A bipinnaria is the first stage in the

larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...

l development of most

starfish Starfish or sea stars are Star polygon, star-shaped echinoderms belonging to the class (biology), class Asteroidea (). Common usage frequently finds these names being also applied to brittle star, ophiuroids, which are correctly referred to ...

, and is usually followed by a

brachiolaria A brachiolaria is the second stage of larval development in many starfishes. It follows the bipinnaria. Brachiolaria have symmetry (biology)#Bilateral_symmetry, bilateral symmetry, unlike the adult starfish, which have a symmetry (biology)#Pentame ...

stage. Movement and feeding is accomplished by the bands of

cilia The cilium (: cilia; ; in Medieval Latin and in anatomy, ''cilium'') is a short hair-like membrane protrusion from many types of eukaryotic cell. (Cilia are absent in bacteria and archaea.) The cilium has the shape of a slender threadlike proj ...

. Starfish that brood their young generally lack a bipinnaria stage, with the eggs developing directly into miniature adults. The bipinnaria is free-living, swimming as part of the

zooplankton Zooplankton are the heterotrophic component of the planktonic community (the " zoo-" prefix comes from ), having to consume other organisms to thrive. Plankton are aquatic organisms that are unable to swim effectively against currents. Consequent ...

. When it initially forms, the entire body is covered by cilia, but as it grows, these become confined to a narrow band forming a number of loops over the body surface. A pair of short, stubby arms soon develop on the body, with the ciliated bands extending into them. In addition to propelling the larva through the water, the cilia also catch suspended food particles, and deliver them to the mouth (more correctly called a ''stomodeum''). Eventually, three additional arms develop at the front end of the larva; at this point it becomes a

. In some species, including the common starfish '' Asterias'', the bipinnaria develops directly into an adult. It is very similar in appearance to the

tornaria A tornaria is the planktonic larva of some species of Hemichordata such as the acorn worms. It is very similar in appearance to the bipinnaria larvae of starfishes, with convoluted bands of cilia running around the body. It is an oval shaped, tra ...

larvae of some

Hemichordata Hemichordata ( ) is a phylum which consists of triploblastic, eucoelomate, and bilaterally symmetrical marine deuterostome animals, generally considered the sister group of the echinoderms. They appear in the Lower or Middle Cambrian and incl ...

, reflecting the descent of the

Ambulacraria Ambulacraria , or Coelomopora , is a clade of invertebrate phyla that includes echinoderms and hemichordates; a member of this group is called an ambulacrarian. Phylogenetic analysis suggests the echinoderms and hemichordates separated around ...

from a common ancestor.

References

Echinoderm biology Larvae {{developmental-biology-stub