Berylline Hummingbird
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The berylline hummingbird (''Saucerottia beryllina'') is a species of
hummingbird Hummingbirds are birds native to the Americas and comprise the Family (biology), biological family Trochilidae. With approximately 366 species and 113 genus, genera, they occur from Alaska to Tierra del Fuego, but most species are found in Cen ...
in the "emeralds", tribe Trochilini of subfamily Trochilinae. It is found in
El Salvador El Salvador, officially the Republic of El Salvador, is a country in Central America. It is bordered on the northeast by Honduras, on the northwest by Guatemala, and on the south by the Pacific Ocean. El Salvador's capital and largest city is S ...
,
Guatemala Guatemala, officially the Republic of Guatemala, is a country in Central America. It is bordered to the north and west by Mexico, to the northeast by Belize, to the east by Honduras, and to the southeast by El Salvador. It is hydrologically b ...
,
Honduras Honduras, officially the Republic of Honduras, is a country in Central America. It is bordered to the west by Guatemala, to the southwest by El Salvador, to the southeast by Nicaragua, to the south by the Pacific Ocean at the Gulf of Fonseca, ...
,
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
, and the
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
.


Taxonomy and systematics

The berylline hummingbird was formerly placed in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''
Amazilia ''Amazilia'' is a hummingbird genus in the subfamily Trochilinae. It is found in tropical Central America, Central and South America. Taxonomy The genus ''Amazilia'' was introduced in 1843 by the French naturalist René Lesson. Lesson had us ...
''. A
molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
study published in 2014 found that the genus ''Amazilia'' was
polyphyletic A polyphyletic group is an assemblage that includes organisms with mixed evolutionary origin but does not include their most recent common ancestor. The term is often applied to groups that share similar features known as Homoplasy, homoplasies ...
. In the revised classification to create
monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent co ...
genera, the berylline hummingbird was moved by most taxonomic systems to the resurrected genus ''
Saucerottia ''Saucerottia'' is a genus of birds in the family (biology), family Trochilidae, or hummingbirds. Species The species now placed in this genus were formerly placed in ''Amazilia''. A molecular phylogenetic study published in 2014 found that the ...
''.Clements, J. F., T. S. Schulenberg, M. J. Iliff, S. M. Billerman, T. A. Fredericks, J. A. Gerbracht, D. Lepage, B. L. Sullivan, and C. L. Wood. 2021. The eBird/Clements checklist of Birds of the World: v2021. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ Retrieved August 25, 2021 However,
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding i ...
's ''
Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. ...
'' retains it in ''Amazilia''. The berylline hummingbird has these five subspecies: *''S. b. viola'' ( Miller, W, 1905) *''S. b. beryllina'' ( Deppe, 1830) *''S. b. lichtensteini'' ( Moore, RT, 1950) *''S. b. sumichrasti'' ( Salvin, 1891) *''S. b. devillei'' ( Bourcier & Mulsant, 1848)


Description

The berylline hummingbird is long. Males weigh about and females about . The bills of males of all subspecies have a black
maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...
and a pinkish
mandible In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...
with a black outer half. Adult males of the
nominate subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...
''S. b. beryllina'' have a bronze-green to coppery head, back and rump. Their uppertail
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are small feathers behind t ...
and tail are coppery to rufous. The bases of their
primaries Primary elections or primaries are elections held to determine which candidates will run in an upcoming general election. In a partisan primary, a political party selects a candidate. Depending on the state and/or party, there may be an "open pri ...
and secondaries are chestnut and show as a patch on the closed wing. Their underparts are glittering golden green. Adult nominate females are very similar to males, but with a paler and more grayish throat and belly. Their bills are entirely black. Juveniles are similar to females but with a grayish cinnamon belly.Weller, A.A. and G. M. Kirwan (2021). Berylline Hummingbird (''Saucerottia beryllina''), version 1.1. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.berhum.01.1 retrieved September 6, 2022 Subspecies ''S. b. viola'' has a grayish tinge on its back and rump, a fawn to cinnamon belly, and dark violet uppertail coverts and tail. ''S. b. lichtensteini'' is overall a lighter green than the nominate and its tail is silvery. ''S. b. sumichrasti'' males have duller green plumage than the nominate and a purplish gloss to the tail. Females' central tail feathers have a silvery to purple cast. ''S. b. devillei'' has more bronze on the back and rump than the nominate and a purplish to bronzy chestnut tail.


Distribution and habitat

The subspecies of berylline hummingbird are found thus: *''S. b. viola'', in western Mexico from
Sonora Sonora (), officially Estado Libre y Soberano de Sonora (), is one of the 31 states which, along with Mexico City, comprise the Administrative divisions of Mexico, Federal Entities of Mexico. The state is divided into Municipalities of Sonora, 72 ...
to
Michoacán Michoacán, formally Michoacán de Ocampo, officially the Free and Sovereign State of Michoacán de Ocampo, is one of the 31 states which, together with Mexico City, compose the Political divisions of Mexico, Federal Entities of Mexico. The stat ...
and
Guerrero Guerrero, officially the Free and Sovereign State of Guerrero, is one of the 32 states that compose the administrative divisions of Mexico, 32 Federal Entities of Mexico. It is divided into Municipalities of Guerrero, 85 municipalities. The stat ...
and rarely to the southwestern US. *''S. b. beryllina'', central Mexico from
México state The State of Mexico, officially just Mexico, is one of the 32 federal entities of the United Mexican States. Colloquially known as Edomex (from , the abbreviation of , and ), to distinguish it from the name of the whole country, it is the mos ...
south into
Veracruz Veracruz, formally Veracruz de Ignacio de la Llave, officially the Free and Sovereign State of Veracruz de Ignacio de la Llave, is one of the 31 states which, along with Mexico City, comprise the 32 Political divisions of Mexico, Federal Entit ...
and
Oaxaca Oaxaca, officially the Free and Sovereign State of Oaxaca, is one of the 32 states that compose the political divisions of Mexico, Federative Entities of the Mexico, United Mexican States. It is divided into municipalities of Oaxaca, 570 munici ...
*''S. b. lichtensteini'', western
Chiapas Chiapas, officially the Free and Sovereign State of Chiapas, is one of the states that make up the Political divisions of Mexico, 32 federal entities of Mexico. It comprises Municipalities of Chiapas, 124 municipalities and its capital and large ...
in southern Mexico *''S. b. sumichrasti'', extreme southeastern Oaxaca and northern and central Chiapas in southern Mexico *''S. b. devillei'', discontinuously from the other subspecies in southern Guatemala and through El Salvador into central Honduras. The berylline hummingbird primarily inhabits arid landscapes. The wide variety includes dense oak and pine-oak forest, scrublands, deciduous and
thorn forest A thorn forest is a dense scrubland with vegetation characteristic of dry subtropical and warm temperate areas with a seasonal rainfall averaging . Regions Africa Is present in the southwest of Africa with smaller areas in other places of Africa. ...
,
gallery forest A gallery forest is one formed as a corridor along rivers or wetlands, projecting into landscapes that are otherwise only sparsely treed such as savannas, grasslands, or deserts. The gallery forest maintains a more temperate microclimate above th ...
, plantations, and parks and gardens. In elevation it is found from near sea level to the
submontane zone Altitudinal zonation (or elevational zonation) in mountainous regions describes the natural layering of ecosystems that occurs at distinct elevations due to varying environmental conditions. Temperature, humidity, soil composition, and solar radi ...
, though it is most common between .


Behavior


Movement

The berylline hummingbird has movements that vary with geography. It is sedentary in most of its range except the far north, where it migrates southward for winter. It increasingly has wandered and sometimes bred in southeastern Arizona and has occurred in southwestern New Mexico and western Texas. Even in the bulk of its range it appears to make some seasonal elevational changes.


Feeding

The berylline hummingbird is a generalist, foraging for nectar at a wide variety of flowering plants both native and introduced. It feeds at all levels of the forest. Though it will congregate with other hummingbird species at flowering trees, it tends to be dominant over most and often defends feeding territories. In addition to nectar it feeds on
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s captured by hawking from a perch or picking from spider webs.


Breeding

In Oaxaca the berylline hummingbird breeds between June and October with a peak in September; its breeding season in the rest of its range has not been defined. It makes a solid cup nest of grass and other plant fibers bound with spiderweb with lichens on the outside. It typically places it on a horizontal branch up to above the ground. The female incubates the clutch of two eggs; the incubation period has not been determined. Fledging occurs about 20 days after hatch.


Vocalization

The berylline hummingbird's song is somewhat variable, "slightly gruff, high-pitched twittering notes
ith The Ith () is a ridge in Germany's Central Uplands which is up to 439 m high. It lies about 40 km southwest of Hanover and, at 22 kilometers, is the longest line of crags in North Germany. Geography Location The Ith is i ...
lisping introductory notes...rendered 'ssi kirr-i-rr kirr-i-rr', 'sirrr, ki-ti ki-dik' or 'sssi-ir sssiir chit-chit chit-chit-chit...'." It also makes calls that are described as "a hard, buzzy 'dzzzzir' or 'drrzzzt', and a more liquid 'dzzzzrrt' that may be repeated several times".


Status

The
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
has assessed the berylline hummingbird as being of Least Concern. It has a large range and a stable population of about two million mature individuals. No immediate threats have been identified. It varies from uncommon to common in Mexico and south but is very local and rare in the US.


References


External links


Berylline hummingbird photoArticle
state.tx.us

Oiseaux {{Taxonbar, from=Q793835 Saucerottia Hummingbird species of Middle America Birds described in 1830 Taxobox binomials not recognized by IUCN