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Bison are large
bovine Bovines (subfamily Bovinae) comprise a diverse group of 10 genera of medium to large-sized ungulates, including cattle, bison, African buffalo, water buffalos, and the four-horned and spiral-horned antelopes. The evolutionary relationship be ...
s in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
''Bison'' (Greek: "wild ox" (bison)) within the tribe
Bovini The tribe Bovini, or wild cattle, are medium to massive bovines that are native to North America, Eurasia, and Africa. These include the enigmatic, antelope-like saola, the African and Asiatic buffalos, and a clade that consists of bison and the ...
. Two
extant Extant is the opposite of the word extinct. It may refer to: * Extant hereditary titles * Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English * Extant taxon, a taxon which is not extinct, ...
and numerous extinct species are recognised. Of the two surviving species, the
American bison The American bison (''Bison bison'') is a species of bison native to North America. Sometimes colloquially referred to as American buffalo or simply buffalo (a different clade of bovine), it is one of two extant species of bison, alongside the ...
, ''B. bison'', found only in North America, is the more numerous. Although colloquially referred to as a buffalo in the
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
and
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...
, it is only distantly related to the
true buffalo Bubalina is a subtribe of wild cattle that includes the various species of true buffalo. Species include the African buffalo, the anoas, and the wild water buffalo (including the domesticated variant water buffalo). Buffaloes can be found na ...
. The North American species is composed of two subspecies, the
Plains bison The Plains bison (''Bison bison bison'') is one of two subspecies/ecotypes of the American bison, the other being the wood bison (''B. b. athabascae''). A natural population of Plains bison survives in Yellowstone National Park (the Yellowstone P ...
, ''B. b. bison'', and the
wood bison The wood bison (''Bison bison athabascae'') or mountain bison (often called the wood buffalo or mountain buffalo), is a distinct northern subspecies or ecotype of the American bison. Its original range included much of the boreal forest regions ...
, ''B. b. athabascae'', which is the namesake of
Wood Buffalo National Park Wood Buffalo National Park is the largest national park of Canada at . It is located in northeastern Alberta and the southern Northwest Territories. Larger in area than Switzerland, it is the second-largest national park in the world. The park ...
in Canada. A third subspecies, the eastern bison (''B. b. pennsylvanicus'') is no longer considered a valid
taxon In biology, a taxon ( back-formation from '' taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular n ...
, being a junior synonym of ''B. b. bison''. References to "woods bison" or "wood bison" from the eastern United States refer to this subspecies, not ''B. b. athabascae'', which was not found in the region. The
European bison The European bison (''Bison bonasus'') or the European wood bison, also known as the wisent ( or ), the zubr (), or sometimes colloquially as the European buffalo, is a European species of bison. It is one of two extant species of bison, alon ...
, ''B. bonasus'', or wisent, or zubr, or colloquially European buffalo, is found in Europe and the
Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia (country), Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range ...
, reintroduced after being
extinct in the wild A species that is extinct in the wild (EW) is one that has been categorized by the International Union for Conservation of Nature as known only by living members kept in captivity or as a naturalized population outside its historic range due t ...
. While bison species have been traditionally classified in their own
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
, modern genetics indicates that they are nested within the genus ''
Bos ''Bos'' (from Latin '' bōs'': cow, ox, bull) is the genus of wild and domestic cattle. ''Bos'' is often divided into four subgenera: ''Bos'', ''Bibos'', ''Novibos'', and ''Poephagus'', but including these last three divisions within the gen ...
,'' which includes, among others, cattle,
yak The domestic yak (''Bos grunniens''), also known as the Tartary ox, grunting ox or hairy cattle, is a species of long-haired domesticated cattle found throughout the Himalayan region of the Indian subcontinent, the Tibetan Plateau, Kachin St ...
s and
gaur The gaur (''Bos gaurus''; ), also known as the Indian bison, is a bovine native to South Asia and Southeast Asia, and has been listed as Vulnerable on the IUCN Red List since 1986. The global population was estimated at a maximum of 21,000 ...
, being most closely related to yaks. Bison are sometimes bred with domestic cattle and produce offspring called
beefalo Beefalo constitute a hybrid offspring of domestic cattle (''Bos taurus''), usually a male in managed breeding programs, and the American bison (''Bison bison''), usually a female in managed breeding programs. The breed was created to combine the ...
or zubron.


Description

The American bison and the European bison (wisent) are the largest surviving terrestrial animals in North America and Europe. They are typical
artiodactyl The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing poste ...
(cloven hooved) ungulates, and are similar in appearance to other bovines such as cattle and true buffalo. They are broad and muscular with shaggy coats of long hair. Adults grow up to in height and in length for American bisonGennady G. Boeskorov, Olga R. Potapova, Albert V. Protopopov, Valery V. Plotnikov, Larry D. Agenbroad, Konstantin S. Kirikov, Innokenty S. Pavlov, Marina V. Shchelchkova, Innocenty N. Belolyubskii, Mikhail D. Tomshin, Rafal Kowalczyk, Sergey P. Davydov, Stanislav D. Kolesov, Alexey N. Tikhonov, Johannes van der Plicht, 2016, The Yukagir Bison: The exterior morphology of a complete frozen mummy of the extinct steppe bison, ''Bison priscus'' from the early Holocene of northern Yakutia, Russia, pp.7,
Quaternary International ''Quaternary International'' is a peer-reviewed scientific journal on quaternary science published by Elsevier on behalf of the International Union for Quaternary Research. The journal was established in 1989 and covers full spectrum of the physica ...
, Vol.406 (25 June 2016), Part B, pp.94-110

"American Bison", The National Wildlife Federation.
and up to in heightSemenov U.A. of World Wide Fund for Nature, WWF-Russia, 2014, "The Wisents of Karachay-Cherkessia", Proceedings of the
Sochi National Park Sochi National Park (russian: Сочинский национальный парк, also Sochinsky National Park) is Russia's oldest national park, established on May 5, 1983. It is located in the Western Caucasus, near the city of Sochi, in Southe ...
(8), pp.23-24, , KMK Scientific Press
and in length for European bison.Hendricks, K
"''Bison bonasus'', European bison"
Animal Diversity Web, 2013.
American bison can weigh from around and European bison can weigh from . European bison tend to be taller than American bison. Bison are nomadic grazers and travel in herds. The bulls leave the herds of females at two or three years of age, and join a herd of males, which are generally smaller than female herds. Mature bulls rarely travel alone. Towards the end of the summer, for the reproductive season, the sexes necessarily commingle. American bison are known for living in the Great Plains, but formerly had a much larger range, including much of the eastern United States and parts of Mexico. Both species were hunted close to
extinction Extinction is the termination of a kind of organism or of a group of kinds ( taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed ...
during the 19th and 20th centuries, but have since rebounded. The wisent in part owes its survival to the Chernobyl disaster, as the
Chernobyl Exclusion Zone The Chernobyl Nuclear Power Plant Zone of Alienation, Belarusian: Хона адчужэння Чарнобыльскай АЭС, ''Zona adčužennia Čarnobyĺskaj AES'', russian: Зона отчуждения Чернобыльской АЭС, ...
has become a kind of wildlife preserve for wisent and other rare megafauna such as the
Przewalski's horse Przewalski's horse (, , (Пржевальский ), ) (''Equus ferus przewalskii'' or ''Equus przewalskii''), also called the takhi, Mongolian wild horse or Dzungarian horse, is a rare and endangered horse originally native to the steppes of Ce ...
, though poaching has become a threat in recent years. The American Plains bison is no longer listed as
endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inv ...
, but this does not mean the species is secure. Genetically pure ''B. b. bison'' currently number only about 20,000, separated into fragmented herds—all of which require active conservation measures. The wood bison is on the endangered species list in Canada and is listed as threatened in the United States, though numerous attempts have been made by
beefalo Beefalo constitute a hybrid offspring of domestic cattle (''Bos taurus''), usually a male in managed breeding programs, and the American bison (''Bison bison''), usually a female in managed breeding programs. The breed was created to combine the ...
ranchers to have it entirely removed from the
Endangered Species List On 29 January 2010, the IUCN Red List of Threatened Species identified 5220 (2754 animals, 1 fungus, 2464 plant, 1 protist) endangered species, subspecies and varieties, stocks and sub-populations. For IUCN lists of endangered species by kin ...
. Although superficially similar, physical and behavioural differences exist between the American and European bison. The American species has 15 ribs, while the European bison has 14. The American bison has four lumbar vertebrae, while the European has five. (The difference in this case is that what would be the first lumbar vertebra has ribs attached to it in American bison and is thus counted as the 15th thoracic vertebra, compared to 14 thoracic vertebrae in wisent.) Adult American bison are less slim in build and have shorter legs. American bison tend to graze more, and
browse Browsing is a kind of orienting strategy. It is supposed to identify something of relevance for the browsing organism. When used about human beings it is a metaphor taken from the animal kingdom. It is used, for example, about people browsing o ...
less than their European relatives. Their anatomies reflect this behavioural difference; the American bison's head hangs lower than the European's. The body of the American bison is typically hairier, though its tail has less hair than that of the European bison. The horns of the European bison point through the plane of their faces, making them more adept at fighting through the interlocking of horns in the same manner as domestic cattle, unlike the American bison, which favours butting. American bison are more easily tamed than their European cousins, and breed with domestic cattle more readily.


Evolution and genetic history

The bovine tribe (Bovini) split about 5 to 10 million years ago into the buffalos (''
Bubalus ''Bubalus'' is a genus of Asiatic bovines that was proposed by Charles Hamilton Smith in 1827. ''Bubalus'' and '' Syncerus'' form the subtribe Bubalina, the true buffaloes. The International Code of Zoological Nomenclature and classificatio ...
'' and ''
Syncerus ''Syncerus'' is a genus of African bovid that contains the living Cape buffalo ''(Syncerus caffer