William George Hill (7 August 1940 – 17 December 2021) was an English geneticist and statistician. He was a professor at

University of Edinburgh The University of Edinburgh (, ; abbreviated as ''Edin.'' in Post-nominal letters, post-nominals) is a Public university, public research university based in Edinburgh, Scotland. Founded by the City of Edinburgh Council, town council under th ...

. He is credited as co-discoverer of the Hill–Robertson effect with his doctoral advisor, Alan Robertson.

Education

Hill was educated at

St Albans School, Hertfordshire St Albans School is a public school (traditional English fee-charging day and former boarding school) in the city of St Albans in Hertfordshire. Entry before Sixth Form is for boys only, but the Sixth Form has been co-educational since 1991. ...

and studied agriculture at

Wye College The College of St Gregory and St Martin at Wye, commonly known as Wye College, was an education and research institution in the village of Wye, Kent. In 1447, Cardinal (Catholic Church), Cardinal John Kempe founded his chantry there which also ...

University of London The University of London (UoL; abbreviated as Lond or more rarely Londin in Post-nominal letters, post-nominals) is a collegiate university, federal Public university, public research university located in London, England, United Kingdom. The ...

graduating with a

Bachelor of Science A Bachelor of Science (BS, BSc, B.S., B.Sc., SB, or ScB; from the Latin ') is a bachelor's degree that is awarded for programs that generally last three to five years. The first university to admit a student to the degree of Bachelor of Scienc ...

degree in 1961. He studied genetics at the

University of California, Davis The University of California, Davis (UC Davis, UCD, or Davis) is a Public university, public Land-grant university, land-grant research university in Davis, California, United States. It is the northernmost of the ten campuses of the University ...

, graduating with a

Master of Science A Master of Science (; abbreviated MS, M.S., MSc, M.Sc., SM, S.M., ScM or Sc.M.) is a master's degree. In contrast to the Master of Arts degree, the Master of Science degree is typically granted for studies in sciences, engineering and medici ...

degree in 1963, then moved to Edinburgh to pursue a

PhD A Doctor of Philosophy (PhD, DPhil; or ) is a terminal degree that usually denotes the highest level of academic achievement in a given discipline and is awarded following a course of graduate study and original research. The name of the deg ...

population genetics Population genetics is a subfield of genetics that deals with genetic differences within and among populations, and is a part of evolutionary biology. Studies in this branch of biology examine such phenomena as Adaptation (biology), adaptation, s ...

with Alan Robertson. His presented thesis was ''"Studies on artificial selection"''. He was awarded a

Doctor of Science A Doctor of Science (; most commonly abbreviated DSc or ScD) is a science doctorate awarded in a number of countries throughout the world. Africa Algeria and Morocco In Algeria, Morocco, Libya and Tunisia, all universities accredited by the s ...

degree in 1976 for research on

quantitative genetics Quantitative genetics is the study of quantitative traits, which are phenotypes that vary continuously—such as height or mass—as opposed to phenotypes and gene-products that are Categorical variable, discretely identifiable—such as eye-col ...

Research and career

Hill was distinguished for his theoretical contributions to the study of the population and quantitative genetics of finite populations, in particular with respect to multilocus problems. He was the first to present formulae for the expected association of linked genes in finite populations due to random sampling of gametes and for the estimation of these associations from

genotype The genotype of an organism is its complete set of genetic material. Genotype can also be used to refer to the alleles or variants an individual carries in a particular gene or genetic location. The number of alleles an individual can have in a ...

frequencies. He has made major contributions to the analysis of quantitative variation in random breeding

population Population is a set of humans or other organisms in a given region or area. Governments conduct a census to quantify the resident population size within a given jurisdiction. The term is also applied to non-human animals, microorganisms, and pl ...

s, both in the design and interpretation of selection experiments and in the analysis of similarity between relatives. He applied these concepts in his own selection experiments in the laboratory and in farm animal improvement programmes. Hill served as

editor in chief An editor-in-chief (EIC), also known as lead editor or chief editor, is a publication's editorial leader who has final responsibility for its operations and policies. The editor-in-chief heads all departments of the organization and is held account ...

of the ''

Proceedings of the Royal Society B ''Proceedings of the Royal Society'' is the main research journal of the Royal Society. The journal began in 1831 and was split into two series in 1905: * Series A: for papers in physical sciences and mathematics. * Series B: for papers in life s ...

'' from 2005 to 2009.

Awards and honours

Hill was elected a

Fellow of the Royal Society of Edinburgh Fellowship of the Royal Society of Edinburgh (FRSE) is an award granted to individuals that the Royal Society of Edinburgh, Scotland's national academy of science and Literature, letters, judged to be "eminently distinguished in their subject". ...

in 1979, a Fellow of the Royal Society (FRS) in 1985 and appointed OBE in 2004. One or more of the preceding sentences incorporates text from the royalsociety.org website where: In 2018 he was awarded The Royal Society's

Darwin Medal The Darwin Medal is one of the medals awarded by the Royal Society for "distinction in evolution, biological diversity and developmental, population and organismal biology". In 1885, International Darwin Memorial Fund was transferred to the ...

for his research in quantitative genetics. In 2019 he was awarded The Genetics Society's Mendel Medal at The Centenary of Genetics Conference, for his contribution to quantitative genetics.

References

{{DEFAULTSORT:Hill, Bill 1940 births 2021 deaths English geneticists Academics of the University of Edinburgh Fellows of the Royal Society Fellows of the Royal Society of Edinburgh Officers of the Order of the British Empire English statisticians People educated at St Albans School, Hertfordshire University of California, Davis alumni Place of birth missing Alumni of the University of London British academic journal editors Alumni of Wye College Alumni of the University of Edinburgh 20th-century English scientists