Black Leopard
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A black panther is the
melanistic Melanism is the congenital excess of melanin in an organism resulting in dark pigment. Pseudomelanism, also called abundism, is another variant of pigmentation, identifiable by dark spots or enlarged stripes, which cover a large part of the bod ...
colour variant of the
leopard The leopard (''Panthera pardus'') is one of the five extant cat species in the genus ''Panthera''. It has a pale yellowish to dark golden fur with dark spots grouped in rosettes. Its body is slender and muscular reaching a length of with a ...
(''Panthera pardus'') and the
jaguar The jaguar (''Panthera onca'') is a large felidae, cat species and the only extant taxon, living member of the genus ''Panthera'' that is native to the Americas. With a body length of up to and a weight of up to , it is the biggest cat spe ...
(''Panthera onca''). Black panthers of both species have excess black
pigment A pigment is a powder used to add or alter color or change visual appearance. Pigments are completely or nearly solubility, insoluble and reactivity (chemistry), chemically unreactive in water or another medium; in contrast, dyes are colored sub ...
s, but their typical rosettes are also present. They have been documented mostly in
tropical forest Tropical forests are forested ecoregions with tropical climates – that is, land areas approximately bounded by the Tropic of Cancer, tropics of Cancer and Tropic of Capricorn, Capricorn, but possibly affected by other factors such as prevailing ...
s, with black leopards in Africa and Asia, and black jaguars in Central and South America. Melanism is caused by a
recessive allele In genetics, dominance is the phenomenon of one variant (allele) of a gene on a chromosome masking or overriding the effect of a different variant of the same gene on the other copy of the chromosome. The first variant is termed dominant and ...
in the leopard, and by a
dominant allele In genetics, dominance is the phenomenon of one variant (allele) of a gene on a chromosome masking or overriding the effect of a different variant of the same gene on the other copy of the chromosome. The first variant is termed dominant and ...
in the jaguar.


Leopard

In 1788, Jean-Claude Delamétherie described a black leopard that was kept in the
Tower of London The Tower of London, officially His Majesty's Royal Palace and Fortress of the Tower of London, is a historic citadel and castle on the north bank of the River Thames in central London, England. It lies within the London Borough of Tower Hamle ...
and had been brought from
Bengal Bengal ( ) is a Historical geography, historical geographical, ethnolinguistic and cultural term referring to a region in the Eastern South Asia, eastern part of the Indian subcontinent at the apex of the Bay of Bengal. The region of Benga ...
. In 1794, Friedrich Albrecht Anton Meyer proposed the
scientific name In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...
''Felis fusca'' for this cat, the
Indian leopard The Indian leopard (''Panthera pardus fusca'') is a subspecies of the leopard (''P. pardus''). It is widely distributed on the Indian subcontinent. It is threatened by illegal trade of skins and body parts, and persecution due to human-leopard ...
(''P. p. fusca''). In 1809,
Georges Cuvier Jean Léopold Nicolas Frédéric, baron Cuvier (23 August 1769 – 13 May 1832), known as Georges Cuvier (; ), was a French natural history, naturalist and zoology, zoologist, sometimes referred to as the "founding father of paleontology". Cuv ...
described a black leopard kept in the
Ménagerie du Jardin des plantes The Ménagerie du Jardin des Plantes () is a zoo in Paris, France, belonging to the botanical garden Jardin des Plantes. Founded in 1794, largely with animals brought from the royal zoo of the Palace of Versailles, abandoned because of the French ...
that had been brought from
Java Java is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea (a part of Pacific Ocean) to the north. With a population of 156.9 million people (including Madura) in mid 2024, proje ...
. Cuvier proposed the name ''Felis melas'', the Javan leopard (''P. p. melas''). By the late 19th century, the occurrence of black and spotted leopard cubs in the same
litter Litter consists of waste products that have been discarded incorrectly, without consent, at an unsuitable location. The waste is objects, often man-made, such as aluminum cans, paper cups, food wrappers, cardboard boxes or plastic bottles, but ...
had been repeatedly recorded in
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
. Black leopards were thought to be more common in
Travancore The kingdom of Travancore (), also known as the kingdom of Thiruvithamkoor () or later as Travancore State, was a kingdom that lasted from until 1949. It was ruled by the Travancore Royal Family from Padmanabhapuram, and later Thiruvanan ...
and in the hills of
southern India South India, also known as Southern India or Peninsular India, is the southern part of the Deccan Peninsula in India encompassing the states of Andhra Pradesh, Karnataka, Kerala, Tamil Nadu and Telangana as well as the union territories of ...
than in other parts of the country. Black leopards were also frequently encountered in southern
Myanmar Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and has ...
. By 1929, the
Natural History Museum, London The Natural History Museum in London is a museum that exhibits a vast range of specimens from various segments of natural history. It is one of three major museums on Exhibition Road in South Kensington, the others being the Science Museum (Lo ...
had skins of black leopards collected in
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
,
Nepal Nepal, officially the Federal Democratic Republic of Nepal, is a landlocked country in South Asia. It is mainly situated in the Himalayas, but also includes parts of the Indo-Gangetic Plain. It borders the Tibet Autonomous Region of China Ch ...
,
Assam Assam (, , ) is a state in Northeast India, northeastern India, south of the eastern Himalayas along the Brahmaputra Valley, Brahmaputra and Barak River valleys. Assam covers an area of . It is the second largest state in Northeast India, nor ...
and
Kanara Kanara or Canara, also known as Karāvali, is the historically significant stretch of land situated by the southwestern Konkan coast of India, alongside the Arabian Sea in the present-day Indian state of Karnataka. The subregion comprises thr ...
in India. Black leopards were thought to be common on the
Malay Peninsula The Malay Peninsula is located in Mainland Southeast Asia. The landmass runs approximately north–south, and at its terminus, it is the southernmost point of the Asian continental mainland. The area contains Peninsular Malaysia, Southern Tha ...
and on Java. A black
African leopard The African leopard (''Panthera pardus pardus'') is the nominate subspecies of the leopard, native to many countries in Africa. It is widely distributed in most of sub-Saharan Africa, but the historical range has been Habitat fragmentation, fragm ...
(''P. p. pardus'') was sighted in the
alpine zone Alpine tundra is a type of natural region or biome that does not contain trees because it is at high elevation, with an associated alpine climate, harsh climate. As the latitude of a location approaches the poles, the threshold elevation for alp ...
of
Mount Kenya Mount Kenya (Meru people, Meru: ''Kĩrĩmaara,'' Kikuyu people, Kikuyu: ''Kĩrĩnyaga'', Kamba language, Kamba: ''Ki nyaa'', Embu language, Embu: ''Kĩ nyaga'') is an extinct volcano in Kenya and the Highest mountain peaks of Africa, second-highe ...
in the winter of 1989–1990. In Kenya's
Laikipia County Laikipia County is one of the 47 counties of Kenya, located on the Equator in the former Rift Valley Province. Laikipia is a cosmopolitan county and is listed as county number 31. The county has two major urban centres: Nanyuki to the southea ...
, a black leopard was photographed by a
camera trap A camera trap is a camera that is automatically triggered by motion in its vicinity, like the presence of an animal or a human being. It is typically equipped with a motion sensor—usually a passive infrared (PIR) sensor or an active infrared ...
in 2007; in 2018, a female subadult black leopard was repeatedly recorded together with a spotted leopard about farther east in a grassland. In India's Western Ghats, black leopards were sighted and photographed in 2010 and 2012 in the Kas Plateau Reserved Forest, and in
Bhadra Wildlife Sanctuary Bhadra Wildlife Sanctuary is a Protected areas of India, protected area and tiger reserves of India, tiger reserve as part of the Project Tiger, situated in Chikkamagaluru district, south of Bhadravati, Karnataka, Bhadravathi city, 20 km from ...
in 2012. In 2015, a dead black leopard was found on a highway near Satara in
Maharashtra Maharashtra () is a state in the western peninsular region of India occupying a substantial portion of the Deccan Plateau. It is bordered by the Arabian Sea to the west, the Indian states of Karnataka and Goa to the south, Telangana to th ...
. In May 2012, a black leopard was photographed at an elevation of in Nepal's
Kanchenjunga Conservation Area Kanchenjunga Conservation Area is a protected area in the Himalayas of eastern Nepal that was established in 1997. It covers in the Taplejung District and comprises two peaks of Kanchenjunga. In the north it adjoins the Qomolangma National Nat ...
. At least one black leopard was photographed in mixed
deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...
forest in Thailand's
Kaeng Krachan National Park Kaeng Krachan National Park (, , ) is the largest List of national parks of Thailand, national park of Thailand. It is on the border with Myanmar (Burma), contiguous with the Tanintharyi Nature Reserve. It is a popular park owing to its proximit ...
during a one-year-long camera trapping survey from 2003 to 2004. In 2009, black leopards were photographed more often than spotted leopards in Kui Buri National Park. Most leopards recorded at 16 sites south of the
Kra Isthmus The Kra Isthmus (, ; ), also called the Isthmus of Kra in Thailand, is the narrowest part of the Malay Peninsula. The western part of the isthmus belongs to Ranong Province and the eastern part to Chumphon Province, both in Southern Thailan ...
between 1996 and 2009 were black, indicating a near- fixation of melanism in Peninsular Malaysia. In 2019, a black individual was photographed outside a protected area in Jeli District. Both black and spotted leopards were recorded in Gunung Gede Pangrango National Park in
West Java West Java (, ) is an Indonesian Provinces of Indonesia, province on the western part of the island of Java, with its provincial capital in Bandung. West Java is bordered by the province of Banten and the country's capital region of Jakarta to t ...
between 2005 and 2017. Frequency of melanism appears to be approximately 11% over the leopard's range. Data on the distribution of leopard populations indicates that melanism occurs in five subspecies in the wild: the Indian leopard, Javan leopard, African leopard, Indochinese leopard (''P. p. delacouri'') and Sri Lankan leopard (''P. p. kotiya''). Based on records from camera traps, melanistic leopards occur foremost in
tropical and subtropical moist broadleaf forests Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature (WWF). Description TSMF is generally found in la ...
. Melanism in the leopard is conferred by a
recessive allele In genetics, dominance is the phenomenon of one variant (allele) of a gene on a chromosome masking or overriding the effect of a different variant of the same gene on the other copy of the chromosome. The first variant is termed dominant and ...
. It is thought that melanism confers a selective advantage under certain conditions since it is more common in regions of dense forest, where light levels are lower. Preliminary studies also suggest that melanism might be linked to beneficial mutations in the
immune system The immune system is a network of biological systems that protects an organism from diseases. It detects and responds to a wide variety of pathogens, from viruses to bacteria, as well as Tumor immunology, cancer cells, Parasitic worm, parasitic ...
. The typical spots and rosettes are present but hidden due to the excess melanin. The
taxonomic 280px, Generalized scheme of taxonomy Taxonomy is a practice and science concerned with classification or categorization. Typically, there are two parts to it: the development of an underlying scheme of classes (a taxonomy) and the allocation ...
status of captive black leopards and the extent of hybridization between the Javan leopard and other leopard subspecies is uncertain. Therefore, coordinated breeding programs for black leopards do not exist in European and North American zoos. Black leopards occupy space needed for breeding endangered leopard subspecies and are not included within the North American
Species Survival Plan The American Species Survival Plan or SSP program was developed in 1981 by the (American) Association of Zoos and Aquariums to help ensure the survival of selected species in zoos and aquariums, most of which are threatened or endangered in the w ...
. A black Amur leopard (''P. p. orientalis'') was exhibited at the
San Diego Zoo The San Diego Zoo is a zoo in San Diego, California, United States, located in Balboa Park (San Diego), Balboa Park. It began with a collection of animals left over from the 1915 Panama–California Exposition that were brought together by its ...
in 2017. A pseudo-melanistic leopard has a normal background color, but the spots are more densely packed than normal, and merge to obscure the golden-brown background color. Any spots on the flanks and limbs that have not merged into the mass of swirls and stripes are unusually small and discrete, rather than forming rosettes. The face and underparts are paler and dappled, like those of ordinary spotted leopards.


Jaguar

In 1801,
Félix de Azara Félix de Azara y Perera (18 May 1742 – 20 October 1821) was a Spanish military officer, naturalist, and engineer. Life Félix de Azara y Perera was born on 18 May 1746 in Barbuñales, Aragon. He joined the army and attended a Spanish milita ...
described a black jaguar observed by local people near the
Paraná River The Paraná River ( ; ; ) is a river in south-central South America, running through Brazil, Paraguay, and Argentina for some ."Parana River". Encyclopædia Britannica. Encyclopædia Britannica Online. Encyclopædia Britannica Inc., 2012. Web. ...
in Paraguay. In 2004, a female black jaguar was recorded in Mexico's
Sierra Madre Occidental The Sierra Madre Occidental is a major mountain range system of the North American Cordillera, that runs northwest–southeast through northwestern and western Mexico, and along the Gulf of California. The Sierra Madre is part of the American C ...
. In 2009, a black jaguar was photographed by a camera trap for the first time in Costa Rica's Alberto Manuel Brenes Biological Reserve. In
Barbilla National Park Barbilla National Park is a national park in the Caribbean La Amistad Conservation Area of Costa Rica located on the eastern slopes of the Cordillera de Talamanca. It protects forests as well as Laguna Ayil and Cerro Tigre and the Dantas River wat ...
, black jaguars were recorded in 2013. In the mountains of the
Cordillera de Talamanca The Cordillera de Talamanca is a mountain range that lies in the southeast half of Costa Rica and the far west of Panama. Much of the range and the area around it is included in La Amistad International Park, which also is shared between the two ...
, 104 records of jaguars were obtained between 2010 and 2019; 26 of them showed melanistic jaguars. In eastern Panama, black jaguars were repeatedly photographed in the Mamoní River Valley between 2016 and 2018, mostly in
primary forest An old-growth forest or primary forest is a forest that has developed over a long period of time without Disturbance (ecology), disturbance. Due to this, old-growth forests exhibit unique ecological features. The Food and Agriculture Organizati ...
. Five black jaguars have been monitored in the
Várzea forest A várzea forest is a seasonal floodplain forest inundated by whitewater rivers that occurs in the Amazon biome. Until the late 1970s, the definition was less clear and várzea was often used for all periodically flooded Amazonian forests. Althoug ...
of
Mamirauá Sustainable Development Reserve The Mamirauá Sustainable Development Reserve () in the Brazilian state of Amazonas (Brazilian state), Amazonas, near the city of Tefé, is a reserve near the village of Boca do Mamirauá. It includes mostly Amazon rainforest, Amazonian flooded f ...
in the Brazilian State of Amazonas between 2003 and 2018. Black jaguars were also recorded in the Brazilian
Pará Pará () is a Federative units of Brazil, state of Brazil, located in northern Brazil and traversed by the lower Amazon River. It borders the Brazilian states of Amapá, Maranhão, Tocantins (state), Tocantins, Mato Grosso, Amazonas (Brazilian st ...
state. Melanism in the jaguar is caused by deletions in the
melanocortin 1 receptor The melanocortin 1 receptor (MC1R), also known as melanocyte-stimulating hormone receptor (MSHR), melanin-activating peptide receptor, or melanotropin receptor, is a G protein–coupled receptor that binds to a class of pituitary peptide hormon ...
gene and conferred by a
dominant allele In genetics, dominance is the phenomenon of one variant (allele) of a gene on a chromosome masking or overriding the effect of a different variant of the same gene on the other copy of the chromosome. The first variant is termed dominant and ...
.


In the United States

There have been numerous reported black panther sightings in the
New Orleans New Orleans (commonly known as NOLA or The Big Easy among other nicknames) is a Consolidated city-county, consolidated city-parish located along the Mississippi River in the U.S. state of Louisiana. With a population of 383,997 at the 2020 ...
area since late 2010. Recent photographs are still under examination by the Louisiana Department of Wildlife & Fisheries. In Florida, a few melanistic
bobcat The bobcat (''Lynx rufus''), also known as the wildcat, bay lynx, or red lynx, is one of the four extant species within the medium-sized wild cat genus '' Lynx''. Native to North America, it ranges from southern Canada through most of the c ...
s have been captured; these have apparently been mistaken for
Florida panther The Florida panther is a North American cougar (''P. c. couguar'') population in South Florida. It lives in pinelands, tropical hardwood hammocks and mixed freshwater swamp forests. Its range includes the Big Cypress National Preserve, Everglade ...
s. Ulmer (1941) presents photographs and descriptions of two animals captured in Martin County in 1939 and 1940. In the photographs, they appear black, and one of the hunters called them black.
The Academy specimen, upon close examination, is far from black. The most heavily pigmented portions are the crown and dorsal area. In most lights these areas appear black, but at certain angles the dorsal strip has a decidedly mahogany tint. The mahogany coloring becomes lighter and richer on the sides. The underparts are lightest, being almost ferruginous in color. The chin, throat and cheeks are dark chocolate-brown, but the facial stripes can be seen clearly. The limbs are dark mahogany. In certain lights the typical spot-pattern of the Florida bobcat can be distinctly seen on the side, underparts and limbs. The Bronx Park animal appears darker and the spots are not visible, although the poor light in the quarantine cage may have been the reason.


In Australia

Black panther sightings are frequently recorded in rural Victoria and
New South Wales New South Wales (commonly abbreviated as NSW) is a States and territories of Australia, state on the Eastern states of Australia, east coast of :Australia. It borders Queensland to the north, Victoria (state), Victoria to the south, and South ...
The ''Animal X Natural Mysteries Unit'' led an investigation into the phantom panther. Mike Williams, a local researcher, said he had sent scat and hair found by locals to labs for analysis, which identified it as scat from dogs that had feasted on
swamp wallaby The swamp wallaby (''Wallabia bicolor'') is a small macropod marsupial of eastern Australia. This wallaby is also commonly known as the black wallaby, with other names including black-tailed wallaby, fern wallaby, black pademelon, stinker (in Qu ...
, and hair from a domestic cat. Williams said he also had leopard scat and hair collected from a private zoo tested by one of the same labs, but that these samples came back with the same results of dog scat and domestic cat hair. The lab used was not identified in the episode.


Cougar

There is no authenticated case of a truly melanistic
cougar The cougar (''Puma concolor'') (, ''Help:Pronunciation respelling key, KOO-gər''), also called puma, mountain lion, catamount and panther is a large small cat native to the Americas. It inhabits North America, North, Central America, Cent ...
. No specimen has been photographed or killed in the wild, nor has it ever been bred in captivity. Unconfirmed sightings known as the "North American black panther" are currently attributed to errors in species identification by non-experts, and by the
mimetic Mimesis (; , ''mīmēsis'') is a term used in literary criticism and philosophy that carries a wide range of meanings, including ''imitatio'', imitation, Similarity (philosophy), similarity, receptivity, representation (arts), representation, m ...
exaggeration of size.


See also

*
Black cat A black cat is a Cat, domestic cat with black fur. They may be a specific Purebred, breed, or a common domestic cat of no particular or mixed breed. Most black cats have golden iris (anatomy), irises due to their high melanin pigment content. Bl ...
* Black tiger * White panther *
Bagheera Bagheera ( / ''Baghīrā'') is a fictional character in Rudyard Kipling's Mowgli stories in '' The Jungle Book'' (coll. 1894) and '' The Second Jungle Book'' (coll. 1895). He is a black panther ( melanistic Indian leopard) who serves as frie ...
, a black leopard in Rudyard Kipling's 1894 short story collection ''
The Jungle Book ''The Jungle Book'' is an 1894 collection of stories by the English author Rudyard Kipling. Most of the characters are animals such as Shere Khan the tiger and Baloo the bear, though a principal character is the boy or "man-cub" Mowgli, who ...
'' *
Black Panther A black panther is the Melanism, melanistic colour variant of the leopard (''Panthera pardus'') and the jaguar (''Panthera onca''). Black panthers of both species have excess black pigments, but their typical Rosette (zoology), rosettes are al ...
, a Marvel superhero and king of
Wakanda Wakanda (), officially the Kingdom of Wakanda, is a fictional country appearing in American comic books published by Marvel Comics. Created by Stan Lee and Jack Kirby, the country first appeared in ''Fantastic Four (comic book), Fantastic Four'' ...


References


External links

* * * {{DEFAULTSORT:Black panther Felid color morphs Leopards Jaguars