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Bird wings are a paired forelimb in
birds Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
. The wings give the birds the ability to fly, creating
lift Lift or LIFT may refer to: Physical devices * Elevator, or lift, a device used for raising and lowering people or goods ** Paternoster lift, a type of lift using a continuous chain of cars which do not stop ** Patient lift, or Hoyer lift, mobile ...
. Terrestrial
flightless birds Flightless birds are birds that through evolution lost the ability to fly. There are over 60 extant species, including the well known ratites ( ostriches, emu, cassowaries, rheas, and kiwi) and penguins. The smallest flightless bird is ...
have reduced wings or none at all (for example, moa). In aquatic flightless birds (
penguins Penguins (order Sphenisciformes , family Spheniscidae ) are a group of aquatic flightless birds. They live almost exclusively in the Southern Hemisphere: only one species, the Galápagos penguin, is found north of the Equator. Highly adap ...
), wings can serve as flippers.


Anatomy

Like most other
tetrapods Tetrapods (; ) are four-limb (anatomy), limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant taxon, extant and extinct amphibians, sauropsids (reptiles, including dinosaurs and therefore birds) and synapsids (p ...
, the forelimb of birds consists of the
shoulder The human shoulder is made up of three bones: the clavicle (collarbone), the scapula (shoulder blade), and the humerus (upper arm bone) as well as associated muscles, ligaments and tendons. The articulations between the bones of the shoulder m ...
(with the
humerus The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a roun ...
), the
forearm The forearm is the region of the upper limb between the elbow and the wrist. The term forearm is used in anatomy to distinguish it from the arm, a word which is most often used to describe the entire appendage of the upper limb, but which in ...
(with the
ulna The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of t ...
and the
radius In classical geometry, a radius ( : radii) of a circle or sphere is any of the line segments from its center to its perimeter, and in more modern usage, it is also their length. The name comes from the latin ''radius'', meaning ray but also the ...
), and the hand. The hand of birds is substantially transformed: some of its bones have been reduced, and some others have merged with each other. Three bones of the
metacarpus In human anatomy, the metacarpal bones or metacarpus form the intermediate part of the skeletal hand located between the phalanges of the fingers and the carpal bones of the wrist, which forms the connection to the forearm. The metacarpal bones ...
and part of the
carpal bones The carpal bones are the eight small bones that make up the wrist (or carpus) that connects the hand to the forearm. The term "carpus" is derived from the Latin carpus and the Greek καρπός (karpós), meaning "wrist". In human anatomy, ...
merge into a carpometacarpus. The bones of three fingers are attached to it. The frontmost one bears an alula—a group of feathers that act like the slats of an airplane. Usually, this finger has one
phalanx bone The phalanges (singular: ''phalanx'' ) are digital bones in the hands and feet of most vertebrates. In primates, the thumbs and big toes have two phalanges while the other digits have three phalanges. The phalanges are classed as long bones ...
, the next has two, and the back has one (but some birds have one more phalanx on the first two fingers—the claw).


Finger identity problem

The bones of three fingers are preserved in the bird wing. The question of which fingers they are has been discussed for about 150 years, and an extensive literature is devoted to it. The anatomical, paleontological, and molecular data suggests that these are fingers 1–3, but embryological data suggests that these are actually fingers 2–4. Several hypotheses have been proposed to explain this discrepancy. Most likely, in birds, finger buds 2–4 began to follow the genetic program for the development of fingers 1–3.


Wing shape

The shape of the wing is important in determining the flight capabilities of a bird. Different shapes correspond to different trade-offs between advantages such as speed, low energy use, and maneuverability. Two important parameters are the aspect ratio and
wing loading In aerodynamics, wing loading is the total mass of an aircraft or flying animal divided by the area of its wing. The stalling speed of an aircraft in straight, level flight is partly determined by its wing loading. An aircraft or animal with a ...
. Aspect ratio is the ratio of
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan ...
to the mean of its
chord Chord may refer to: * Chord (music), an aggregate of musical pitches sounded simultaneously ** Guitar chord a chord played on a guitar, which has a particular tuning * Chord (geometry), a line segment joining two points on a curve * Chord ( ...
(or the square of the wingspan divided by wing area). Wing loading is the ratio of weight to wing area. Most kinds of bird wings can be grouped into four types, with some falling between two of these types. These types of wings are elliptical wings, high-speed wings, high aspect ratio wings and soaring wings with slots.


Elliptical wings

Elliptical wings are rounded and short. This type of wing allows for tight maneuvering in confined spaces such as dense vegetation. Elliptical wings are common in forest raptors (such as ''
Accipiter ''Accipiter'' is a genus of birds of prey in the family Accipitridae. With 51 recognized species it is the most diverse genus in its family. Most species are called goshawks or sparrowhawks, although almost all New World species (excepting the no ...
'' hawks), and many
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped'), which includes more than half of all bird species. Sometimes known as perching birds, passerines are distinguished from other orders of birds by th ...
s, particularly non-migratory ones (migratory species have longer wings). They are also common in species that use a rapid takeoff to evade predators, such as
pheasant Pheasants ( ) are birds of several genera within the family Phasianidae in the order Galliformes. Although they can be found all over the world in introduced (and captive) populations, the pheasant genera native range is restricted to Eurasia ...
s and
partridge A partridge is a medium-sized galliform bird in any of several genera, with a wide native distribution throughout parts of Europe, Asia and Africa. Several species have been introduced to the Americas. They are sometimes grouped in the Perdic ...
s.


High speed wings

High-speed wings are short, pointed wings that when combined with a heavy wing loading and rapid wingbeats provide an energetically expensive, but high-speed flight. This type of wing is present in fast-flying birds such as
duck Duck is the common name for numerous species of waterfowl in the family Anatidae. Ducks are generally smaller and shorter-necked than swans and geese, which are members of the same family. Divided among several subfamilies, they are a fo ...
s. Birds that use their wings to "fly" underwater such as the auks also have small and elongated wings. The peregrine falcon has the highest recorded dive speed of 242 mph (389 km/h). Peregrine falcons have relatively large wings but they partially close their wings during dives. The fastest straight, powered flight is the
spine-tailed swift The white-throated needletail (''Hirundapus caudacutus''), also known as needle-tailed swift or spine-tailed swift, is a large swift in the genus Hirundapus. It is reputed to reach speeds of up to 170 km/h (105 mph) in horizontal flight, but this ...
at 105 mph (170 km/h).


High aspect ratio wings

High aspect ratio (elongated) wings confer high flight efficiency for flights of long duration. When combined with a low wing loading, they are used for slow flight. This may take the form of almost hovering (as used by
kestrel The term kestrel (from french: crécerelle, derivative from , i.e. ratchet) is the common name given to several species of predatory birds from the falcon genus ''Falco''. Kestrels are most easily distinguished by their typical hunting behaviour ...
s,
tern Terns are seabirds in the family Laridae that have a worldwide distribution and are normally found near the sea, rivers, or wetlands. Terns are treated as a subgroup of the family Laridae which includes gulls and skimmers and consists o ...
s and
nightjar Nightjars are medium-sized nocturnal or crepuscular birds in the family Caprimulgidae and order Caprimulgiformes, characterised by long wings, short legs, and very short bills. They are sometimes called goatsuckers, due to the ancient folk tal ...
s) or in soaring and
gliding Gliding is a recreational activity and competitive air sport in which pilots fly unpowered aircraft known as gliders or sailplanes using naturally occurring currents of rising air in the atmosphere to remain airborne. The word ''soaring'' is ...
flight, particularly the dynamic soaring used by
seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same envir ...
s, which takes advantage of wind speed variation at different altitudes (
wind shear Wind shear (or windshear), sometimes referred to as wind gradient, is a difference in wind speed and/or direction over a relatively short distance in the atmosphere. Atmospheric wind shear is normally described as either vertical or horizon ...
) above ocean waves to provide lift. Low-speed flight is also important for birds that plunge-dive for fish.


Soaring wings with deep slots

These wings are favored by larger species of inland birds, such as
eagle Eagle is the common name for many large birds of prey of the family Accipitridae. Eagles belong to several groups of genera, some of which are closely related. Most of the 68 species of eagle are from Eurasia and Africa. Outside this area, j ...
s,
vulture A vulture is a bird of prey that scavenges on carrion. There are 23 extant species of vulture (including Condors). Old World vultures include 16 living species native to Europe, Africa, and Asia; New World vultures are restricted to North a ...
s,
pelican Pelicans (genus ''Pelecanus'') are a genus of large water birds that make up the family Pelecanidae. They are characterized by a long beak and a large throat pouch used for catching prey and draining water from the scooped-up contents before ...
s, and
stork Storks are large, long-legged, long-necked wading birds with long, stout bills. They belong to the family called Ciconiidae, and make up the order Ciconiiformes . Ciconiiformes previously included a number of other families, such as herons a ...
s. The slots at the end of the wings, between the primaries, reduce the
induced drag In aerodynamics, lift-induced drag, induced drag, vortex drag, or sometimes drag due to lift, is an aerodynamic drag force that occurs whenever a moving object redirects the airflow coming at it. This drag force occurs in airplanes due to wings or ...
and
wingtip vortices Wingtip vortices are circular patterns of rotating air left behind a wing as it generates lift.Clancy, L.J., ''Aerodynamics'', section 5.14 One wingtip vortex trails from the tip of each wing. Wingtip vortices are sometimes named ''trailing' ...
by "capturing" the energy in air flowing from the lower to upper wing surface at the tips, whilst the shorter size of the wings aids in takeoff (high aspect ratio wings require a long
taxi A taxi, also known as a taxicab or simply a cab, is a type of vehicle for hire with a driver, used by a single passenger or small group of passengers, often for a non-shared ride. A taxicab conveys passengers between locations of their choic ...
to get airborne).


References

{{fins, limbs and wings Bird anatomy Limbs (anatomy)