curruca communis'') is a common and widespread typical warbler which breeds throughout Europe and across much of temperate western Palearctic, Asia. This small passerine bird is strongly bird migra ...
. The nearest relatives of the garden warbler outside the sister group are believed to be the African hill babbler and Dohrn's thrush-babbler, both of which should probably be placed in ''Sylvia'' rather than their current genera, ''Pseudoalcippe'' and ''Horizorhinus'' respectively. The blackcap was one of the many bird species originally described by
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
in his landmark 1758
10th edition 1 (one, unit, unity) is a number, numeral, and glyph. It is the first and smallest positive integer of the infinite sequence of natural numbers. This fundamental property has led to its unique uses in other fields, ranging from science to sp ...
of ''
Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...
'', as ''Motacilla atricapilla''.Linnaeus (1758) p. 187. The current genus name is from
Modern Latin Neo-LatinSidwell, Keith ''Classical Latin-Medieval Latin-Neo Latin'' in ; others, throughout. (also known as New Latin and Modern Latin) is the style of written Latin used in original literary, scholarly, and scientific works, first in Italy d ...
''silvia'', a woodland sprite, related to ''silva'', a wood.Jobling (2010) p. 376. The species name, like the English name, refers to the male's black cap. ''Atricapilla'' is from the Latin ''ater'', "black", and ''capillus'', "hair (of the head)".Jobling (2010) p. 59.
Fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
s and
subfossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
s of the blackcap have been found in a number of European countries; the oldest, dated to 1.2–1.0 million years ago, are from the
Early Pleistocene The Early Pleistocene is an unofficial epoch (geology), sub-epoch in the international geologic timescale in chronostratigraphy, representing the earliest division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently esti ...
of Bulgaria. Fossils from France show that the genus ''Sylvia'' dates back at least 20 million years.Mason (1995) p. 11.


Subspecies

The differences between subspecies are small, making subspecific boundaries hard to define, and the exact distribution of ''S. a. heineken'' is unclear, since birds from northwest Africa may be of this form.Mason (1995) pp. 15–18. About 2% of male blackcaps on
Madeira Madeira ( ; ), officially the Autonomous Region of Madeira (), is an autonomous Regions of Portugal, autonomous region of Portugal. It is an archipelago situated in the North Atlantic Ocean, in the region of Macaronesia, just under north of ...
and the
Azores The Azores ( , , ; , ), officially the Autonomous Region of the Azores (), is one of the two autonomous regions of Portugal (along with Madeira). It is an archipelago composed of nine volcanic islands in the Macaronesia region of the North Atl ...
are
melanistic Melanism is the congenital excess of melanin in an organism resulting in dark pigment. Pseudomelanism, also called abundism, is another variant of pigmentation, identifiable by dark spots or enlarged stripes, which cover a large part of the bod ...
, with black plumage on the whole head and upper breast. Melanistic females are rarer, and are characterised by grey-brown underparts. This dark morph has also been recorded from the
Canary Islands The Canary Islands (; ) or Canaries are an archipelago in the Atlantic Ocean and the southernmost Autonomous communities of Spain, Autonomous Community of Spain. They are located in the northwest of Africa, with the closest point to the cont ...
, but not from
Cape Verde Cape Verde or Cabo Verde, officially the Republic of Cabo Verde, is an island country and archipelagic state of West Africa in the central Atlantic Ocean, consisting of ten volcanic islands with a combined land area of about . These islands ...
. The melanistic birds were formerly sometimes considered to be a distinct subspecies, ''S. a. obscura''.Shirihai ''et al'' (2010) pp. 54–56.


Description

The blackcap is a mainly grey warbler with distinct male and female
plumage Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...
s. The
nominate subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...
is about long with a wing length. The weight is typically , but can be up to for birds preparing to migrate. The adult male has olive-grey upperparts, other than a paler grey nape and a neat black cap on the head. The underparts are light grey, becoming silvery white on the chin, throat and upper breast. The tail is dark grey, with an olive tint to the outer edge of each feather. The bill and long legs are grey, and the iris is reddish brown. The female resembles the male, but has a reddish-brown cap and a slightly browner tone to the grey of the upperparts. Juveniles are similar to the female, but their upperparts have a slight rufous tinge, and the breast and flanks have a more olive tone; young males have a darker brown cap than their female counterparts. This species is unmistakable; other dark-headed ''Sylvia'' species, such Sardinian and Orphean warblers have extensive black on the head instead of a small cap. They are also larger and have white edges on the tail.Simms (1985) pp. 68–80. Blackcaps have a complete
moult In biology, moulting (British English), or molting (American English), also known as sloughing, shedding, or in many invertebrates, ecdysis, is a process by which an animal casts off parts of its body to serve some beneficial purpose, either at ...
in their breeding areas in August and September prior to migration. Some birds, typically those migrating the greatest distances, have a further partial moult between December and March. Juveniles replace their loosely structured body feathers with adult plumage, starting earlier, but taking longer to complete, than the adults. Blackcaps breeding in the north of the range have an earlier and shorter post-juvenile moult than those further south, and
cross-breeding A crossbreed is an organism with purebred parents of two different breeds, varieties, or populations. A domestic animal of unknown ancestry, where the breed status of only one parent or grandparent is known, may also be called a crossbreed though ...
of captive birds shows that the timing is
gene In biology, the word gene has two meanings. The Mendelian gene is a basic unit of heredity. The molecular gene is a sequence of nucleotides in DNA that is transcribed to produce a functional RNA. There are two types of molecular genes: protei ...
tically controlled.Mason (1995) pp. 92–96.


Voice

The male's song is a rich musical warbling, often ending in a loud high-pitched crescendo, which is given in bursts of up to 30 seconds. The song is repeated for about two-and-a-half minutes, with a short pause before each repetition. In some geographically isolated areas, such as islands,
peninsula A peninsula is a landform that extends from a mainland and is only connected to land on one side. Peninsulas exist on each continent. The largest peninsula in the world is the Arabian Peninsula. Etymology The word ''peninsula'' derives , . T ...
s and valleys in the
Alps The Alps () are some of the highest and most extensive mountain ranges in Europe, stretching approximately across eight Alpine countries (from west to east): Monaco, France, Switzerland, Italy, Liechtenstein, Germany, Austria and Slovenia. ...
, a simplified fluting song occurs, named the ''Leiern'' (drawling) song by the German
ornithologists __NOTOC__ This is a list of ornithologists who have articles, in alphabetical order by surname. See also :Ornithologists. A * John Abbot – US * Clinton Gilbert Abbott – US * William Louis Abbott – US * Humayun Abdulali — India * Joseph ...
who first described it. The song's introduction is like that of other blackcaps, but the final warbling part is a simple alternation between two notes, as in a
great tit The great tit (''Parus major'') is a small passerine bird in the tit family Paridae. It is a widespread and common species throughout Europe, the Middle East, Central Asia and east across the Palearctic to the Amur River, south to parts of No ...
's call but more fluting.Snow & Perrins (1998) pp. 1316–1319.Mason (1995) pp. 85–91. The main song is confusable with that of the garden warbler, but it is slightly higher pitched than in that species, more broken into discrete song segments, and less mellow. Both species have a quiet subsong, a muted version of the full song, which is even harder to separate. The blackcap occasionally mimics the song of other birds,Simms (1985) pp. 56–67. the most frequently copied including the garden warbler and the
common nightingale The common nightingale, rufous nightingale or simply nightingale (''Luscinia megarhynchos''), is a small passerine bird which is best known for its powerful and beautiful Bird vocalization, song. It was formerly classed as a member of the Thrush ...
. The main call is a hard ''tac-tac'', like stones knocking together, and other vocalisations include a squeaking ''sweet'' alarm, and a low-pitched trill similar to that of a garden warbler. Male blackcaps will sometimes sing even when incubating, especially with the second brood. This appears to be intended to maintain the bond with the female. Wintering birds in Africa are quiet initially, but start singing in January or February prior to their return north.Barlow (1997) p. 318.Ash & Atkins (2009) p. 289.


Distribution and habitat


Distribution

The continental breeding range of the blackcap lies between the 14–30° July isotherms, and is occupied by the nominate subspecies, the other forms being restricted to islands or fringe areas in the
Caucasus The Caucasus () or Caucasia (), is a region spanning Eastern Europe and Western Asia. It is situated between the Black Sea and the Caspian Sea, comprising parts of Southern Russia, Georgia, Armenia, and Azerbaijan. The Caucasus Mountains, i ...
and eastern Iberia. Birds on the Mediterranean and Atlantic islands and in the milder west and south of the main Eurasian distribution often winter within the nesting range, but populations elsewhere are migratory. The blackcap is a leap-frog migrant; birds from the north of the breeding range travel furthest south, whereas Mediterranean breeders move much shorter distances. The wintering areas overlap with the breeding range, but also include extensive areas in
West Africa West Africa, also known as Western Africa, is the westernmost region of Africa. The United Nations geoscheme for Africa#Western Africa, United Nations defines Western Africa as the 16 countries of Benin, Burkina Faso, Cape Verde, The Gambia, Gha ...
,
East Africa East Africa, also known as Eastern Africa or the East of Africa, is a region at the eastern edge of the Africa, African continent, distinguished by its unique geographical, historical, and cultural landscape. Defined in varying scopes, the regi ...
south to
Lake Malawi Lake Malawi, also known as Lake Nyasa in Tanzania and Lago Niassa in Mozambique, () is an African Great Lakes, African Great Lake and the southernmost lake in the East African Rift system, located between Malawi, Mozambique and Tanzania. It is ...
, and further north in Ethiopia, South Sudan and Eritrea. The large majority of birds wintering in eastern Africa are of the southwest Asian race, ''S. a. dammholzi''.Zimmerman ''et al'' (1996) p. 462. There is a migratory divide in Europe at
longitude Longitude (, ) is a geographic coordinate that specifies the east- west position of a point on the surface of the Earth, or another celestial body. It is an angular measurement, usually expressed in degrees and denoted by the Greek lett ...
10–11°E. Birds to the west of this line head southwest towards Iberia or West Africa, whereas populations to the east migrate to the eastern Mediterranean and on to East Africa. Cross-breeding in captivity of birds from the resident population on the Canary Islands with migratory blackcaps from Germany showed the urge to migrate is genetically controlled, the offspring showing intermediate behaviour in terms of restlessness at migration time.Mason (1995) pp. 107–112. Similar experiments using birds from southern Germany and eastern Austria, on opposite sides of the migratory divide, demonstrated that the direction of migration is also genetically determined.Newton (2010) pp. 320–329. Climate change appears to be affecting the migration pattern of the garden warbler and blackcap. Both are arriving in Europe earlier than previously, and blackcaps and juvenile (but not adult) garden warblers are departing nearly two weeks later than in the 1980s. Birds of both species are longer-winged and lighter than in the past, suggesting a longer migration as the breeding range expands northwards. In recent decades, substantial numbers of continental European birds have taken to wintering in gardens in Great Britain, and, to a lesser extent, Ireland, where the blackcap was formerly just a summer visitor. Although the British climate is sub-optimal, compensatory factors include the ready availability of food, (particularly from bird tables), a shorter migration distance, and the avoidance of the Alps and the
Sahara Desert The Sahara (, ) is a desert spanning across North Africa. With an area of , it is the largest hot desert in the world and the list of deserts by area, third-largest desert overall, smaller only than the deserts of Antarctica and the northern Ar ...
. It was originally thought that most of these wintering birds come from Germany, and
isotope analysis Isotope analysis is the identification of isotopic signature, abundance of certain stable isotopes of chemical elements within organic and inorganic compounds. Isotopic analysis can be used to understand the flow of energy through a food we ...
(which enables the wintering location to be determined) showed that the continental birds wintering in Britain tend to mate only among themselves, and do not usually interbreed with those wintering in the Mediterranean or western Africa. This is because the British migrants arrive back on the breeding grounds earlier than blackcaps wintering around the Mediterranean, and form pairs before the southern birds arrive. Mixed pairings are also selected against because the hybrid young would migrate in an intermediate direction, which would take them into the
Bay of Biscay The Bay of Biscay ( ) is a gulf of the northeast Atlantic Ocean located south of the Celtic Sea. It lies along the western coast of France from Point Penmarc'h to the Spanish border, and along the northern coast of Spain, extending westward ...
. It now appears that the wintering birds in the UK come from a much wider area than previously thought. The majority come from France, and some individuals come from as far away as Spain and Poland. The steady supply of winter food in gardens gives even Spanish-breeding birds an opportunity to put on weight quicker than in their home range. A 2021 paper showed that blackcaps, particularly adults, wintering in Britain and Ireland showed high site fidelity and low movement between wintering sites, in contrast to blackcaps wintering in their traditional winter ranges. Adults that frequented gardens had better body condition, smaller fat stores, longer bills, and rounder wing tips. The bill and wing tip shapes reflected a more generalist diet than that of birds in traditional winter sites. Attribution 4.0 International (CC BY 4.0) licensed Blackcaps did not exclusively feed in gardens; visits were linked to harsher weather. Individuals generally stayed at garden sites until immediately before spring departure, and supplemental feeding may have benefits for winter survival, When preparing for migration, abundant supplemental food may allow blackcaps to attain better body condition and may facilitate earlier and more successful breeding attempts. In the Iberian Peninsula, migratory blackcaps (and
European robin The European robin (''Erithacus rubecula''), known simply as the robin or robin redbreast in the British Isles, is a small insectivorous passerine bird that belongs to the Old World flycatcher family Muscicapidae. It is found across Europe, ea ...
s) track similar climatic conditions over the season, which sedentary individuals must cope with great variation in climate over the year. This suggests a trade-off between the cost of travelling long distances of migrants, and the flexibility required by sedentary individuals to tolerate a wide variety of environmental conditions.


Habitat

The blackcap's main breeding habitat is mature deciduous woodland, with good scrub cover below the trees. Other habitats, such as parks, large gardens and overgrown hedges, are used as long as they meet the essential requirements of tall trees for songposts and an established
understory In forestry and ecology, understory (American English), or understorey (English in the Commonwealth of Nations, Commonwealth English), also known as underbrush or undergrowth, includes plant life growing beneath the Canopy (biology), forest ca ...
. Where other ''Sylvia'' warblers also breed, blackcaps tend to use taller trees than their relatives, preferably those with a good canopy, such as
pedunculate oak ''Quercus robur'', the pedunculate oak, is a species of flowering plant in the beech and oak family, Fagaceae. It is a large tree, native to most of Europe and western Asia, and is widely cultivated in other temperate regions. It grows on soi ...
. In prime habitat, breeding densities reach 100–200 pairs per square kilometre (250–500 pairs per square mile) in northern Europe, and 500–900 pairs per square kilometre (1,250–2,250 pairs per square mile) in Italy. Densities are much lower in poorer habitats such as conifer forests.Mason (1995) pp. 20–27. Breeding occurs in Europe at altitudes up to . The preferred winter habitat around the Mediterranean is scrub and olive orchards, where densities approach the levels found in the best breeding areas. The British wintering population is atypical, with 95% found in gardens, mostly in towns at altitudes below . In Africa, habitats include cultivated land,
acacia ''Acacia'', commonly known as wattles or acacias, is a genus of about of shrubs and trees in the subfamily Mimosoideae of the pea family Fabaceae. Initially, it comprised a group of plant species native to Africa, South America, and Austral ...
scrub,
mangrove A mangrove is a shrub or tree that grows mainly in coastal saline water, saline or brackish water. Mangroves grow in an equatorial climate, typically along coastlines and tidal rivers. They have particular adaptations to take in extra oxygen a ...
s and forest, and these warblers are found at altitudes up to in the east of the continent. Wintering birds wander in search of good fruit supplies, but often stay in good feeding areas, and return in subsequent winters. Migrants may occur in a wide variety of habitats, such as
reed bed A reedbed or reed bed is a natural habitat found in floodplains, waterlogged depressions and estuaries. Reedbeds are part of a succession from young reeds colonising open water or wet ground through a gradation of increasingly dry ground. As ...
and
fen A fen is a type of peat-accumulating wetland fed by mineral-rich ground or surface water. It is one of the main types of wetland along with marshes, swamps, and bogs. Bogs and fens, both peat-forming ecosystems, are also known as mires ...
, but show a preference for shrubland.Mason (1995) pp. 36–37.


Behaviour


Territory

When male blackcaps return to their breeding areas, they establish a
territory A territory is an area of land, sea, or space, belonging or connected to a particular country, person, or animal. In international politics, a territory is usually a geographic area which has not been granted the powers of self-government, ...
. Adults that have previously bred return to the site they have used in previous summers, whereas inexperienced birds either wander until they find a suitable area, or establish a very large initial territory which contracts under pressure from neighbours. Territorial boundaries are established initially by loud singing, performed while the male displays with his crown raised, tail fanned and slow wingbeats. This display is followed, if necessary, by a chase, often leading to a fight. The typical territory size in a French study was , but in insect-rich tall maquis in Gibraltar, the average was only . Females feed within a home range which may overlap other blackcap territories, and covers up to six times the area of the defended zone.Mason (1995) pp. 38–41. ''Sylvia'' warblers are unusual in that they vigorously defend their territories against other members of their genus as well as
conspecifics Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organism ...
. Blackcaps and garden warblers use identical habits in the same wood, yet aggressive interactions mean that their territories never overlap.Mason (1995) pp. 42–45. Similar songs are a feature of the ''Sylvia'' warblers as a group, and it has been suggested that this promotes
interspecific competition Interspecific competition, in ecology, is a form of competition in which individuals of ''different'' species compete for the same resources in an ecosystem (e.g. food or living space). This can be contrasted with mutualism, a type of symbiosis. ...
and helps to segregate territories between related species. It appears more likely from later studies that segregation of
sympatric In biology, two closely related species or populations are considered sympatric when they exist in the same geographic area and thus frequently encounter each other. An initially interbreeding population that splits into two or more distinct spe ...
species, other than the blackcap and garden warbler, is due to subtle habitat preferences rather than interspecies aggression.


Breeding

Blackcaps first breed when they are one year old, and are mainly
monogamous Monogamy ( ) is a relationship of two individuals in which they form a mutual and exclusive intimate partnership. Having only one partner at any one time, whether for life or serial monogamy, contrasts with various forms of non-monogamy (e.g. ...
, although both sexes may sometimes deviate from this. A male attracts a female to his territory through songMason (1995) pp. 57–59. and a display involving raising the black crown feathers, fluffing the tail, slow wingbeats, and a short flapping flight. He also builds one or more simple nests (cock nests), usually near his songpost. The final nest, which may be one of the cock nests or built from scratch, is a neat cup of roots, stems and grasses lined with fine material such as hair. The nest is typically deep and across, and is built in the cover of
bramble ''Rubus'' is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, most commonly known as brambles. Fruits of various species are known as raspberries, blackberries, dewberries, and bristleberries. I ...
, scrubs or trees. It is constructed mainly by the female, and may be up to above the ground, although lower than is more typical. The
clutch A clutch is a mechanical device that allows an output shaft to be disconnected from a rotating input shaft. The clutch's input shaft is typically attached to a motor, while the clutch's output shaft is connected to the mechanism that does th ...
is typically 4–6 eggs (range 2–7), which are usually buff with grey and brown blotches and a few dark brown spots. The average size of the egg is .Mason (1995) pp. 63–64. The eggs are incubated for an average of 11 days (range 10–16). Both adults incubate, although only the female stays on the nest at night. The chicks are
altricial Precocial species in birds and mammals are those in which the young are relatively mature and mobile from the moment of birth or hatching. They are normally nidifugous, meaning that they leave the nest shortly after birth or hatching. Altricial ...
, hatching naked and with closed eyes, and are fed by both parents. They
fledge Fledging is the stage in a flying animal's life between egg, hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnera ...
about 11–12 days after hatching, leaving the nest shortly before they are able to fly. They are assisted with feeding for a further two or three weeks. If the nest is threatened, the non-incubating bird gives an alarm call so that the sitting parent and chicks stay still and quiet. A male blackcap may mob a potential predator, or try to lure it away with disjointed runs and flaps on the ground.Mason (1995) pp. 65–68. The blackcap normally raises just one brood, but second nestings are sometimes recorded, particularly in the milder climate of the Mediterranean and the Atlantic islands; triple brooding has been observed once, the female laying a total of 23 eggs in the season. Of eggs laid, 65–93% hatch successfully, and 75–92% of the chicks go on to fledge.Mason (1995) pp. 69–71. The productivity (young fledged per nest) varies with location, level of predation and quality of habitat, but the national figure for the UK was 2.5. The adult annual survival rate is 43% (males 46%, females 29%), and 36% of juveniles live through their first year. The typical life expectancy is two years, but the record is 13 years and 10 months for a bird in the Czech Republic.


Feeding

The blackcap feeds mainly on insects during the breeding season, then switches to fruit in late summer, the change being triggered by an internal
biological rhythm Biological rhythms are repetitive biological processes. Some types of biological rhythms have been described as biological clocks. They can range in frequency from microseconds to less than one repetitive event per decade. Biological rhythms are st ...
. When migrants arrive on their territories they initially take berries,
pollen Pollen is a powdery substance produced by most types of flowers of seed plants for the purpose of sexual reproduction. It consists of pollen grains (highly reduced Gametophyte#Heterospory, microgametophytes), which produce male gametes (sperm ...
and
nectar Nectar is a viscous, sugar-rich liquid produced by Plant, plants in glands called nectaries, either within the flowers with which it attracts pollination, pollinating animals, or by extrafloral nectaries, which provide a nutrient source to an ...
if there are insufficient insects available, then soon switch to their preferred diet. They mainly pick prey off
foliage A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, f ...
and twigs, but may occasionally hover, flycatch or feed on the ground. Blackcaps eat a wide range of
invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...
prey, although
aphids Aphids are small sap-sucking insects in the Taxonomic rank, family Aphididae. Common names include greenfly and blackfly, although individuals within a species can vary widely in color. The group includes the fluffy white Eriosomatinae, woolly ...
are particularly important early in the season, and
flies Flies are insects of the Order (biology), order Diptera, the name being derived from the Ancient Greek, Greek δι- ''di-'' "two", and πτερόν ''pteron'' "wing". Insects of this order use only a single pair of wings to fly, the hindwin ...
,
beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
s and
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder ...
s are also taken in large numbers. Small snails are swallowed whole, since the shell is a source of calcium for the bird's eggs. Chicks are mainly fed soft-bodied insects, fruit only if invertebrates are scarce.Mason (1995) pp. 72–80. In July, the diet switches increasingly to fruit. The
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residue (biochemistry), residues. Proteins perform a vast array of functions within organisms, including Enzyme catalysis, catalysing metab ...
needed for egg-laying and for the chicks to grow is replaced by fruit sugar which helps the birds to fatten for migration. Aphids are still taken while they are available, since they often contain sugars from the plant sap on which they feed. Blackcaps eat a wide range of small fruit, and squeeze out any seeds on a branch before consuming the pulp. This technique makes them an important propagator of
mistletoe Mistletoe is the common name for obligate parasite, obligate parasitic plant, hemiparasitic plants in the Order (biology), order Santalales. They are attached to their host tree or shrub by a structure called the haustorium, through which they ...
. The
mistle thrush The mistle thrush (''Turdus viscivorus'') is a bird common to much of Europe, Palearctic, temperate Asia and North Africa. It is a year-round resident in a large part of its range, but northern and eastern populations bird migration, migrate s ...
, which also favours that plant, is less beneficial since it tends to crush the seeds. Although any suitable fruit may be eaten, some have seasonal or local importance; elder makes up a large proportion of the diet of northern birds preparing for migration, and energy-rich
olive The olive, botanical name ''Olea europaea'' ("European olive"), is a species of Subtropics, subtropical evergreen tree in the Family (biology), family Oleaceae. Originating in Anatolia, Asia Minor, it is abundant throughout the Mediterranean ...
s and lentisc are favoured by blackcaps wintering in the Mediterranean. The continental birds wintering in British gardens rely on provided food, and the major items are bread and fat, each making up around 20% of the diet; one bird survived the whole winter eating only
Christmas cake Christmas cake is a type of cake, often fruitcake, served at Christmas time in many countries. British variations Christmas cake is an English tradition that began as plum porridge. A traditional English Christmas cake is made with moist Zan ...
. Fruit is also eaten, notably
cotoneaster ''Cotoneaster'' is a genus of flowering plants in the rose family, Rosaceae, native to the Palaearctic region (temperate Asia, Europe, north Africa), with a strong concentration of diversity in the genus in the mountains of southwestern China a ...
(41% of the fruit consumed),
ivy ''Hedera'', commonly called ivy (plural ivies), is a genus of 12–15 species of evergreen climbing or ground-creeping woody plants in the family Araliaceae, native to Western Europe, Central Europe, Southern Europe, Macaronesia, northwestern ...
and
honeysuckle Honeysuckles are arching shrubs or Vine#Twining vines, twining vines in the genus ''Lonicera'' () of the family Caprifoliaceae. The genus includes 158 species native to northern latitudes in North America, Eurasia, and North Africa. Widely kno ...
, and apple if available. Some birds have learned to take
peanut The peanut (''Arachis hypogaea''), also known as the groundnut, goober (US), goober pea, pindar (US) or monkey nut (UK), is a legume crop grown mainly for its edible seeds. It is widely grown in the tropics and subtropics by small and large ...
s from feeders.Mason (1995) pp. 80–84. Blackcaps defend good winter food sources in the wild, and at garden feeding stations they repel competitors as large as
starlings Starlings are small to medium-sized passerine (perching) birds known for the often dark, glossy iridescent sheen of their plumage; their complex vocalizations including mimicking; and their distinctive, often elaborate swarming behavior, known ...
and blackbirds. Birds occasionally become tame enough to feed from the hand.


Predators and parasites

Blackcaps are caught by
Eurasian sparrowhawk The Eurasian sparrowhawk (''Accipiter nisus''), also known as the northern sparrowhawk or simply the sparrowhawk, is a small bird of prey in the family Accipitridae. Adult male Eurasian sparrowhawks have bluish grey upperparts and orange-barred ...
s in the breeding range, and by
Eleonora's falcon Eleonora's falcon (''Falco eleonorae'') is a medium-sized falcon. It belongs to the hobby group, a rather close-knit number of similar falcons often considered a subgenus ''Hypotriorchis''. The sooty falcon is sometimes considered its closest rel ...
s on migration.
Eurasian jay The Eurasian jay (''Garrulus glandarius'') is a species of passerine bird in the crow family Corvidae. It has pinkish brown plumage with a black stripe on each side of a whitish throat, a bright blue panel on the upper wing and a black tail. The ...
s and
Eurasian magpie The Eurasian magpie or common magpie (''Pica pica'') is a resident breeding bird throughout the northern part of the Eurasian continent. It is one of several birds in the crow family (corvids) designated magpies, and belongs to the Holarctic r ...
s take eggs and young, as do mammals such as
stoat The stoat (''Mustela erminea''), also known as the Eurasian ermine or ermine, is a species of mustelid native to Eurasia and the northern regions of North America. Because of its wide circumpolar distribution, it is listed as Least Concern on th ...
s,
weasels Weasels are mammals of the genus ''Mustela'' of the family Mustelidae. The genus ''Mustela'' includes the least weasels, polecats, stoats, ferrets, and European mink. Members of this genus are small, active predators, with long and slender b ...
and
squirrel Squirrels are members of the family Sciuridae (), a family that includes small or medium-sized rodents. The squirrel family includes tree squirrels, ground squirrels (including chipmunks and prairie dogs, among others), and flying squirrel ...
s. Domestic cats are the most important predator, possibly killing up to 10% of blackcaps.Mason (1995) pp. 49–53. Blackcaps are occasionally hosts of the
common cuckoo The cuckoo, common cuckoo, European cuckoo or Eurasian cuckoo (''Cuculus canorus'') is a member of the cuckoo order of birds, Cuculiformes, which includes the Geococcyx, roadrunners, the ani (bird), anis and the coucals. This species is a widesp ...
, a
brood parasite Brood may refer to: Nature * Brood, a collective term for offspring * Brooding, the incubation of bird eggs by their parents * Bee brood, the young of a beehive * Individual broods of North American periodical cicadas: ** Brood X, the largest ...
. The level of parasitism is low because the cuckoo's eggs are often rejected. Blackcaps have evolved adaptations which make it difficult for the parasitic species to succeed, despite the cuckoo's tendency to lay eggs which resemble those of their host. Blackcaps are good at spotting alien eggs, and their own eggs are very alike within a clutch, making it easier to spot the intruder. There is, however, considerable variation between different clutches, making it harder for the cuckoo to convincingly mimic a blackcap egg. The open habitat and cup nest of the warbler make it a potential target for the cuckoo; it may have experienced much higher levels of parasitism in the past, and countermeasures would have spread rapidly once they evolved. The only blood parasites found in a study of blackcaps trapped on migration were
protozoa Protozoa (: protozoan or protozoon; alternative plural: protozoans) are a polyphyletic group of single-celled eukaryotes, either free-living or parasitic, that feed on organic matter such as other microorganisms or organic debris. Historically ...
ns from the genera ''
Haemoproteus ''Haemoproteus'' is a genus of alveolates that are parasitic in birds, reptiles, and amphibians. Its name is derived from Greek: ''haima'' 'blood' and ''Proteus'', a sea god that had the power to assume various shapes. The name ''Haemoproteus'' ...
'' and ''
Plasmodium ''Plasmodium'' is a genus of unicellular eukaryotes that are obligate parasites of vertebrates and insects. The life cycles of ''Plasmodium'' species involve development in a Hematophagy, blood-feeding insect host (biology), host which then inj ...
''. The study concluded that 45.5% of the males and 22.7% of the females were affected, but the number of parasites was small, and the ability to store fat for the migration flight was unimpaired. Seventeen strains of '' H. parabelopolskyi'' are found only in the blackcap, and form a
monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent co ...
group; three further members of that group are found only in the garden warbler, and another three occur in the African hill babbler, supporting the shared ancestry of the three bird species. The protozoan '' Isospora ashmoonensis'' was first identified in a blackcap in Egypt. Blackcaps may carry
parasitic worm Parasitic worms, also known as helminths, are a polyphyletic group of large macroparasites; adults can generally be seen with the naked eye. Many are intestinal worms that are soil-transmitted and infect the gastrointestinal tract. Other par ...
s that sometimes kill their hosts. External parasites include
chewing lice The Mallophaga are a possibly paraphyletic section of lice, known as chewing lice, biting lice, or bird lice, containing more than 3000 species. These lice are external parasites that feed mainly on birds, although some species also feed on mamma ...
and
feather mite Feather mites are the members of diverse mite Superfamily (zoology), superfamilies: * superorder Acariformes ** Psoroptidia *** Analgoidea *** Freyanoidea *** Pterolichoidea * superorder Parasitiformes ** Dermanyssoidea They are ectoparasites on ...
s. The latter do little damage, although heavy infestations cause individual tail feathers to develop asymmetrically.


In culture

Aristotle Aristotle (; 384–322 BC) was an Ancient Greek philosophy, Ancient Greek philosopher and polymath. His writings cover a broad range of subjects spanning the natural sciences, philosophy, linguistics, economics, politics, psychology, a ...
, in his ''
History of Animals ''History of Animals'' (, ''Ton peri ta zoia historion'', "Inquiries on Animals"; , "History of Animals") is one of the major texts on biology by the ancient Greek philosopher Aristotle. It was written in sometime between the mid-fourth centur ...
'', considered that the
garden warbler The garden warbler (''Sylvia borin'') is a common and widespread small bird that breeds in most of Europe and in the Palearctic to western Siberia. It is a plain, long-winged and long-tailed typical warbler with brown upperparts and dull white ...
eventually
metamorphosed Metamorphic rocks arise from the transformation of existing rock to new types of rock in a process called metamorphism. The original rock (protolith) is subjected to temperatures greater than and, often, elevated pressure of or more, causi ...
into a blackcap.Arnason & Murphy (2001) p. 131. The blackcap's song has led to it being described as the "mock nightingale" or "country nightingale", and
John Clare John Clare (13 July 1793 – 20 May 1864) was an English poet. The son of a farm labourer, he became known for his celebrations of the English countryside and his sorrows at its disruption. His work underwent major re-evaluation in the late 20t ...
, in "The March Nightingale" describes the listener as believing that the rarer species has arrived prematurely. "He stops his own and thinks the nightingale/Hath of her monthly reckoning counted wrong".Cocker (2005) pp. 374–376. The song is also the topic of Italian poet
Giovanni Pascoli Giovanni Placido Agostino Pascoli (; 31 December 1855 – 6 April 1912) was an Italian poet, classical scholar and an emblematic figure of Italian literature in the late nineteenth century. Alongside Gabriele D'Annunzio, he was one of the grea ...
's "La Capinera" he Blackcap
Giovanni Verga Giovanni Carmelo Verga di Fontanabianca (; 2 September 1840 – 27 January 1922) was an Italian Literary realism, realist (''Verismo (literature), verista'') writer. His novels ''I Malavoglia'' (1881) and ''Mastro-don Gesualdo'' (1889) are widel ...
's 1871 novel ''Storia di una capinera'', according to its author, was inspired by a story of a blackcap trapped and caged by children. The bird, silent and pining for its lost freedom, eventually dies. In the book, a nun evacuated from her convent by
cholera Cholera () is an infection of the small intestine by some Strain (biology), strains of the Bacteria, bacterium ''Vibrio cholerae''. Symptoms may range from none, to mild, to severe. The classic symptom is large amounts of watery diarrhea last ...
falls in love with a family friend, only to have to return to her confinement when the disease wanes. The novel was adapted as films of the same name in 1917, 1943 and 1993.Cardillo (1987) p. 31.Chiti ''et al'' (2002) p. 351. The last version was directed by
Franco Zeffirelli Gian Franco Corsi Zeffirelli (; 12 February 1923 – 15 June 2019) was an Italian stage and film director, producer, production designer and politician. He was one of the most significant opera and theatre directors of the post–World War II e ...
, and its English-language version was retitled as '' Sparrow''.Chiti ''et al'' (2002) p. 231. In ''
Saint François d'Assise ''Saint François d'Assise : Scènes Franciscaines'' (English: Franciscan Scenes of Saint Francis of Assisi), or simply ''Saint François d'Assise'', is an opera in three acts and eight scenes by French composer Olivier Messiaen, who was also ...
'', an opera by
Messiaen Olivier Eugène Prosper Charles Messiaen (, ; ; 10 December 1908 – 27 April 1992) was a French composer, organist, and ornithology, ornithologist. One of the major composers of the 20th-century classical music, 20th century, he was also an ou ...
, the orchestration is based on bird song. St Francis himself is represented by the blackcap.Hill (1995) p. 259. In Finland, a caged blackcap is featured in a Christmas song ''Sylvian Joululaulu'' (compare with the Latin name ''Sylvia atricapilla''), and is said to represent Finland before it claimed independence from Russia in 1917. Folk names for the blackcap often refer to its most obvious plumage feature (black-headed peggy, King Harry black cap and coal hoodie) or to its song, as in the "nightingale" names above. Other old names are based on its choice of nesting material (Jack Straw, hay bird, hay chat and hay Jack).Mason (1995) p. 13. There is a tradition of the
Royal Navy The Royal Navy (RN) is the naval warfare force of the United Kingdom. It is a component of His Majesty's Naval Service, and its officers hold their commissions from the King of the United Kingdom, King. Although warships were used by Kingdom ...
's
Fleet Air Arm The Fleet Air Arm (FAA) is the naval aviation component of the United Kingdom's Royal Navy (RN). The FAA is one of five :Fighting Arms of the Royal Navy, RN fighting arms. it is a primarily helicopter force, though also operating the Lockhee ...
bases being named for birds. A former base near Stretton in Cheshire was called HMS ''Blackcap''.Ferguson (2008) pp. 202–203.


Status

The blackcap has a very large range, and its population in Europe is estimated at 41–65 million breeding pairs. Allowing for birds breeding in Africa and Asia, the total population estimate is between 101 and 161 million individuals. It is therefore classified by the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the stat ...
as being of
least concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
. Blackcaps and other small birds are illegally trapped and hunted in large numbers in Mediterranean countries, particularly in Lebanon, Syria, Palestine, Malta, Libya, Egypt and Cyprus, where they are considered as a
delicacy A delicacy is a rare food item that is considered highly desirable, sophisticated, or peculiarly distinctive within a given culture or region. A delicacy may have an unusual flavor or be expensive compared to everyday foods. Delicacies va ...
. Despite hunting and natural hazards, the European population of the blackcap has been rising for several decades as the range extends northwards, for example to Scotland and Denmark. There are occasional nesting records from outside the main range, such as in northern Israel and the
Faroes The Faroe Islands ( ) (alt. the Faroes) are an archipelago in the North Atlantic Ocean and an autonomous territory of the Danish Realm, Kingdom of Denmark. Located between Iceland, Norway, and the United Kingdom, the islands have a populat ...
, and wandering birds may appear further afield in Iceland or on the islands of Arctic Russia. In the Baltic, the spread of the blackcap appears to have been helped by the availability of territories formerly occupied by the declining
barred warbler The barred warbler (''Curruca nisoria'') is a typical warbler which breeds across temperate regions of central and eastern Europe and western and central Asia. This passerine bird is strongly migratory, and winters in tropical eastern Africa.De ...
. Blackcap (Sylvia atricapilla) male ringed Malta.jpg, ringed male, Malta Blackcap (Sylvia atricapilla) female ringed Malta.jpg, ringed female, Malta


Notes


References


Cited texts

* * * * * * * * * * * * * * * * *


External links


Blackcap videos
on the Internet Bird Collection * Madeira Birds
Blackcap
including images of melanistic birds
Ageing and sexing (PDF; 4.5 MB) by Javier Blasco-Zumeta & Gerd-Michael Heinze



Blackcap male song video, Lodz(Poland)
{{Authority control Sylvia (bird) Birds of Eurasia Birds of Europe Birds of Macaronesia Wintering birds of Africa Birds described in 1758 Taxa named by Carl Linnaeus