Bematistes Epaea
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''Bematistes epaea'', the common bematistes, is a
butterfly Butterflies are winged insects from the lepidopteran superfamily Papilionoidea, characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The oldest butterfly fossi ...
in the family
Nymphalidae The Nymphalidae are the largest family of butterflies, with more than 6,000 species distributed throughout most of the world. Belonging to the superfamily Papilionoidea, they are usually medium-sized to large butterflies. Most species ha ...
. It is found in
Senegal Senegal, officially the Republic of Senegal, is the westernmost country in West Africa, situated on the Atlantic Ocean coastline. It borders Mauritania to Mauritania–Senegal border, the north, Mali to Mali–Senegal border, the east, Guinea t ...
,
Guinea-Bissau Guinea-Bissau, officially the Republic of Guinea-Bissau, is a country in West Africa that covers with an estimated population of 2,026,778. It borders Senegal to Guinea-Bissau–Senegal border, its north and Guinea to Guinea–Guinea-Bissau b ...
,
Sierra Leone Sierra Leone, officially the Republic of Sierra Leone, is a country on the southwest coast of West Africa. It is bordered to the southeast by Liberia and by Guinea to the north. Sierra Leone's land area is . It has a tropical climate and envi ...
,
Liberia Liberia, officially the Republic of Liberia, is a country on the West African coast. It is bordered by Sierra Leone to Liberia–Sierra Leone border, its northwest, Guinea to Guinea–Liberia border, its north, Ivory Coast to Ivory Coast–Lib ...
,
Ivory Coast Ivory Coast, also known as Côte d'Ivoire and officially the Republic of Côte d'Ivoire, is a country on the southern coast of West Africa. Its capital city of Yamoussoukro is located in the centre of the country, while its largest List of ci ...
,
Ghana Ghana, officially the Republic of Ghana, is a country in West Africa. It is situated along the Gulf of Guinea and the Atlantic Ocean to the south, and shares borders with Côte d’Ivoire to the west, Burkina Faso to the north, and Togo to t ...
,
Togo Togo, officially the Togolese Republic, is a country in West Africa. It is bordered by Ghana to Ghana–Togo border, the west, Benin to Benin–Togo border, the east and Burkina Faso to Burkina Faso–Togo border, the north. It is one of the le ...
,
Benin Benin, officially the Republic of Benin, is a country in West Africa. It was formerly known as Dahomey. It is bordered by Togo to the west, Nigeria to the east, Burkina Faso to the north-west, and Niger to the north-east. The majority of its po ...
,
Nigeria Nigeria, officially the Federal Republic of Nigeria, is a country in West Africa. It is situated between the Sahel to the north and the Gulf of Guinea in the Atlantic Ocean to the south. It covers an area of . With Demographics of Nigeria, ...
,
Cameroon Cameroon, officially the Republic of Cameroon, is a country in Central Africa. It shares boundaries with Nigeria to the west and north, Chad to the northeast, the Central African Republic to the east, and Equatorial Guinea, Gabon, and the R ...
,
Equatorial Guinea Equatorial Guinea, officially the Republic of Equatorial Guinea, is a country on the west coast of Central Africa. It has an area of . Formerly the colony of Spanish Guinea, its post-independence name refers to its location both near the Equ ...
, the
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,
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, the
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Uganda Uganda, officially the Republic of Uganda, is a landlocked country in East Africa. It is bordered to the east by Kenya, to the north by South Sudan, to the west by the Democratic Republic of the Congo, to the south-west by Rwanda, and to the ...
,
Kenya Kenya, officially the Republic of Kenya, is a country located in East Africa. With an estimated population of more than 52.4 million as of mid-2024, Kenya is the 27th-most-populous country in the world and the 7th most populous in Africa. ...
,
Tanzania Tanzania, officially the United Republic of Tanzania, is a country in East Africa within the African Great Lakes region. It is bordered by Uganda to the northwest; Kenya to the northeast; the Indian Ocean to the east; Mozambique and Malawi to t ...
,
Malawi Malawi, officially the Republic of Malawi, is a landlocked country in Southeastern Africa. It is bordered by Zambia to the west, Tanzania to the north and northeast, and Mozambique to the east, south, and southwest. Malawi spans over and ...
and
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.


Description

''P. epaea'' is distinguished by having the hindmarginal spot of the forewing narrow and of fairly uniform breadth, at the hindmargin not or scarcely broader than half the hindmargin and not filling up the base of cellule 2, while the black marginal band on the upperside of the hindwing is in the d suddenly much widened at the apex, but in the female often very broad. On the under surface of the hinclwing the basal area is brighter red-yellow than the other parts of the wing. - ''epaea'' Cr. (58 a). male- Ground-colour of the hindwing above, hindmarginal spot and subapical band of the fore wing yellow-brown; hind wing beneath in the distal part more or less suffused with brown-grey and in the middle somewhat lighter. female Subapical band and hindmarginal spot of the forewing white and also a median band on the hindwing, 7 mm. in breadth, which forms a continuation of the hindmarginal spot of the forewing; the whole distal half of the hindwing, on the other hand, black-grey on both surfaces, with black veins and streaks. Senegambia to Angola and the upper Congo. -female -ab. ''lutosa'' Suff. is a female form which differs in having the white markings of the typical female light clay-yellow and the proximal part of the dark distal half of the hindwing above broadly light brown to dark yellow-brown or olive-brown; the pure black outer part of the distal half consequently retains almost the same shape as in the male. Cameroons to the Congo. - ''insularis'' Auriv. is a race in which the male only differs from the type-form in having the subapical band of the forewing whitish yellow above and white beneath; in the female the subapical band of the forewing is pure white, but the hindmarginal spot of the forewing and the hindwing are coloured and marked quite as in the male; the base of the hindwing beneath is light yellow instead of red- yellow. Island of Fernando Po. - ''melina'' Thur. is only known in the male; the light markings of the upper surface are dull ochre-yellow and the hindmarginal spot of the forewing is differently shaped, as its proximal side runs quite parallel with the costal margin and if produced would intersect the apex of the wing; in addition the distal side of the hindmarginal spot is bordered by two straight lines and forms a distinct angle. Nyassaland. - ''kivuana'' Jord. the markings of the forewing nearly as large as in the type-form, but the subapical band light cream-coloured and the hindmarginal spot light tan-yellow; the hindwing before the middle with a narrow yellowish white transverse band, not sharply defined distally; all the markings beneath yellowish white. -female : all the markings pure white; the subapical band of the fore wing narrower than in the type- form; the transverse band of the hindwing also narrower and more sharply defined. Island of Kwidgwi in Lake Kivu. - ''paragea'' Sm. (59c) is distinguished by having the markings very narrow and also in the male white; the subapical band of the forewing is only 1-2mm. in breadth and often almost broken up into spots; the hindmarginal spot is about 3 mm. in breadth and sometimes indistinct or obsolete at the hindmargin; the transverse band of the hindwing is 5mm. in breadth, sharply (or not sharply -- ''angustifasciata'' Grunb.) defined and not or scarcely extending beyond the apex of the cell. Uganda and Sesse Islands. - ''homochroa'' Jord. (59 e). Both sexes similarly coloured and marked and agreeing with the male of the type-form. The orange-yellow colour of the upper surface is somewhat lighter than in the type-form and the dark marginal band on the upperside of the hindwing is much narrower and terminates at the middle of the hindmargin. Abyssinia.


Subspecies

*''Bematistes epaea epaea'' (Senegal, Guinea-Bissau, Liberia, Ivory Coast, Ghana, Togo, Nigeria, Cameroon, Congo, Angola, western Democratic Republic of the Congo, north-western Tanzania, Zambia) *''Bematistes epaea angustifasciata'' d'Abrera, 1980 (central and southern Uganda, western Kenya) *''Bematistes epaea bicolorata'' (Le Doux, 1937) (south-eastern Democratic Republic of the Congo) *''Bematistes epaea epitellus'' Staudinger, 1896 (coast of Kenya, coast of Tanzania) *''Bematistes epaea homochroa'' (Rothschild & Jordan, 1905) (southern Ethiopia) *''Bematistes epaea insulana'' Ackery, 1995 (Bioko) *''Bematistes epaea kivuana'' (Jordan, 1910) (Democratic Republic of the Congo: east to Kivu) *''Bematistes epaea lutosa'' (Suffert, 1904) (eastern Democratic Republic of the Congo, Uganda: west to the Bwamba Valley) *''Bematistes epaea melina'' (Thurau, 1903) (south-western Tanzania, Malawi: north to the shores of Lake Malawi) *''Bematistes epaea paragea'' (Grose-Smith, 1900) (Sudan, western Uganda, western Kenya, north-western Tanzania)


Biology

The habitat consists of forests and dense Guinea savanna. Both sexes are
mimicked In evolutionary biology, mimicry is an evolved resemblance between an organism and another object, often an organism of another species. Mimicry may evolve between different species, or between individuals of the same species. In the simples ...
by '' Elymniopsis bammakoo'', ''
Pseudacraea eurytus ''Pseudacraea eurytus'', the false wanderer, is a butterfly of the family Nymphalidae. It is found in Africa. Description The numerous forms of this group dolomena'' and ''Pseudacraea rubrobasalis">rubrobasalis''may be known at once by the l ...
'' and '' Mimacraea darwinia'', while the female is also the model for the female of '' Papilio cynorta''. '' Acraea jodutta'' is probably a co-mimic. The larvae feed on '' Lindaeckeria schweinfurthii'', '' Lindaeckeria dentata'' and '' Adenia cisampelloides''.


Taxonomy

See Pierre & Bernaud, 2014 Pierre & Bernau, 2014 Classification et Liste Synonymique des Taxons du Genre ''Acraea'
pdf
/ref>


References


External links


''Die Gross-Schmetterlinge der Erde'' 13: Die Afrikanischen Tagfalter. Plate XIII 58
''a''
''Die Gross-Schmetterlinge der Erde'' 13: Die Afrikanischen Tagfalter. Plate XIII 59
''c'' ''paragea''
''Die Gross-Schmetterlinge der Erde'' 13: Die Afrikanischen Tagfalter. Plate XIII 59
''e'' ''homochroa''
Images representing ''Bematistes epaea''
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Images representing ''Bematistes epaea melina''
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Images representing ''Bematistes epaea paragea''
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Images representing ''Acraea paragea''
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Images representing ''Acraea paragea kivuana''
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at Pteron *Jacques Pierre et Dominique Bernaud, 2012 Révision des ''Acraea'' (''Acraea'') du groupe "epaea" (Lepidoptera, Nymphalidae) ''Bulletin de la Société entomologique de France'' Année 2012 117(1) pp. 7–3
online
Butterflies described in 1779 Acraeini Butterflies of Africa Taxa named by Pieter Cramer {{Heliconiinae-stub