A biome () is a distinct geographical region with specific

climate Climate is the long-term weather pattern in a region, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteoro ...

vegetation Vegetation is an assemblage of plants and the ground cover they provide. It is a general term, without specific reference to particular Taxon, taxa, life forms, structure, Spatial ecology, spatial extent, or any other specific Botany, botanic ...

, and animal life. It consists of a biological

community A community is a social unit (a group of people) with a shared socially-significant characteristic, such as place, set of norms, culture, religion, values, customs, or identity. Communities may share a sense of place situated in a given g ...

that has formed in response to its physical environment and regional

. In 1935, Tansley added the climatic and soil aspects to the idea, calling it ''

ecosystem An ecosystem (or ecological system) is a system formed by Organism, organisms in interaction with their Biophysical environment, environment. The Biotic material, biotic and abiotic components are linked together through nutrient cycles and en ...

''. The International Biological Program (1964–74) projects popularized the concept of biome. However, in some contexts, the term ''biome'' is used in a different manner. In German literature, particularly in the

Walter Walter may refer to: People and fictional characters * Walter (name), including a list of people and fictional and mythical characters with the given name or surname * Little Walter, American blues harmonica player Marion Walter Jacobs (1930–19 ...

terminology, the term is used similarly as ''

biotope A biotope is an area of uniform environmental conditions providing a living place for a specific assemblage of flora (plants), plants and fauna (animals), animals. ''Biotope'' is almost synonymous with the term habitat (ecology), "habitat", which ...

'' (a concrete geographical unit), while the biome definition used in this article is used as an international, non-regional, terminology—irrespectively of the continent in which an area is present, it takes the same biome name—and corresponds to his "zonobiome", "orobiome" and "pedobiome" (biomes determined by climate zone, altitude or soil). In the Brazilian literature, the term ''biome'' is sometimes used as a synonym of '' biogeographic province'', an area based on

species composition Relative species abundance is a component of biodiversity and is a measure of how common or rare a species is relative to other species in a defined location or community.Hubbell, S. P. 2001. ''The unified neutral theory of biodiversity and biogeog ...

(the term '' floristic province'' being used when plant species are considered), or also as synonym of the "morphoclimatic and phytogeographical domain" of Ab'Sáber, a geographic space with subcontinental dimensions, with the predominance of similar geomorphologic and climatic characteristics, and of a certain vegetation form. Both include many biomes in fact.

Classifications

To divide the world into a few ecological zones is difficult, notably because of the small-scale variations that exist everywhere on earth and because of the gradual changeover from one biome to the other. Their boundaries must therefore be drawn arbitrarily and their characterization made according to the average conditions that predominate in them. A 1978 study on North American

grasslands A grassland is an area where the vegetation is dominated by grasses (Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes such as clover, and other herbs. Grasslands occur ...

found a positive logistic correlation between

evapotranspiration Evapotranspiration (ET) refers to the combined processes which move water from the Earth's surface (open water and ice surfaces, bare soil and vegetation) into the Atmosphere of Earth, atmosphere. It covers both water evaporation (movement of w ...

in mm/yr and above-ground net primary production in g/m²/yr. The general results from the study were that precipitation and water use led to above-ground primary production, while solar irradiation and temperature lead to below-ground primary production (roots), and temperature and water lead to cool and warm season growth habit. These findings help explain the categories used in Holdridge's bioclassification scheme (see below), which were then later simplified by Whittaker. The number of classification schemes and the variety of determinants used in those schemes, however, should be taken as strong indicators that biomes do not fit perfectly into the classification schemes created.

Holdridge (1947, 1964) life zones

In 1947, the American botanist and climatologist Leslie Holdridge classified climates based on the biological effects of temperature and rainfall on

under the assumption that these two abiotic factors are the largest determinants of the types of vegetation found in a habitat. Holdridge uses the four axes to define 30 so-called "humidity provinces", which are clearly visible in his diagram. While this scheme largely ignores soil and sun exposure, Holdridge acknowledged that these were important.

Allee (1949) biome-types

The principal biome-types by Allee (1949): *

Tundra In physical geography, a tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. There are three regions and associated types of tundra: #Arctic, Arctic, Alpine tundra, Alpine, and #Antarctic ...

Taiga Taiga or tayga ( ; , ), also known as boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruces, and larches. The taiga, or boreal forest, is the world's largest land biome. In North A ...

Deciduous forest In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, after flo ...

Grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...

s *

Desert A desert is a landscape where little precipitation occurs and, consequently, living conditions create unique biomes and ecosystems. The lack of vegetation exposes the unprotected surface of the ground to denudation. About one-third of the la ...

* High plateaus *

Tropical forest Tropical forests are forested ecoregions with tropical climates – that is, land areas approximately bounded by the Tropic of Cancer, tropics of Cancer and Tropic of Capricorn, Capricorn, but possibly affected by other factors such as prevailing ...

* Minor terrestrial biomes

Kendeigh (1961) biomes

The principal biomes of the world by Kendeigh (1961): * ''Terrestrial'' ** Temperate deciduous forest ** Coniferous forest **

Woodland A woodland () is, in the broad sense, land covered with woody plants (trees and shrubs), or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunli ...

Chaparral Chaparral ( ) is a shrubland plant plant community, community found primarily in California, southern Oregon, and northern Baja California. It is shaped by a Mediterranean climate (mild wet winters and hot dry summers) and infrequent, high-intens ...

** Tropical savanna **

* Marine **

Ocean The ocean is the body of salt water that covers approximately 70.8% of Earth. The ocean is conventionally divided into large bodies of water, which are also referred to as ''oceans'' (the Pacific, Atlantic, Indian Ocean, Indian, Southern Ocean ...

plankton Plankton are the diverse collection of organisms that drift in Hydrosphere, water (or atmosphere, air) but are unable to actively propel themselves against ocean current, currents (or wind). The individual organisms constituting plankton are ca ...

and nekton ** Balanoid-gastropod- thallophyte ** Pelecypod-

annelid The annelids (), also known as the segmented worms, are animals that comprise the phylum Annelida (; ). The phylum contains over 22,000 extant species, including ragworms, earthworms, and leeches. The species exist in and have adapted to vario ...

Coral reef A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of colonies of coral polyps held together by calcium carbonate. Most coral reefs are built from stony corals, whose polyps cluster in group ...

Whittaker (1962, 1970, 1975) biome-types

Whittaker classified biomes using two abiotic factors: precipitation and temperature. His scheme can be seen as a simplification of Holdridge's; more readily accessible, but missing Holdridge's greater specificity. Whittaker based his approach on theoretical assertions and empirical sampling. He had previously compiled a review of biome classifications.

Key definitions for understanding Whittaker's scheme

Physiognomy Physiognomy () or face reading is the practice of assessing a person's character or personality from their outer appearance—especially the face. The term can also refer to the general appearance of a person, object, or terrain without referenc ...

: sometimes referring to the plants' appearance; or the biome's apparent characteristics, outward features, or appearance of ecological communities or species – including plants. * Biome: a grouping of terrestrial ecosystems on a given continent that is similar in vegetation structure, physiognomy, features of the environment and characteristics of their animal communities. * Formation: a major kind of community of plants on a given continent. * Biome-type: grouping of convergent biomes or formations of different continents, defined by physiognomy. * Formation-type: a grouping of convergent formations. Whittaker's distinction between biome and formation can be simplified: formation is used when applied to

plant communities A plant community is a collection or Association (ecology), association of plant species within a designated geographical unit, which forms a relatively uniform patch, distinguishable from neighboring patches of different vegetation types. The comp ...

only, while biome is used when concerned with both plants and animals. Whittaker's convention of biome-type or formation-type is a broader method to categorize similar communities.

Whittaker's parameters for classifying biome-types

Whittaker used what he called "gradient analysis" of ecocline patterns to relate communities to climate on a worldwide scale. Whittaker considered four main ecoclines in the terrestrial realm. # Intertidal levels: The wetness gradient of areas that are exposed to alternating water and dryness with intensities that vary by location from high to low tide # Climatic moisture gradient # Temperature gradient by altitude # Temperature gradient by latitude Along these gradients, Whittaker noted several trends that allowed him to qualitatively establish biome-types: * The gradient runs from favorable to the extreme, with corresponding changes in productivity. * Changes in physiognomic complexity vary with how favorable of an environment exists (decreasing community structure and reduction of stratal differentiation as the environment becomes less favorable). * Trends in the diversity of structure follow trends in species diversity; alpha and beta species diversities decrease from favorable to extreme environments. * Each growth-form (i.e. grasses, shrubs, etc.) has its characteristic place of maximum importance along the ecoclines. * The same growth forms may be dominant in similar environments in widely different parts of the world. Whittaker summed the effects of gradients (3) and (4) to get an overall temperature gradient and combined this with a gradient (2), the moisture gradient, to express the above conclusions in what is known as the Whittaker classification scheme. The scheme graphs average annual precipitation (x-axis) versus average annual temperature (y-axis) to classify biome-types.

Biome-types

Tropical rainforest Tropical rainforests are dense and warm rainforests with high rainfall typically found between 10° north and south of the Equator. They are a subset of the tropical forest biome that occurs roughly within the 28° latitudes (in the torrid zo ...

# Tropical seasonal rainforest #*

deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...

#* semideciduous # Temperate giant rainforest # Montane rainforest # Temperate deciduous forest # Temperate evergreen forest #* needleleaf #*

sclerophyll Sclerophyll is a type of vegetation that is adapted to long periods of dryness and heat. The plants feature hard leaves, short Internode (botany), internodes (the distance between leaves along the stem) and leaf orientation which is parallel or ...

# Subarctic-subalpine needle-leaved forests (

taiga Taiga or tayga ( ; , ), also known as boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, spruces, and larches. The taiga, or boreal forest, is the world's largest land biome. In North A ...

) # Elfin woodland # Thorn forest # Thorn scrub # Temperate woodland # Temperate shrublands #*

#* heath #*

#* subalpine-needleleaf #* subalpine-broadleaf #

Savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...

# Temperate grassland # Alpine grasslands #

# Tropical desert # Warm-temperate desert # Cool temperate desert scrub # Arctic-alpine desert #

Bog A bog or bogland is a wetland that accumulates peat as a deposit of dead plant materials often mosses, typically sphagnum moss. It is one of the four main types of wetlands. Other names for bogs include mire, mosses, quagmire, and musk ...

# Tropical fresh-water swamp forest # Temperate fresh-water swamp forest # Mangrove swamp # Salt marsh # Wetland

Goodall (1974–) ecosystem types

The multi-authored series ''Ecosystems of the World'', edited by David W. Goodall, provides a comprehensive coverage of the major "ecosystem types or biomes" on Earth:

Walter (1976, 2002) zonobiomes

The eponymously named Heinrich Walter classification scheme considers the seasonality of temperature and precipitation. The system, also assessing precipitation and temperature, finds nine major biome types, with the important climate traits and vegetation types. The boundaries of each biome correlate to the conditions of moisture and cold stress that are strong determinants of plant form, and therefore the vegetation that defines the region. Extreme conditions, such as flooding in a swamp, can create different kinds of communities within the same biome.

Schultz (1988) eco-zones

Schultz (1988, 2005) defined nine ''ecozones'' (his concept of ecozone is more similar to the concept of biome than to the concept of ecozone of BBC):Schultz, J. ''Die Ökozonen der Erde'', 1st ed., Ulmer, Stuttgart, Germany, 1988, 488 pp.; 2nd ed., 1995, 535 pp.; 3rd ed., 2002; 4th ed., 2008; 5th ed., 2016. Transl.: ''The Ecozones of the World: The Ecological Divisions of the Geosphere''. Berlin: Springer-Verlag, 1995; 2nd ed., 2005

# polar/subpolar zone # boreal zone # humid mid-latitudes # dry mid-latitudes # subtropics with winter rain # subtropics with year-round rain # dry tropics and subtropics # tropics with summer rain # tropics with year-round rain

Bailey (1989) ecoregions

Robert Bailey (geographer), Robert G. Bailey nearly developed a biogeographical classification system of

ecoregion An ecoregion (ecological region) is an ecological and geographic area that exists on multiple different levels, defined by type, quality, and quantity of environmental resources. Ecoregions cover relatively large areas of land or water, and c ...

s for the United States in a map published in 1976. He subsequently expanded the system to include the rest of North America in 1981, and the world in 1989. The Bailey system, based on climate, is divided into four domains (polar, humid temperate, dry, and humid tropical), with further divisions based on other climate characteristics (subarctic, warm temperate, hot temperate, and subtropical; marine and continental; lowland and mountain). * 100 Polar Domain ** 120

Division (Köppen: Ft) ** M120 Tundra Division – Mountain Provinces ** 130 Subarctic Division (Köppen: E) ** M130 Subarctic Division – Mountain Provinces * 200 Humid Temperate Domain ** 210 Warm Continental Division (Köppen: portion of Dcb) ** M210 Warm Continental Division – Mountain Provinces ** 220 Hot Continental Division (Köppen: portion of Dca) ** M220 Hot Continental Division – Mountain Provinces ** 230 Subtropical Division (Köppen: portion of Cf) ** M230 Subtropical Division – Mountain Provinces ** 240 Marine Division (Köppen: Do) ** M240 Marine Division – Mountain Provinces ** 250 Prairie Division (Köppen: arid portions of Cf, Dca, Dcb) ** 260 Mediterranean Division (Köppen: Cs) ** M260 Mediterranean Division – Mountain Provinces * 300 Dry Domain ** 310 Tropical/Subtropical Steppe Division ** M310 Tropical/Subtropical Steppe Division – Mountain Provinces ** 320 Tropical/Subtropical Desert Division ** 330 Temperate Steppe Division ** 340 Temperate Desert Division * 400 Humid Tropical Domain ** 410 Savanna Division ** 420 Rainforest Division

Olson & Dinerstein (1998) biomes for WWF / Global 200

A team of biologists convened by the

World Wildlife Fund The World Wide Fund for Nature (WWF) is a Swiss-based international non-governmental organization founded in 1961 that works in the field of wilderness preservation and the reduction of human impact on the environment. It was formerly named the ...

(WWF) developed a scheme that divided the world's land area into

biogeographic realm A biogeographic realm is the broadest biogeography, biogeographic division of Earth's land surface, based on distributional patterns of terrestrial animal, terrestrial organisms. They are subdivided into bioregions, which are further subdivid ...

s (called "ecozones" in a BBC scheme), and these into

s (Olson & Dinerstein, 1998, etc.). Each ecoregion is characterized by a main biome (also called major habitat type).Olson, D. M. & E. Dinerstein (1998). The Global 200: A representation approach to conserving the Earth's most biologically valuable ecoregions. ''Conservation Biol.'' 12:502–515

.Olson, D. M., Dinerstein, E., Wikramanayake, E. D., Burgess, N. D., Powell, G. V. N., Underwood, E. C., D'Amico, J. A., Itoua, I., Strand, H. E., Morrison, J. C., Loucks, C. J., Allnutt, T. F., Ricketts, T. H., Kura, Y., Lamoreux, J. F., Wettengel, W. W., Hedao, P., Kassem, K. R. (2001). Terrestrial ecoregions of the world: a new map of life on Earth. ''Bioscience'' 51(11):933–938

. This classification is used to define the

Global 200 The Global 200 is the list of ecoregions identified by the World Wide Fund for Nature ( WWF), the global conservation organization, as priorities for conservation. According to WWF, an ecoregion is defined as a "relatively large unit of land or w ...

list of

s identified by the WWF as priorities for conservation. For the

terrestrial ecoregions An ecoregion (ecological region) is an ecology, ecological and Geography, geographic area that exists on multiple different levels, defined by type, quality, and quantity of environmental resources. Ecoregions cover relatively large areas of la ...

, there is a specific EcoID, format XXnnNN (XX is the

, nn is the biome number, NN is the individual number).

Biogeographic realm A biogeographic realm is the broadest biogeography, biogeographic division of Earth's land surface, based on distributional patterns of terrestrial animal, terrestrial organisms. They are subdivided into bioregions, which are further subdivid ...
s (terrestrial and freshwater)

* NA:

Nearctic The Nearctic realm is one of the eight biogeographic realms constituting the Earth's land surface. The Nearctic realm covers most of North America, including Greenland, Central Florida, and the highlands of Mexico. The parts of North America ...

* PA:

Palearctic The Palearctic or Palaearctic is a biogeographic realm of the Earth, the largest of eight. Confined almost entirely to the Eastern Hemisphere, it stretches across Europe and Asia, north of the foothills of the Himalayas, and North Africa. Th ...

* AT: Afrotropic * IM:

Indomalaya The Indomalayan realm is one of the eight biogeographic realms. It extends across most of South and Southeast Asia and into the southern parts of East Asia. Also called the Oriental realm by biogeographers, Indomalaya spreads all over the Ind ...

* AA:

Australasia Australasia is a subregion of Oceania, comprising Australia, New Zealand (overlapping with Polynesia), and sometimes including New Guinea and surrounding islands (overlapping with Melanesia). The term is used in a number of different context ...

* NT: Neotropic * OC:

Oceania Oceania ( , ) is a region, geographical region including Australasia, Melanesia, Micronesia, and Polynesia. Outside of the English-speaking world, Oceania is generally considered a continent, while Mainland Australia is regarded as its co ...

* AN:

Antarctic The Antarctic (, ; commonly ) is the polar regions of Earth, polar region of Earth that surrounds the South Pole, lying within the Antarctic Circle. It is antipodes, diametrically opposite of the Arctic region around the North Pole. The Antar ...

The applicability of the realms scheme above – based on Udvardy (1975)—to most freshwater taxa is unresolved.Abell, R., M. Thieme, C. Revenga, M. Bryer, M. Kottelat, N. Bogutskaya, B. Coad, N. Mandrak, S. Contreras-Balderas, W. Bussing, M. L. J. Stiassny, P. Skelton, G. R. Allen, P. Unmack, A. Naseka, R. Ng, N. Sindorf, J. Robertson, E. Armijo, J. Higgins, T. J. Heibel, E. Wikramanayake, D. Olson, H. L. Lopez, R. E. d. Reis, J. G. Lundberg, M. H. Sabaj Perez, and P. Petry. (2008). Freshwater ecoregions of the world: A new map of biogeographic units for freshwater biodiversity conservation. ''BioScience'' 58:403–414

.

Biogeographic realms ( Marine ecoregion, marine)

Arctic The Arctic (; . ) is the polar regions of Earth, polar region of Earth that surrounds the North Pole, lying within the Arctic Circle. The Arctic region, from the IERS Reference Meridian travelling east, consists of parts of northern Norway ( ...

* Temperate Northern Atlantic * Temperate Northern Pacific * Tropical Atlantic * Western Indo-Pacific * Central Indo-Pacific * Eastern Indo-Pacific * Tropical Eastern Pacific * Temperate South America * Temperate Southern Africa * Temperate Australasia *

Southern Ocean The Southern Ocean, also known as the Antarctic Ocean, comprises the southernmost waters of the world ocean, generally taken to be south of 60th parallel south, 60° S latitude and encircling Antarctica. With a size of , it is the seco ...

Spalding, M. D. et al. (2007). Marine ecoregions of the world: a bioregionalization of coastal and shelf areas. ''BioScience'' 57: 573–583

.

Biomes (terrestrial)

Tropical and subtropical moist broadleaf forests Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature (WWF). Description TSMF is generally found in la ...

(tropical and subtropical, humid) #

Tropical and subtropical dry broadleaf forests The tropical and subtropical dry broadleaf forest is a habitat (ecology), habitat type defined by the World Wide Fund for Nature and is located at tropical and subtropical latitudes. Though these forests occur in climates that are warm year-roun ...

(tropical and subtropical, semihumid) #

Tropical and subtropical coniferous forests Tropical and subtropical coniferous forests are a tropical forest habitat (ecology), habitat type defined by the World Wide Fund for Nature. These forests are found predominantly in North America, North and Central America and experience low level ...

(tropical and subtropical, semihumid) # Temperate broadleaf and mixed forests (temperate, humid) #

Temperate coniferous forests Temperate coniferous forest is a terrestrial ecoregion, terrestrial biome defined by the World Wide Fund for Nature. Temperate coniferous forests are found predominantly in areas with warm summers and cool winters, and vary in their kinds of plant ...

(temperate, humid to semihumid) # Boreal forests/taiga (subarctic, humid) #

Tropical and subtropical grasslands, savannas, and shrublands Tropical and subtropical grasslands, savannas, and shrublands is a terrestrial biome defined by the World Wide Fund for Nature. The biome is dominated by grass and/or shrubs located in semi-arid to semi- humid climate regions of subtropical and ...

(tropical and subtropical, semiarid) #

Temperate grasslands, savannas, and shrublands Temperate grasslands, savannas, and shrublands are terrestrial biomes defined by the World Wide Fund for Nature. The predominant vegetation in these biomes consists of grass and/or shrubs. The climate is temperate and ranges from Semi-arid clima ...

(temperate, semiarid) # Flooded grasslands and savannas (temperate to tropical, fresh or brackish water inundated) #

Montane grasslands and shrublands Montane grasslands and shrublands are a biome defined by the World Wildlife Fund. The biome includes high elevation grasslands and shrublands around the world. The term "montane" in the name of the biome refers to "high elevation", rather than t ...

(alpine or montane climate) #

(Arctic) # Mediterranean forests, woodlands, and scrub or sclerophyll forests (temperate warm, semihumid to semiarid with winter rainfall) #

Deserts and xeric shrublands Deserts and xeric shrublands are a biome defined by the World Wide Fund for Nature. Deserts and xeric (Ancient Greek 'dry') shrublands form the largest terrestrial biome, covering 19% of Earth's land surface area. Ecoregions in this habitat ...

(temperate to tropical, arid) #

Mangrove A mangrove is a shrub or tree that grows mainly in coastal saline water, saline or brackish water. Mangroves grow in an equatorial climate, typically along coastlines and tidal rivers. They have particular adaptations to take in extra oxygen a ...

(subtropical and tropical, salt water inundated)

Biomes (freshwater)

According to the WWF, the following are classified as

freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. The term excludes seawater and brackish water, but it does include non-salty mi ...

biomes: * Large

lake A lake is often a naturally occurring, relatively large and fixed body of water on or near the Earth's surface. It is localized in a basin or interconnected basins surrounded by dry land. Lakes lie completely on land and are separate from ...

s * Large

river deltas A river delta is a landform, wikt:archetype#Noun, archetypically triangular, created by the deposition (geology), deposition of the sediments that are carried by the waters of a river, where the river merges with a body of slow-moving water or ...

* Polar

s * Montane

s * Temperate coastal rivers * Temperate floodplain rivers and

wetland A wetland is a distinct semi-aquatic ecosystem whose groundcovers are flooded or saturated in water, either permanently, for years or decades, or only seasonally. Flooding results in oxygen-poor ( anoxic) processes taking place, especially ...

s * Temperate upland rivers * Tropical and subtropical coastal rivers * Tropical and subtropical floodplain rivers and

s * Tropical and subtropical upland rivers * Xeric freshwaters and

endorheic basin An endorheic basin ( ; also endoreic basin and endorreic basin) is a drainage basin that normally retains water and allows no outflow to other external bodies of water (e.g. rivers and oceans); instead, the water drainage flows into permanent ...

s * Oceanic islands

Biomes (marine)

Biomes of the coastal and

continental shelf A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an islan ...

areas (

neritic zone The neritic zone (or sublittoral zone) is the relatively shallow part of the ocean above the drop-off of the continental shelf, approximately in depth. From the point of view of marine biology it forms a relatively stable and well-illuminate ...

): * Polar * Temperate shelves and sea * Temperate

upwelling Upwelling is an physical oceanography, oceanographic phenomenon that involves wind-driven motion of dense, cooler, and usually nutrient-rich water from deep water towards the ocean surface. It replaces the warmer and usually nutrient-depleted sur ...

* Tropical

* Tropical coral

Summary of the scheme

Biosphere The biosphere (), also called the ecosphere (), is the worldwide sum of all ecosystems. It can also be termed the zone of life on the Earth. The biosphere (which is technically a spherical shell) is virtually a closed system with regard to mat ...

Biogeographic realm A biogeographic realm is the broadest biogeography, biogeographic division of Earth's land surface, based on distributional patterns of terrestrial animal, terrestrial organisms. They are subdivided into bioregions, which are further subdivid ...

s (terrestrial) (8) ***

Ecoregions An ecoregion (ecological region) is an ecology, ecological and Geography, geographic area that exists on multiple different levels, defined by type, quality, and quantity of environmental resources. Ecoregions cover relatively large areas of la ...

(867), each characterized by a biome, a major habitat type (14) ****

Ecosystems An ecosystem (or ecological system) is a system formed by Organism, organisms in interaction with their Biophysical environment, environment. The Biotic material, biotic and abiotic components are linked together through nutrient cycles and en ...

(

s) *

s (freshwater) (8) ***

(426), each characterized by a biome, a major habitat type (12) ****

(biotopes) *

** Biogeographic realms (marine) (12) *** ( Marine provinces) (62) ****

(232), each characterized by a biome, a major habitat type (5) *****

(biotopes) Example: *

***

Ecoregion An ecoregion (ecological region) is an ecological and geographic area that exists on multiple different levels, defined by type, quality, and quantity of environmental resources. Ecoregions cover relatively large areas of land or water, and c ...

Dinaric Mountains mixed forests The Dinaric Mountains mixed forests are a terrestrial ecoregion of the temperate broadleaf and mixed forests biome in Southeastern Europe, according to both the WWF and Digital Map of European Ecological Regions by the European Environment Age ...

(PA0418); biome type: temperate broadleaf and mixed forests ****

Ecosystem An ecosystem (or ecological system) is a system formed by Organism, organisms in interaction with their Biophysical environment, environment. The Biotic material, biotic and abiotic components are linked together through nutrient cycles and en ...

: Orjen, vegetation belt between 1,100 and 1,450 m, Oromediterranean zone, nemoral zone (temperate zone) *****

Biotope A biotope is an area of uniform environmental conditions providing a living place for a specific assemblage of flora (plants), plants and fauna (animals), animals. ''Biotope'' is almost synonymous with the term habitat (ecology), "habitat", which ...

: ''Oreoherzogio-Abietetum illyricae'' Fuk. ( Plant list) ****** Plant: Silver fir ('' Abies alba'')

Other biomes

Marine biomes

Pruvot (1896) zones or "systems": *

Littoral zone The littoral zone, also called litoral or nearshore, is the part of a sea, lake, or river that is close to the shore. In coastal ecology, the littoral zone includes the intertidal zone extending from the high water mark (which is rarely flood ...

Pelagic zone The pelagic zone consists of the water column of the open ocean and can be further divided into regions by depth. The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or water column between the sur ...

Abyssal zone The abyssal zone or abyssopelagic zone is a layer of the pelagic zone of the ocean. The word ''abyss'' comes from the Greek word (), meaning "bottomless". At depths of , this zone remains in perpetual darkness. It covers 83% of the total area ...

Longhurst (1998) biomes: * Coastal * Polar * Trade wind * Westerly Other marine habitat types (not covered yet by the Global 200/WWF scheme): * Open sea *

Deep sea The deep sea is broadly defined as the ocean depth where light begins to fade, at an approximate depth of or the point of transition from continental shelves to continental slopes. Conditions within the deep sea are a combination of low tempe ...

Hydrothermal vent Hydrothermal vents are fissures on the seabed from which geothermally heated water discharges. They are commonly found near volcanically active places, areas where tectonic plates are moving apart at mid-ocean ridges, ocean basins, and hot ...

s * Cold seeps *

Benthic zone The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from the Ancient Greek word (), meaning "the depths". ...

(trades and westerlies) * Abyssal * Hadal (ocean trench) * Littoral/

Intertidal zone The intertidal zone or foreshore is the area above water level at low tide and underwater at high tide; in other words, it is the part of the littoral zone within the tidal range. This area can include several types of habitats with various ...

Salt marsh A salt marsh, saltmarsh or salting, also known as a coastal salt marsh or a tidal marsh, is a coastal ecosystem in the upper coastal intertidal zone between land and open saltwater or brackish water that is regularly flooded by the tides. I ...

Estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environm ...

* Coastal lagoons/Atoll lagoons *

Kelp forest Kelp forests are underwater areas with a high density of kelp, which covers a large part of the world's coastlines. Smaller areas of anchored kelp are called kelp beds. They are recognized as one of the most productive and dynamic ecosystems on E ...

* Pack ice

Anthropogenic biomes

Humans have altered global patterns of

biodiversity Biodiversity is the variability of life, life on Earth. It can be measured on various levels. There is for example genetic variability, species diversity, ecosystem diversity and Phylogenetics, phylogenetic diversity. Diversity is not distribut ...

and

processes. As a result, vegetation forms predicted by conventional biome systems can no longer be observed across much of Earth's land surface as they have been replaced by crops and rangelands or cities. Anthropogenic biomes provide an alternative view of the terrestrial biosphere based on global patterns of sustained direct human interaction with ecosystems, including

agriculture Agriculture encompasses crop and livestock production, aquaculture, and forestry for food and non-food products. Agriculture was a key factor in the rise of sedentary human civilization, whereby farming of domesticated species created ...

human settlement In geography, statistics and archaeology, a settlement, locality or populated place is a community of people living in a particular location, place. The complexity of a settlement can range from a minuscule number of Dwelling, dwellings gro ...

urbanization Urbanization (or urbanisation in British English) is the population shift from Rural area, rural to urban areas, the corresponding decrease in the proportion of people living in rural areas, and the ways in which societies adapt to this change. ...

forestry Forestry is the science and craft of creating, managing, planting, using, conserving and repairing forests and woodlands for associated resources for human and Natural environment, environmental benefits. Forestry is practiced in plantations and ...

and other uses of land. Anthropogenic biomes offer a way to recognize the irreversible coupling of human and ecological systems at global scales and manage Earth's biosphere and anthropogenic biomes. Land cover IGBP

Major anthropogenic biomes: * Dense settlements * Croplands *

Rangeland Rangelands are grasslands, shrublands, woodlands, wetlands, and deserts that are grazed by domestic livestock or wild animals. Types of rangelands include tallgrass and shortgrass prairies, desert grasslands and shrublands, woodlands, savanna ...

s * Forested * Indoor

Microbial biomes

Endolithic biomes

The endolithic biome, consisting entirely of microscopic life in rock pores and cracks, kilometers beneath the surface, has only recently been discovered, and does not fit well into most classification schemes.

Effects of climate change

Anthropogenic

climate change Present-day climate change includes both global warming—the ongoing increase in Global surface temperature, global average temperature—and its wider effects on Earth's climate system. Climate variability and change, Climate change in ...

has the potential to greatly alter the distribution of Earth's biomes. Meaning, biomes around the world could change so much that they would be at risk of becoming new biomes entirely. More specifically, between 54% and 22% of global land area will experience climates that correspond to other biomes. 3.6% of land area will experience climates that are completely new or unusual. An example of a biome shift is woody plant encroachment, which can change grass savanna into shrub savanna. Average temperatures have risen more than twice the usual amount in both arctic and mountainous biomes, which leads to the conclusion that arctic and mountainous biomes are currently the most vulnerable to climate change. South American terrestrial biomes have been predicted to go through the same temperature trends as arctic and mountainous biomes. With its annual average temperature continuing to increase, the moisture currently located in forest biomes will dry up.

References