A berry is a small, pulpy, and often edible
fruit
In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering.
Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a
stone
In geology, rock (or stone) is any naturally occurring solid mass or aggregate of minerals or mineraloid matter. It is categorized by the minerals included, its Chemical compound, chemical composition, and the way in which it is formed. Rocks ...
or
pit although many
pips or seeds may be present.
Common examples of berries in the culinary sense are
strawberries
The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown hybrid plant cultivated worldwide for its fruit. The genus ''Fragaria'', the strawberries, is in the rose family, Rosaceae. The fruit is appreciated f ...
,
raspberries
The raspberry is the edible fruit of several plant species in the genus ''Rubus'' of the Rosaceae, rose family, most of which are in the subgenus ''Rubus#Modern classification, Idaeobatus''. The name also applies to these plants themselves. Ras ...
,
blueberries
Blueberries are a widely distributed and widespread group of perennial flowering plants with blue or purple berries. They are classified in the section ''Cyanococcus'' with the genus ''Vaccinium''. Commercial blueberries—both wild (lowbush) ...
,
blackberries
BlackBerry is a discontinued brand of handheld devices and related mobile services, originally developed and maintained by the Canadian company Research In Motion (RIM, later known as BlackBerry Limited) until 2016. The first BlackBerry device ...
,
white currant
The white currant or whitecurrant is a group of cultivars of the red currant (''Ribes rubrum''), a species of flowering plant in the family Grossulariaceae, native to Europe.
It is sometimes mislabelled as '' Ribes glandulosum'',Darina Allen ...
s,
blackcurrant
The blackcurrant (''Ribes nigrum''), also known as black currant or cassis, is a deciduous shrub in the family Grossulariaceae grown for its edible berries. It is native to temperate parts of central and northern Europe and northern Asia, w ...
s, and
redcurrant
The redcurrant or red currant (''Ribes rubrum'') is a member of the genus ''Ribes'' in the gooseberry family. It is native to western Europe. The species is widely cultivated and has escaped into the wild in many regions.
Description
''Ribes ...
s. In Britain, soft fruit is a
horticultural
Horticulture (from ) is the art and science of growing fruits, vegetables, flowers, trees, shrubs and ornamental plants. Horticulture is commonly associated with the more professional and technical aspects of plant cultivation on a smaller and mo ...
term for such fruits.
The common usage of the term "berry" is different from the scientific or
botanical definition of a berry, which refers to a
fleshy fruit
In botany, fleshy fruits are fruits which are fleshy and brightly coloured, making them attractive to animals which eat them and disperse the seeds. The word 'succulent fruit' is synonymous to fleshy fruit and both words are often used interchange ...
produced from the
ovary
The ovary () is a gonad in the female reproductive system that produces ova; when released, an ovum travels through the fallopian tube/ oviduct into the uterus. There is an ovary on the left and the right side of the body. The ovaries are end ...
of a single flower where the outer layer of the ovary wall develops into an edible fleshy portion(
pericarp
Fruits are the mature ovary or ovaries of one or more flowers. They are found in three main anatomical categories: aggregate fruits, multiple fruits, and simple fruits.
Fruitlike structures may develop directly from the seed itself rather th ...
). The botanical definition includes many fruits that are not commonly known or referred to as berries, such as
grape
A grape is a fruit, botanically a berry, of the deciduous woody vines of the flowering plant genus ''Vitis''. Grapes are a non- climacteric type of fruit, generally occurring in clusters.
The cultivation of grapes began approximately 8,0 ...
s,
tomato
The tomato (, ), ''Solanum lycopersicum'', is a plant whose fruit is an edible Berry (botany), berry that is eaten as a vegetable. The tomato is a member of the nightshade family that includes tobacco, potato, and chili peppers. It originate ...
es,
cucumber
The cucumber (''Cucumis sativus'') is a widely-cultivated creeping vine plant in the family Cucurbitaceae that bears cylindrical to spherical fruits, which are used as culinary vegetables.[eggplant
Eggplant (American English, US, Canadian English, CA, Australian English, AU, Philippine English, PH), aubergine (British English, UK, Hiberno English, IE, New Zealand English, NZ), brinjal (Indian English, IN, Singapore English, SG, Malays ...]
s,
banana
A banana is an elongated, edible fruit – botanically a berry – produced by several kinds of large treelike herbaceous flowering plants in the genus '' Musa''. In some countries, cooking bananas are called plantains, distinguishing the ...
s, and
chili peppers
Chili peppers, also spelled chile or chilli ( ), are varieties of berry-fruit plants from the genus ''Capsicum'', which are members of the nightshade family Solanaceae, cultivated for their pungency. They are used as a spice to add pungency ( ...
. Fruits commonly considered berries but excluded by the botanical definition include strawberries, raspberries, and
blackberries
BlackBerry is a discontinued brand of handheld devices and related mobile services, originally developed and maintained by the Canadian company Research In Motion (RIM, later known as BlackBerry Limited) until 2016. The first BlackBerry device ...
, which are
aggregate fruit
A raspberry_beetle.html" ;"title="raspberry fruit (shown with a raspberry beetle">raspberry fruit (shown with a raspberry beetle larva) is an aggregate fruit, an aggregate of drupelets.
image:Aquilegia vulgaris 004.JPG, The fruit of an ''Aquilegi ...
s, and
mulberries
''Morus'', a genus of flowering plants in the family Moraceae, consists of 19 species of deciduous trees commonly known as mulberries, growing wild and under cultivation in many temperate world regions. Generally, the genus has 64 subordinate ...
, which are
multiple fruit
Multiple fruits, also called collective fruits, are fruiting bodies formed from a cluster of flowers, the ''inflorescence''. Each flower in the inflorescence produces a fruit, but these mature into a single mass. After flowering, the mass is call ...
s.
Watermelons
The watermelon (''Citrullus lanatus'') is a species of flowering plant in the family Cucurbitaceae, that has a large, edible fruit. It is a scrambling and trailing vine-like plant, and is widely cultivated worldwide, with more than 1,000 ...
and
pumpkins are giant berries that fall into the category "
pepos". A plant bearing berries is said to be or .
Berries are eaten worldwide and often used in
jam
Fruit preserves are preparations of fruits whose main preserving agent is sugar and sometimes acid, often stored in glass jars and used as a condiment or spread.
There are many varieties of fruit preserves globally, distinguished by the meth ...
s,
preserves,
cake
Cake is a flour confection usually made from flour, sugar, and other ingredients and is usually baked. In their oldest forms, cakes were modifications of bread, but cakes now cover a wide range of preparations that can be simple or elabor ...
s, or
pie
A pie is a baked dish which is usually made of a pastry dough casing that contains a filling of various sweet or savoury ingredients. Sweet pies may be filled with fruit (as in an apple pie), nuts ( pecan pie), fruit preserves ( jam tart ...
s. Some berries are commercially important. The berry industry varies from country to country as do types of berries cultivated or growing in the wild. Some berries such as raspberries and strawberries have been bred for hundreds of years and are distinct from their wild counterparts, while other berries, such as
lingonberries and
cloudberries, grow almost exclusively in the wild.
While many berries are edible, some are
poisonous
A poison is any chemical substance that is harmful or lethal to living organisms. The term is used in a wide range of scientific fields and industries, where it is often specifically defined. It may also be applied colloquially or figurati ...
to humans, such as those of
deadly nightshade
''Atropa bella-donna'', commonly known as deadly nightshade or belladonna, is a toxic perennial herbaceous plant in the nightshade family Solanaceae, which also includes tomatoes, potatoes and eggplant. It is native to Europe and Western Asia, ...
and
pokeweed
''Phytolacca'' is a genus of perennial plants native to North America, South America and East Asia. Some members of the genus are known as pokeweeds or similar names such as pokebush, pokeberry, pokeroot or poke sallet. Other names for species ...
. Others, such as the
white mulberry
White is the lightest color and is achromatic (having no chroma). It is the color of objects such as snow, chalk, and milk, and is the opposite of black. White objects fully (or almost fully) reflect and scatter all the visible wavelen ...
,
red mulberry, and
elderberry
''Sambucus'' is a genus of between 20 and 30 species of flowering plants in the family Adoxaceae. The various species are commonly referred to as elder, with the flowers as elderflower, and the fruit as elderberry.
Description
Elders are most ...
, are poisonous when
unripe, but are edible when ripe.
History

Berries have been valuable as a food source for humans since before the start of agriculture, and remain among the primary food sources of other
primates
Primates is an order of mammals, which is further divided into the strepsirrhines, which include lemurs, galagos, and lorisids; and the haplorhines, which include tarsiers and simians ( monkeys and apes). Primates arose 74–63 ...
. They were a seasonal staple for early
hunter-gatherers
A hunter-gatherer or forager is a human living in a community, or according to an ancestrally derived lifestyle, in which most or all food is obtained by foraging, that is, by gathering food from local naturally occurring sources, especially w ...
for thousands of years, and wild berry gathering remains a popular activity in Europe and North America today. In time, humans learned to store berries so that they could be used in the winter. They may be made into
fruit preserves
Fruit preserves are preparations of fruits whose main preserving agent is sugar and sometimes acid, often stored in glass jars and used as a condiment or spread.
There are many varieties of fruit preserves globally, distinguished by the meth ...
, and among Native Americans, mixed with meat and fats as
pemmican
Pemmican () (also pemican in older sources) is a mixture of tallow, dried meat, and sometimes dried berries. A calorie-rich food, it can be used as a key component in prepared meals or eaten raw. Historically, it was an important part of indigeno ...
.
Berries also began to be cultivated in Europe and other countries. Some species of blackberries and raspberries of the genus ''
Rubus
''Rubus'' is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, most commonly known as brambles. Fruits of various species are known as raspberries, blackberries, dewberries, and bristleberries. ...
'' have been cultivated since the 17thcentury, while smooth-skinned blueberries and cranberries of the genus ''
Vaccinium
''Vaccinium'' is a common and widespread genus of shrubs or dwarf shrubs in the heath family (Ericaceae). The fruits of many species are eaten by humans and some are of commercial importance, including the cranberry, blueberry, bilberry (wh ...
'' have been cultivated in the United States for over a century.
In Japan, between the 10th and 18thcenturies, the terms and
(
kanji
are logographic Chinese characters, adapted from Chinese family of scripts, Chinese script, used in the writing of Japanese language, Japanese. They were made a major part of the Japanese writing system during the time of Old Japanese and are ...
: ;
katakana
is a Japanese syllabary, one component of the Japanese writing system along with hiragana, kanji and in some cases the Latin script (known as rōmaji).
The word ''katakana'' means "fragmentary kana", as the katakana characters are derived fr ...
:
) referred to many berry crops. The most widely cultivated berry of modern times is the
strawberry
The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown Hybrid (biology), hybrid plant cultivated worldwide for its fruit. The genus ''Fragaria'', the strawberries, is in the rose family, Rosaceae. The fruit ...
, which is produced globally at twice the amount of all other berry crops combined.
The strawberry was mentioned by ancient Romans, who thought it had medicinal properties, but it was then not a staple of agriculture.
Woodland strawberries began to be grown in French gardens in the 14thcentury. The
musk strawberry(''F. moschata''), also known as the strawberry, began to be grown in European gardens in the late 16thcentury. Later, the
Virginia strawberry
''Fragaria virginiana'', known as Virginia strawberry, wild strawberry, common strawberry, or mountain strawberry, is a North American strawberry that grows across much of the United States and southern Canada. It is one of the two species of wi ...
was grown in Europe and the United States. The most commonly consumed strawberry, the
garden strawberry
The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown hybrid plant cultivated worldwide for its fruit. The genus ''Fragaria'', the strawberries, is in the rose family, Rosaceae. The fruit is appreciated ...
(''F. ananassa''), is an accidental hybrid of the Virginia strawberry and a Chilean variety ''
Fragaria chiloensis
''Fragaria chiloensis'', the beach strawberry, Chilean strawberry, or coastal strawberry, is one of two species of wild strawberry that were hybridized to create the modern garden strawberry (''F. × ananassa''). It is native to the Pacific Oce ...
''. It was first noted by a French gardener around the mid 18thcentury that, when ''F. moschata'' and ''F. virginiana'' were planted in between rows of ''F. chiloensis'', the Chilean strawberry would bear abundant and unusually large fruits. Soon after, began to study the
breeding of strawberries and made several discoveries crucial to the science of plant breeding, such as the sexual reproduction of strawberry.
Later, in the early 1800s, English breeders of strawberry made varieties of ''F. ananassa'' which were important in strawberry breeding in Europe, and hundreds of
cultivars
A cultivar is a kind of cultivated plant that people have selected for desired traits and which retains those traits when propagated. Methods used to propagate cultivars include division, root and stem cuttings, offsets, grafting, tissue cult ...
have since been produced through the breeding of strawberries.
Etymology
The
Old English
Old English ( or , or ), or Anglo-Saxon, is the earliest recorded form of the English language, spoken in England and southern and eastern Scotland in the Early Middle Ages. It developed from the languages brought to Great Britain by Anglo-S ...
word () comes from
Proto-Germanic
Proto-Germanic (abbreviated PGmc; also called Common Germanic) is the linguistic reconstruction, reconstructed proto-language of the Germanic languages, Germanic branch of the Indo-European languages.
Proto-Germanic eventually developed from ...
, variously
reconstructed as , , (source also of
Old Norse
Old Norse, also referred to as Old Nordic or Old Scandinavian, was a stage of development of North Germanic languages, North Germanic dialects before their final divergence into separate Nordic languages. Old Norse was spoken by inhabitants ...
,
Middle Dutch
Middle Dutch is a collective name for a number of closely related West Germanic dialects whose ancestor was Old Dutch. It was spoken and written between 1150 and 1500. Until the advent of Modern Dutch after 1500 or , there was no overarching sta ...
,
German
German(s) may refer to:
* Germany, the country of the Germans and German things
**Germania (Roman era)
* Germans, citizens of Germany, people of German ancestry, or native speakers of the German language
** For citizenship in Germany, see also Ge ...
, ,
Old Saxon
Old Saxon (), also known as Old Low German (), was a Germanic language and the earliest recorded form of Low German (spoken nowadays in Northern Germany, the northeastern Netherlands, southern Denmark, the Americas and parts of Eastern Eur ...
,
Gothic, ), which is of unknown origin. This and "
apple
An apple is a round, edible fruit produced by an apple tree (''Malus'' spp.). Fruit trees of the orchard or domestic apple (''Malus domestica''), the most widely grown in the genus, are agriculture, cultivated worldwide. The tree originated ...
" are the only fruit names in modern English which are descended from "native"
Germanic words.
Botanical definition
In
botanical
Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...
terminology, a berry is a simple fruit with
seed
In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
s and pulp produced from the
ovary
The ovary () is a gonad in the female reproductive system that produces ova; when released, an ovum travels through the fallopian tube/ oviduct into the uterus. There is an ovary on the left and the right side of the body. The ovaries are end ...
of a single flower. It is fleshy throughout, except for the seeds. It does not have a special "line of weakness" along which it splits to release the seeds when ripe (i.e. it is
indehiscent
Dehiscence is the splitting of a mature plant structure along a built-in line of weakness to release its contents. This is common among fruits, anthers and sporangia. Sometimes this involves the complete detachment of a part. Structures that ...
). A berry may develop from an ovary with one or more
carpels
Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ...
(the female reproductive structures of a flower). The seeds are usually embedded in the fleshy interior of the ovary, but there are some non-fleshy examples such as
peppers, with air rather than pulp around their seeds. The differences between the everyday and botanical uses of "berry" result in three categories: those fruits that are berries under both definitions; those fruits that are botanical berries but not commonly known as berries; and those parts of plants commonly known as berries that are not botanical berries, and may not even be fruits.
Berries under both definitions include
blueberries
Blueberries are a widely distributed and widespread group of perennial flowering plants with blue or purple berries. They are classified in the section ''Cyanococcus'' with the genus ''Vaccinium''. Commercial blueberries—both wild (lowbush) ...
,
cranberries,
lingonberries, and the fruits of many other members of the
heather family, as well as
gooseberries,
goji berries
Goji, goji berry, or wolfberry () is the sweet fruit of either ''Lycium barbarum'' or ''Lycium chinense'', two closely related species of Lycium, boxthorn in the nightshade family, Solanaceae. ''L. barbarum'' and ''L. chinense'' frui ...
and
elderberries
''Sambucus'' is a genus of between 20 and 30 species of flowering plants in the family Adoxaceae. The various species are commonly referred to as elder, with the flowers as elderflower, and the fruit as elderberry.
Description
Elders are most ...
. The fruits of some "currants" (''
Ribes
''Ribes'' () is a genus of about 200 known species of flowering plants, most of them native to the temperate regions of the Northern Hemisphere. The species may be known as various kinds of currants, such as redcurrants, blackcurrants, and White ...
'' species), such as
blackcurrant
The blackcurrant (''Ribes nigrum''), also known as black currant or cassis, is a deciduous shrub in the family Grossulariaceae grown for its edible berries. It is native to temperate parts of central and northern Europe and northern Asia, w ...
s,
red currants and
white currant
The white currant or whitecurrant is a group of cultivars of the red currant (''Ribes rubrum''), a species of flowering plant in the family Grossulariaceae, native to Europe.
It is sometimes mislabelled as '' Ribes glandulosum'',Darina Allen ...
s, are botanical berries, and are treated as horticultural berries (or as soft fruit in the UK), even though their most commonly used names do not include the word "berry".
Botanical berries not commonly known as berries include
banana
A banana is an elongated, edible fruit – botanically a berry – produced by several kinds of large treelike herbaceous flowering plants in the genus '' Musa''. In some countries, cooking bananas are called plantains, distinguishing the ...
s,
tomato
The tomato (, ), ''Solanum lycopersicum'', is a plant whose fruit is an edible Berry (botany), berry that is eaten as a vegetable. The tomato is a member of the nightshade family that includes tobacco, potato, and chili peppers. It originate ...
es,
grape
A grape is a fruit, botanically a berry, of the deciduous woody vines of the flowering plant genus ''Vitis''. Grapes are a non- climacteric type of fruit, generally occurring in clusters.
The cultivation of grapes began approximately 8,0 ...
s,
eggplant
Eggplant (American English, US, Canadian English, CA, Australian English, AU, Philippine English, PH), aubergine (British English, UK, Hiberno English, IE, New Zealand English, NZ), brinjal (Indian English, IN, Singapore English, SG, Malays ...
s (aubergines),
persimmon
The persimmon () is the edible fruit of a number of species of trees in the genus '' Diospyros''. The most widely cultivated of these is the Chinese and Japanese kaki persimmon, ''Diospyros kaki''. In 2022, China produced 77% of the world's p ...
s,
watermelon
The watermelon (''Citrullus lanatus'') is a species of flowering plant in the family Cucurbitaceae, that has a large, edible fruit. It is a Glossary of botanical terms#scandent, scrambling and trailing vine-like plant, and is plant breeding ...
s, and
pumpkin
A pumpkin is a cultivar, cultivated winter squash in the genus ''Cucurbita''. The term is most commonly applied to round, orange-colored squash varieties, but does not possess a scientific definition. It may be used in reference to many dif ...
s.
There are several different kinds of fruits which are commonly called berries, but are not botanical berries.
Blackberries
BlackBerry is a discontinued brand of handheld devices and related mobile services, originally developed and maintained by the Canadian company Research In Motion (RIM, later known as BlackBerry Limited) until 2016. The first BlackBerry device ...
,
raspberries
The raspberry is the edible fruit of several plant species in the genus ''Rubus'' of the Rosaceae, rose family, most of which are in the subgenus ''Rubus#Modern classification, Idaeobatus''. The name also applies to these plants themselves. Ras ...
, and
strawberries
The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown hybrid plant cultivated worldwide for its fruit. The genus ''Fragaria'', the strawberries, is in the rose family, Rosaceae. The fruit is appreciated f ...
are kinds of
aggregate fruit
A raspberry_beetle.html" ;"title="raspberry fruit (shown with a raspberry beetle">raspberry fruit (shown with a raspberry beetle larva) is an aggregate fruit, an aggregate of drupelets.
image:Aquilegia vulgaris 004.JPG, The fruit of an ''Aquilegi ...
;
they contain seeds from different ovaries of a single flower. In aggregate fruits like blackberries, the individual "fruitlets" making up the fruit can be clearly seen. The fruits of
blackthorn
''Prunus spinosa'', called blackthorn or sloe, is an Old World species of flowering plant in the rose family, Rosaceae. It is locally naturalized in parts of the New World.
The fruits are used to make sloe gin in Britain and patxaran in Basq ...
may be called "sloe berries", but botanically are small
stone fruit
In botany, a drupe (or stone fruit) is a type of fruit in which an outer fleshy part (exocarp, or skin, and mesocarp, or flesh) surrounds a single shell (the ''pip'' (UK), ''pit'' (US), ''stone'', or ''pyrena'') of hardened endocarp with a seed ...
s or drupes, like
plum
A plum is a fruit of some species in Prunus subg. Prunus, ''Prunus'' subg. ''Prunus'.'' Dried plums are often called prunes, though in the United States they may be labeled as 'dried plums', especially during the 21st century.
Plums are ...
s or
apricot
An apricot (, ) is a fruit, or the tree that bears the fruit, of several species in the genus ''Prunus''.
Usually an apricot is from the species '' P. armeniaca'', but the fruits of the other species in ''Prunus'' sect. ''Armeniaca'' are also ...
s.
Juniper
Junipers are coniferous trees and shrubs in the genus ''Juniperus'' ( ) of the cypress family Cupressaceae. Depending on the taxonomy, between 50 and 67 species of junipers are widely distributed throughout the Northern Hemisphere as far south ...
s and
yews are commonly said to have berries, but these plants do not produce botanical fruits at all: they are
gymnosperms
The gymnosperms ( ; ) are a group of woody, perennial Seed plant, seed-producing plants, typically lacking the protective outer covering which surrounds the seeds in flowering plants, that include Pinophyta, conifers, cycads, Ginkgo, and gnetoph ...
, specifically
conifers
Conifers () are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All e ...
, not
angiosperms
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed within a fruit. T ...
(flowering plants). Their "berries" are highly-modified seed-bearing
cones
In geometry, a cone is a three-dimensional figure that tapers smoothly from a flat base (typically a circle) to a point not contained in the base, called the ''apex'' or '' vertex''.
A cone is formed by a set of line segments, half-lines, ...
. In
juniper berries
A juniper berry is the female seed cone produced by the various species of junipers. It is not a true berry but a cone with unusually fleshy and merged scales called a galbulus, which gives it a berry-like appearance. The cones from a handful of ...
, used to flavour
gin
Gin () is a distilled alcoholic drink flavoured with juniper berries and other botanical ingredients.
Gin originated as a medicinal liquor made by monks and alchemists across Europe. The modern gin was modified in Flanders and the Netherlands ...
, the cone scales, which are hard and woody in most conifers, are instead soft and fleshy when ripe. The bright red berries of yews consist of a fleshy outgrowth(
aril
An aril (), also called arillus, is a specialized outgrowth from a seed that partly or completely covers the seed. An arillode, or false aril, is sometimes distinguished: whereas an aril grows from the attachment point of the seed to the ova ...
) almost enclosing the poisonous seed. The resemblance of these plant structures to botanical berries provides a striking example of
convergent evolution
Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last comm ...
in different plant
clades
In biology, a clade (), also known as a monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach to taxonomy ...
.
Cultivation
Strawberries
The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown hybrid plant cultivated worldwide for its fruit. The genus ''Fragaria'', the strawberries, is in the rose family, Rosaceae. The fruit is appreciated f ...
have been grown in gardens in Europe since the 14thcentury.
Blueberries
Blueberries are a widely distributed and widespread group of perennial flowering plants with blue or purple berries. They are classified in the section ''Cyanococcus'' with the genus ''Vaccinium''. Commercial blueberries—both wild (lowbush) ...
were domesticated starting in 1911, with the first commercial crop in 1916.
Huckleberries
Huckleberry is a name used in North America for several plants in the family Ericaceae, in two closely related genera: ''Vaccinium'' and ''Gaylussacia''.
Nomenclature
The name 'huckleberry' is a North American variation of the English dialectal ...
of all varieties are not fully domesticated, but domestication was attempted from 1994 to 2010 for the economically significant
western huckleberry. Many other varieties of ''
Vaccinium
''Vaccinium'' is a common and widespread genus of shrubs or dwarf shrubs in the heath family (Ericaceae). The fruits of many species are eaten by humans and some are of commercial importance, including the cranberry, blueberry, bilberry (wh ...
'' are likewise not domesticated, with some being of commercial importance.
Agricultural methods
Like most other food crops, berries are commercially grown, with both conventional pest management and
integrated pest management
Integrated pest management (IPM), also known as integrated pest control (IPC) integrates both chemical and non-chemical practices for economic control of pests. The UN's Food and Agriculture Organization defines IPM as "the careful consideratio ...
(IPM) practices.
Organically certified berries are becoming more widely available.
:5
Many soft fruit berries require a period of temperatures between for breaking dormancy. In general,
strawberries
The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown hybrid plant cultivated worldwide for its fruit. The genus ''Fragaria'', the strawberries, is in the rose family, Rosaceae. The fruit is appreciated f ...
require 200–300hours,
blueberries
Blueberries are a widely distributed and widespread group of perennial flowering plants with blue or purple berries. They are classified in the section ''Cyanococcus'' with the genus ''Vaccinium''. Commercial blueberries—both wild (lowbush) ...
650–850hours,
blackberries
BlackBerry is a discontinued brand of handheld devices and related mobile services, originally developed and maintained by the Canadian company Research In Motion (RIM, later known as BlackBerry Limited) until 2016. The first BlackBerry device ...
700hours,
raspberries
The raspberry is the edible fruit of several plant species in the genus ''Rubus'' of the Rosaceae, rose family, most of which are in the subgenus ''Rubus#Modern classification, Idaeobatus''. The name also applies to these plants themselves. Ras ...
800–1700hours,
currants and
gooseberries 800–1500hours, and
cranberries 2000hours.
However, too low a temperature will kill the crops: blueberries do not tolerate temperatures below , raspberries, depending on variety, may tolerate as low as , and blackberries are injured below .
Spring frosts are, however, much more damaging to berry crops than low winter temperatures. Sites with moderate slopes(), facing north or east in the
Northern Hemisphere
The Northern Hemisphere is the half of Earth that is north of the equator. For other planets in the Solar System, north is defined by humans as being in the same celestial sphere, celestial hemisphere relative to the invariable plane of the Solar ...
, near large bodies of water, which regulate spring temperature, are considered ideal in preventing spring frost injury to the new leaves and flowers.
All berry crops have shallow
root systems
In vascular plants, the roots are the organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often below the surfa ...
.
Many US
land-grant university
A land-grant university (also called land-grant college or land-grant institution) is an institution of higher education in the United States designated by a state to receive the benefits of the Morrill Land-Grant Acts, Morrill Acts of 1862 and ...
extension offices suggest that strawberries should not be planted more than five years on the same site, due to the danger of
black root rot (though many other illnesses go by the same name), which in the past has been controlled in major commercial production by annual
methyl bromide
Bromomethane, commonly known as methyl bromide, is an organobromine compound with chemical formula, formula Carbon, CHydrogen, H3Bromine, Br. This colorless, odorless, nonflammable gas is Bromine cycle, produced both industrially and biologically ...
fumigation
but which is largely prohibited now. Besides the number of years in production,
soil compaction
In geotechnical engineering, soil compaction is the process in which stress applied to a soil causes densification as air is displaced from the pores between the soil grains. When stress is applied that causes densification due to water (or other ...
, the frequency of fumigation, and herbicide usages increase the appearance of black root rot in strawberries.
Raspberries, blackberries, strawberries, and many other berries are susceptible to
verticillium wilt
Verticillium wilt is a wilt disease affecting over 350 species of eudicot plants. It is caused by six species of '' Verticillium'' fungi: ''V. dahliae'', ''V. albo-atrum'', ''V. longisporum'', ''V. nubilum'', ''V. theobr ...
. Blueberries and cranberries grow poorly if the clay or silt content of the soil is more than 20%, while most other berries tolerate a wide range of
soil types
A soil type is a taxonomic unit in soil science. All soils that share a certain set of well-defined properties form a distinctive soil type. Soil type is a technical term of soil classification, the science that deals with the systematic categor ...
.
For most berry crops, the ideal soil is well drained sandy
loam
Loam (in geology and soil science) is soil composed mostly of sand (particle size > ), silt (particle size > ), and a smaller amount of clay (particle size < ). By weight, its mineral composition is about 40–40–20% concentration of sand–si ...
, with
a pH of 6.2–6.8 and a moderate to high organic content; however, blueberries have an ideal pH of 4.2–4.8 and can be grown on muck soils, while blueberries and cranberries prefer poorer soils with lower
cation exchange
Ion exchange is a reversible interchange of one species of ion present in an insoluble solid with another of like charge present in a solution surrounding the solid. Ion exchange is used in softening or demineralizing of water, purification of ch ...
, lower calcium, and lower levels of phosphorus.
Growing most berries organically requires the use of proper
crop rotation
Crop rotation is the practice of growing a series of different types of crops in the same area across a sequence of growing seasons. This practice reduces the reliance of crops on one set of nutrients, pest and weed pressure, along with the pro ...
, the right mix of
cover crops, and the cultivation of the correct
beneficial microorganisms in the soil.
As blueberries and cranberries thrive in soils that are not hospitable to most other plants, and conventional fertilizers are toxic to them, the primary concern when growing them organically is bird management.
Postharvest small fruit berries are generally stored at
relative humidity
Humidity is the concentration of water vapor present in the air. Water vapor, the gaseous state of water, is generally invisible to the human eye. Humidity indicates the likelihood for precipitation (meteorology), precipitation, dew, or fog t ...
and .
Cranberries, however, are frost sensitive, and should be stored at .
Blueberries are the only berries that respond to
ethylene
Ethylene (IUPAC name: ethene) is a hydrocarbon which has the formula or . It is a colourless, flammable gas with a faint "sweet and musky" odour when pure. It is the simplest alkene (a hydrocarbon with carbon–carbon bond, carbon–carbon doub ...
, but flavor does not improve after harvest, so they require the same treatment as other berries. Removal of ethylene may reduce disease and spoilage in all berries.
Precooling within one to two hours post-harvest to storage temperature, generally, via forced air cooling increases the storage life of berries by about a third.
Under optimum storage conditions, raspberries and blackberries last for two to five days, strawberries 7–10days, blueberries two to four weeks, and cranberries two to four months.
Berries can be shipped under high carbon dioxide or
modified atmosphere of carbon dioxide for high carbon dioxide or carbon dioxide and oxygen for a modified atmosphere container to increase shelf life and prevent
grey mold rot.
Breeding
Several discoveries in the science of breeding berries were made in the 18thcentury by in his work on strawberries.
In the traditional technique of
plant breeding
Plant breeding is the science of changing the traits of plants in order to produce desired characteristics. It is used to improve the quality of plant products for use by humans and animals. The goals of plant breeding are to produce crop varie ...
, berries with specific desirable characteristics were chosen and allowed to sexually reproduce with other berries, and offspring with improved traits could then be selected and used for further crossing. Plants may be
hybridized with different species within the same
genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
; hybridization between different genus may also be possible, but more difficult. Breeding may seek to increase the size and yield of the fruit, improve the flavor and quality of its nutrient content, such as antioxidants, expand the harvest season, and produce
cultivars
A cultivar is a kind of cultivated plant that people have selected for desired traits and which retains those traits when propagated. Methods used to propagate cultivars include division, root and stem cuttings, offsets, grafting, tissue cult ...
with resistance to diseases, tolerance of hot or cold conditions, and other desirable traits.
Advancements in
molecular biology
Molecular biology is a branch of biology that seeks to understand the molecule, molecular basis of biological activity in and between Cell (biology), cells, including biomolecule, biomolecular synthesis, modification, mechanisms, and interactio ...
and
genetic engineering
Genetic engineering, also called genetic modification or genetic manipulation, is the modification and manipulation of an organism's genes using technology. It is a set of Genetic engineering techniques, technologies used to change the genet ...
allow for a more efficient and better targeted approach in the selection for a desirable
genotype
The genotype of an organism is its complete set of genetic material. Genotype can also be used to refer to the alleles or variants an individual carries in a particular gene or genetic location. The number of alleles an individual can have in a ...
, via
marker-assisted selection
Marker assisted selection or marker aided selection (MAS) is an indirect selection process where a trait of interest is selected based on a marker ( morphological, biochemical or DNA/RNA variation) linked to a trait of interest (e.g. productivit ...
, for example.
Genetic modification
Genetic engineering, also called genetic modification or genetic manipulation, is the modification and manipulation of an organism's genes using technology. It is a set of technologies used to change the genetic makeup of cells, including th ...
techniques can also be used for breeding berries.
[
]
Horticultural soft fruit berries
Some fruit not commonly referred to as berries and not always botanically berries are included by US land-grant university
A land-grant university (also called land-grant college or land-grant institution) is an institution of higher education in the United States designated by a state to receive the benefits of the Morrill Land-Grant Acts, Morrill Acts of 1862 and ...
extension offices in their guides for berry cultivation, or in guides for identifying local wild edible and non-edible berries. Examples include beach plums, American persimmons, pawpaws, Pacific crabapples, and prickly pears.
Commercial production
In the year 2005, there were of land worldwide cultivating berries, with produced.:4
Economics
In certain regions, berrypicking can be a large part of the economy, and it is becoming increasingly common for western European
Western Europe is the western region of Europe. The region's extent varies depending on context.
The concept of "the West" appeared in Europe in juxtaposition to "the East" and originally applied to the Western half of the ancient Mediterranean ...
countries such as Sweden and Finland to import cheap labor from Thailand or Bulgaria to do berry picking. This practice has come under scrutiny in the past years because of the low wages and poor living standard for the "berry-pickers", as well as the lack of worker safety.
In the late 2010s in the US, reduced migration from Mexico and Central America and increased minimum wage
A minimum wage is the lowest remuneration that employers can legally pay their employees—the price floor below which employees may not sell their labor. List of countries by minimum wage, Most countries had introduced minimum wage legislation b ...
standards have made finding "stoop-work" labourers to pick the strawberry crop difficult and costly.
Phytochemicals and color
Once ripened, berries have a contrasting color to their background (often of green leaves), making them visible and attractive to frugivorous
A frugivore ( ) is an animal that thrives mostly on raw fruits or succulent fruit-like produce of plants such as roots, shoots, nuts and seeds. Approximately 20% of mammalian herbivores eat fruit. Frugivores are highly dependent on the abundance ...
animals and birds. This assists the wide dispersal of the plants' seeds.[
Berry colors are due to natural ]phytochemical
Phytochemicals are naturally-occurring chemicals present in or extracted from plants. Some phytochemicals are nutrients for the plant, while others are metabolites produced to enhance plant survivability and reproduction.
The fields of ext ...
s, including plant pigments, such as anthocyanin
Anthocyanins (), also called anthocyans, are solubility, water-soluble vacuole, vacuolar pigments that, depending on their pH, may appear red, purple, blue, or black. In 1835, the German pharmacist Ludwig Clamor Marquart named a chemical compou ...
s, together with other flavonoid
Flavonoids (or bioflavonoids; from the Latin word ''flavus'', meaning yellow, their color in nature) are a class of polyphenolic secondary metabolites found in plants, and thus commonly consumed in the diets of humans.
Chemically, flavonoids ...
s localized mainly in berry skins, seeds and leaves.[ Although berry pigments have ]antioxidant
Antioxidants are Chemical compound, compounds that inhibit Redox, oxidation, a chemical reaction that can produce Radical (chemistry), free radicals. Autoxidation leads to degradation of organic compounds, including living matter. Antioxidants ...
properties ', there is no physiological
Physiology (; ) is the science, scientific study of function (biology), functions and mechanism (biology), mechanisms in a life, living system. As a branches of science, subdiscipline of biology, physiology focuses on how organisms, organ syst ...
evidence established to date that berry pigments have actual antioxidant or any other functions within the human body. Consequently, it is not permitted to claim that foods containing polyphenols have antioxidant health value on product labels in the United States or Europe.
Culinary significance
Use in baked goods
Berries are commonly used in pies or tarts
A tart is a baked dish consisting of a filling over a pastry base with an open top not covered with pastry. The pastry is usually shortcrust pastry; the filling may be sweet or savoury, though modern tarts are usually fruit-based, sometimes with ...
, such as blueberry pie
Blueberry pie is a pie with a blueberry filling. Blueberry pie is readily made because it does not require pitting or peeling of fruit. It usually has a top and bottom crust. The top crust can be circular, but the pie can also have a crumble c ...
, blackberry pie
Blackberry pie is a pie composed of blackberry filling, usually in the form of either blackberry jam, actual blackberries themselves, or some combination thereof. Blackberry pie is Tartness, tart, so it requires more sugar than blueberry pie. Bla ...
, and strawberry pie
Strawberry pie is a dessert food consisting mainly of strawberries and sugar in a pie crust, sometimes with gelatin. It is often served with whipped cream.
Preparation
Fresh strawberry pies are usually made with in-season sweet berries, while ...
.
Berries are often used in baking, such as blueberry muffins, blackberry muffins, berry cobblers, berry crisps
Potato chips (North American English and Australian English; often just chip) or crisp (British English and Hiberno-English) are thin slices of potato (or a thin deposit of potato paste) that has been deep fried, baked, or air fried until ...
, berry cakes, berry buckles, berry crumb cakes, berry tea cakes, and berry cookies. Berries are commonly incorporated whole into the batter for baking, and care is often taken so as to not burst the berries. Frozen or dried berries may be preferable for some baked berry products. Fresh berries are also often incorporated into baked berry desserts, sometimes with cream
Cream is a dairy product composed of the higher-fat layer skimmed from the top of milk before homogenization. In un-homogenized milk, the fat, which is less dense, eventually rises to the top. In the industrial production of cream, this proces ...
, either as a filling to the dessert or as a topping.
Beverages
Berries are often added to water and/or juiced, as in cranberry juice
Cranberry juice is the liquid juice of the cranberry a fruit recognized for its bright red color, tart taste, and versatility for product manufacturing. Major cranberry products include cranberry juice, dried cranberry, cranberry sauce, frozen ...
, which accounts for 95% of cranberry crop usage, blueberry
Blueberries are a widely distributed and widespread group of perennial flowering plants with blue or purple berries. They are classified in the section ''Cyanococcus'' with the genus ''Vaccinium''. Commercial blueberries—both wild (lowbush) ...
juice, raspberry juice, goji berry juice, acai juice, aronia berry juice, and strawberry
The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown Hybrid (biology), hybrid plant cultivated worldwide for its fruit. The genus ''Fragaria'', the strawberries, is in the rose family, Rosaceae. The fruit ...
juice. Wine
Wine is an alcoholic drink made from Fermentation in winemaking, fermented fruit. Yeast in winemaking, Yeast consumes the sugar in the fruit and converts it to ethanol and carbon dioxide, releasing heat in the process. Wine is most often made f ...
is the principal alcoholic beverage
Drinks containing alcohol (drug), alcohol are typically divided into three classes—beers, wines, and Distilled beverage, spirits—with alcohol content typically between 3% and 50%. Drinks with less than 0.5% are sometimes considered Non-al ...
made from berries(grapes). Fruit wine
Fruit wines are fermented alcoholic beverages made from a variety of base ingredients (other than grapes); they may also have additional flavors taken from fruits, flowers, and herbs. This definition is sometimes broadened to include any alcohol ...
s are commonly made out of other berries. In most cases, sugars must be added to the berry juices in the process of chaptalization
Chaptalization is the process of adding sugar to unfermented grape must in order to increase the alcohol content after fermentation. The technique is named after its developer, the French chemist Jean-Antoine-Claude Chaptal. This process is not i ...
to increase the alcohol content of the wine. Examples of fruit wines made from berries include: elderberry wine, strawberry wine, blueberry wine, blackberry wine, redcurrant wine, huckleberry wine, goji wine and cranberry wine. Berries are used in some styles of beer, particularly framboise (made with raspberry) and other fruit lambic
Lambic ( , ; ) is a type of beer brewed in the Pajottenland region of Belgium southwest of Brussels since the 13th century. Types of lambic beer include gueuze, kriek lambic, and framboise. Lambic differs from most other beers in that it is Bre ...
s.
Dried
Currants, raisins
A raisin is a dried grape. Raisins are produced in many regions of the world and may be eaten raw or used in cooking, baking, and brewing. In the United Kingdom, Ireland, New Zealand, Australia and South Africa, the word ''raisin'' is reserv ...
and sultanas are examples of dried grape berries, and many other commercially important berries are available in dried form.
Fruit preserves
Berries are perishable fruits with a short shelf life, and are often preserved by drying
Drying is a mass transfer process consisting of the removal of water or another solvent by evaporation from a solid, semi-solid or liquid. This process is often used as a final production step before selling or packaging products. To be consider ...
, freezing
Freezing is a phase transition in which a liquid turns into a solid when its temperature is lowered below its freezing point.
For most substances, the melting and freezing points are the same temperature; however, certain substances possess dif ...
, pickling
Pickling is the process of food preservation, preserving or extending the shelf life of food by either Anaerobic organism, anaerobic fermentation (food), fermentation in brine or immersion in vinegar. The pickling procedure typically affects t ...
or making fruit preserves
Fruit preserves are preparations of fruits whose main preserving agent is sugar and sometimes acid, often stored in glass jars and used as a condiment or spread.
There are many varieties of fruit preserves globally, distinguished by the meth ...
. Berries such as blackberry
BlackBerry is a discontinued brand of handheld devices and related mobile services, originally developed and maintained by the Canadian company Research In Motion (RIM, later known as BlackBerry Limited) until 2016. The first BlackBerry device ...
, blueberry
Blueberries are a widely distributed and widespread group of perennial flowering plants with blue or purple berries. They are classified in the section ''Cyanococcus'' with the genus ''Vaccinium''. Commercial blueberries—both wild (lowbush) ...
, boysenberry
The boysenberry is a cross between the European raspberry (''Rubus idaeus''), European blackberry ('' Rubus fruticosus''), American dewberry ('' Rubus aboriginum''), and loganberry (''Rubus'' × ''loganobaccus'').
Description
Boysenberrie ...
, lingonberry
''Vaccinium vitis-idaea'' is a small evergreen shrub in the heath family, Ericaceae. It is known colloquially as the lingonberry, partridgeberry, foxberry, mountain cranberry, or cowberry. It is native to boreal forest and Arctic tundra through ...
, loganberry
The loganberry (''Rubus'' × ''loganobaccus'') is a hybrid of the North American blackberry ('' Rubus ursinus'') and the European raspberry (''Rubus idaeus''), accidentally bred in 1881 by James Harvey Logan, for whom they are named. They are ...
, raspberry
The raspberry is the edible fruit of several plant species in the genus ''Rubus'' of the Rosaceae, rose family, most of which are in the subgenus ''Rubus#Modern classification, Idaeobatus''. The name also applies to these plants themselves. Ras ...
, and strawberry
The garden strawberry (or simply strawberry; ''Fragaria × ananassa'') is a widely grown Hybrid (biology), hybrid plant cultivated worldwide for its fruit. The genus ''Fragaria'', the strawberries, is in the rose family, Rosaceae. The fruit ...
are often used in jams and jellies. In the United States, Native Americans were "the first to make preserves from blueberries".
Other usages
Chefs have created quick pickled soft fruit, such as blackberries, strawberries, and blueberries. Strawberries can be battered and quickly fried in a deep fryer
A deep fryer (or deep fat fryer), sometimes referred to by the French name friteuse, is a kitchen appliance used to cook foods by full immersion in hot oil—deep frying. The cooking oil (or fats) are typically between temperatures of .
Long ...
. Sauces made from berries, such as cranberry sauce, can be frozen until hard, battered, and deep fried. Cranberry sauce is a traditional food item for Thanksgiving
Thanksgiving is a national holiday celebrated on various dates in October and November in the United States, Canada, Saint Lucia, Liberia, and unofficially in countries like Brazil and Germany. It is also observed in the Australian territory ...
, and similar sauces can be made from many other berries such as blueberries, raspberries, blackberries, and huckleberries.
In culture
Dye
Berries have been used in some cultures for dyeing
Dyeing is the application of dyes or pigments on textile materials such as fibers, yarns, and fabrics with the goal of achieving color with desired color fastness. Dyeing is normally done in a special solution containing dyes and particular ...
. Many berries contain juices that can easily stain, affording use as a natural dye. For example, blackberries are useful for making dyes, especially when ripe berries can easily release juice to produce a colorfast
Colour fastness is a term—used in the dyeing of textile materials—that characterizes a material's colour's resistance to Color fade, fading or Glossary of dyeing terms#R, running. Colour fastness is the property of dyes and it is directly propo ...
effect. ''Rubus'' berries, such as blackberry, raspberry, black raspberry
Black raspberry is a common name for three species of the genus ''Rubus'':
*''Rubus leucodermis'', native to western North America
*''Rubus occidentalis
''Rubus occidentalis'' is a species of ''Rubus'' native to eastern North America. I ...
, dewberry
The dewberries are a group of species in the genus '' Rubus'', section ''Rubus'', closely related to the blackberries. They are small, trailing (rather than upright or high-arching) brambles with aggregate fruits, reminiscent of the raspbe ...
, loganberry
The loganberry (''Rubus'' × ''loganobaccus'') is a hybrid of the North American blackberry ('' Rubus ursinus'') and the European raspberry (''Rubus idaeus''), accidentally bred in 1881 by James Harvey Logan, for whom they are named. They are ...
, and thimbleberry all produce dye colors. These were once used by Native Americans. In Hawaii, the native Hawaiian raspberry called was used to dye tapa cloth
Tapa cloth (or simply ''tapa'') is a barkcloth made in the islands of the Pacific Ocean, primarily in Tonga, Samoa and Fiji, but as far afield as Niue, Cook Islands, Futuna, Solomon Islands, Java, New Zealand, Vanuatu, Papua New Guinea and Ha ...
with lavender and pink hues, whereas berries from the dianella lily were used for blue coloration, and berries from the black nightshade were used to produce green coloration.
Research
Berry consumption is under preliminary research for the potential to improve nutrition and affect chronic diseases. A 2016 meta-analysis
Meta-analysis is a method of synthesis of quantitative data from multiple independent studies addressing a common research question. An important part of this method involves computing a combined effect size across all of the studies. As such, th ...
found that berry consumption can significantly lower body mass index
Body mass index (BMI) is a value derived from the mass (Mass versus weight, weight) and height of a person. The BMI is defined as the human body weight, body mass divided by the square (algebra), square of the human height, body height, and is ...
, low density lipoprotein
Low-density lipoprotein (LDL) is one of the five major groups of lipoprotein that transport all fat molecules around the body in extracellular water. These groups, from least dense to most dense, are chylomicrons (aka ULDL by the overall densit ...
(LDL) and systolic blood pressure
Blood pressure (BP) is the pressure of circulating blood against the walls of blood vessels. Most of this pressure results from the heart pumping blood through the circulatory system. When used without qualification, the term "blood pressure" r ...
.
See also
* List of berries
References
Further reading
*
External links
* of the National Berry Crops Initiative (United States)
{{Authority control
Fruit morphology
Non-timber forest products