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''Pinus muricata'', the bishop pine, is a
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. ''World Flora Online'' accepts 134 species-rank taxa (119 species and 15 nothospecies) of pines as cu ...
with a very restricted range: mostly in California, including several offshore
Channel Islands The Channel Islands are an archipelago in the English Channel, off the French coast of Normandy. They are divided into two Crown Dependencies: the Jersey, Bailiwick of Jersey, which is the largest of the islands; and the Bailiwick of Guernsey, ...
, and a few locations in Baja California, Mexico. Stands of Bishop Pine are also found in Point Reyes National Seashore, in Marin County north of San Francisco, always on the west side of the San Andreas Fault that runs through the park. It is always on or near the coast. In San Luis Obispo County it is found alone or in stands scattered on the coastal mountains and hills from
Morro Bay Morro Bay (''Morro'', Spanish for "Hill") is a seaside city in San Luis Obispo County, California, United States. Located on the Central Coast of California, the city's population was 10,757 as of the 2020 census, up from 10,234 at the 2010 ...
to Shell Beach. A few stands of the tree are seen on the hills above the Sycamore Canyon Resort in Avila Beach. Within the City of
San Luis Obispo ; ; ; Chumashan languages, Chumash: ''tiłhini'') is a city and county seat of San Luis Obispo County, California, United States. Located on the Central Coast (California), Central Coast of California, San Luis Obispo is roughly halfway betwee ...
, the Terrace Hill Open Space has several scattered specimens. Bishop pine seems to prefer already disturbed, unvegetated areas where it probably faces less competition from oaks and shrubs. The common name "bishop pine" resulted from the tree having been first identified near the Mission of San Luis Obispo in
San Luis Obispo ; ; ; Chumashan languages, Chumash: ''tiłhini'') is a city and county seat of San Luis Obispo County, California, United States. Located on the Central Coast (California), Central Coast of California, San Luis Obispo is roughly halfway betwee ...
, California. This tree has a large number of common names and other prior scientific names, due primarily to numerous variant forms. Other English names that have occasionally been used are prickle cone pine, Obispo pine, Santa Cruz pine and dwarf marine pine.


Description

''Pinus muricata'' is a
coniferous Conifers () are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a sin ...
evergreen In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
growing to a height of , rarely up to , with a trunk diameter of up to . The
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
is often smaller, stunted and twisted in coastal exposures. It is drought-tolerant and grows on dry, rocky soil. The needles are in pairs, green to blue-green, and long.
Cone In geometry, a cone is a three-dimensional figure that tapers smoothly from a flat base (typically a circle) to a point not contained in the base, called the '' apex'' or '' vertex''. A cone is formed by a set of line segments, half-lines ...
s occur in one to five clusters. The cones are strongly reflexed down the branch, long; the scales are stiff, thin on the side of the cone facing the stem, but greatly thickened on the side facing away and with a stout spine; both features adaptive to minimise
squirrel Squirrels are members of the family Sciuridae (), a family that includes small or medium-sized rodents. The squirrel family includes tree squirrels, ground squirrels (including chipmunks and prairie dogs, among others), and flying squirrel ...
predation and fire damage to the cones. The cones remain unopened for many years until fire or strong heat causes them to open and release the seeds. File:Pinus muricata-1.jpg, Needles File:Pinus muricata, Morro Bay State Park, San Luis Obispo, California 4.jpg, Cones File:Growing tip of Pinus muricata.JPG, Growing tips


Forms

There are two ''Pinus muricata'' forms: * a southern form with bright green needles * a northern form with dark blue-green needles. The
resin A resin is a solid or highly viscous liquid that can be converted into a polymer. Resins may be biological or synthetic in origin, but are typically harvested from plants. Resins are mixtures of organic compounds, predominantly terpenes. Commo ...
composition also differs. The dividing line between the two is very sharp, five miles (8 km) south of the boundary between Mendocino County and Sonoma County, California. Experimental attempts to hybridize the two forms have consistently failed, indicating that their taxonomic relationship may be more distant than the very small differences in appearance would suggest.


Ecology

''Pinus muricata'' is found with several oak and cypress associates within the
California Coast Ranges The Coast Ranges of California span from Del Norte County, California, Del Norte or Humboldt County, California, south to Santa Barbara County, California, Santa Barbara County. The other three coastal California mountain ranges are the Trans ...
. There are also a number of common
understory In forestry and ecology, understory (American English), or understorey (English in the Commonwealth of Nations, Commonwealth English), also known as underbrush or undergrowth, includes plant life growing beneath the Canopy (biology), forest ca ...
flora associates including sword fern, salal and western poison oak. Notable occurrences of ''P. muricata'' is in association with Mendocino cypress as a
pygmy forest A "natural National Park in the Philippines, a UNESCO World Heritage Site file:Mount Kemiri (8187817161).jpg, An elfin forest in Sumatra's Gunung Leuser National Park Dwarf forest, elfin forest, or pygmy forest is an uncommon ecosystem feat ...
on coastal terraces in Mendocino County and Sonoma County, including one location within
Salt Point State Park Salt Point State Park is a state park in Sonoma County, California, Sonoma County, California, United States. The park covers on the coast of Northern California, with of hiking trails and over of a rough rocky coastline including Salt Point w ...
. It is classified an
endangered species An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching, inv ...
in Mexico.


Uses

''Pinus muricata'' has been used in
plantation Plantations are farms specializing in cash crops, usually mainly planting a single crop, with perhaps ancillary areas for vegetables for eating and so on. Plantations, centered on a plantation house, grow crops including cotton, cannabis, tob ...
s with resultant growth rates higher than in the wild, but with adverse impacts to
biodiversity Biodiversity is the variability of life, life on Earth. It can be measured on various levels. There is for example genetic variability, species diversity, ecosystem diversity and Phylogenetics, phylogenetic diversity. Diversity is not distribut ...
. This plant has ornamental value, and is cultivated in parks and gardens. It has gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. It includes the full range of cultivated p ...
.RHS Plant Selector Pinus muricata AGM / RHS Gardening
/ref>


References


External links

* * * * * {{Taxonbar, from=Q866150 muricata Trees of Northern America Flora of California Natural history of the California chaparral and woodlands Trees of mild maritime climate