Beagle 2 Evolution
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The Beagle is a small
breed A breed is a specific group of breedable domestic animals having homogeneous appearance (phenotype), homogeneous behavior, and/or other characteristics that distinguish it from other organisms of the same species. In literature, there exist seve ...
of
scent hound Canton_of_Bern.html" ;"title="Franz Rudolf Frisching in the uniform of an officer of the Canton of Bern">Bernese Huntsmen Corps with his Berner Laufhund, painted by Jean Preudhomme in 1785 Scent hounds (or scenthounds) are a Dog type, type of h ...
, similar in appearance to the much larger
foxhound A foxhound is a type of large hunting hound bred for strong hunting instincts, a keen sense of smell, and their barking, energy, drive, and speed. In fox hunting, the foxhound's namesake, packs of foxhounds track quarry, followed—usually on ho ...
. The beagle was developed primarily for
hunting Hunting is the Human activity, human practice of seeking, pursuing, capturing, and killing wildlife or feral animals. The most common reasons for humans to hunt are to obtain the animal's body for meat and useful animal products (fur/hide (sk ...
rabbit or
hare Hares and jackrabbits are mammals belonging to the genus ''Lepus''. They are herbivores and live Solitary animal, solitarily or in pairs. They nest in slight depressions called forms, and their young are precociality, able to fend for themselves ...
, known as
beagling Beagling is mainly the hunting of hares and rabbits by beagles using their strong sense of smell. A beagle pack (10 or more hounds) is usually followed on foot, but in a few cases mounted. Beagling is often enjoyed by 'retired' fox hunters who ...
. Possessing a great sense of smell and superior tracking instincts, the beagle is the primary breed used as a
detection dog A detection dog or sniffer dog is a dog that is trained to use its senses to detect substances such as explosives, illegal drugs, wildlife scat, semen, currency, blood, and contraband electronics such as illicit mobile phones. The sense most use ...
for prohibited agricultural imports and foodstuffs in
quarantine A quarantine is a restriction on the movement of people, animals, and goods which is intended to prevent the spread of disease or pests. It is often used in connection to disease and illness, preventing the movement of those who may have bee ...
around the world. The beagle is a popular
pet A pet, or companion animal, is an animal kept primarily for a person's company or entertainment rather than as a working animal, livestock, or a laboratory animal. Popular pets are often considered to have attractive/ cute appearances, inte ...
due to its size and amiable temperament. The modern breed was developed in Great Britain around the 1830s from several breeds, including the Talbot Hound, the
North Country Beagle The North Country Beagle, Northern Hound or Northern Beagle was a breed of dog that existed in Britain probably until early in the 19th century. The exact date of its extinction is not known; it is likely that it was gradually interbred with othe ...
, the
Southern Hound The Southern Hound was a breed of dog that existed in Britain probably until sometime in the 19th century, now extinct. The exact date of its extinction is not known; it is likely that it was gradually interbred with other breeds until the genuine ...
, and possibly the Harrier. Beagles have been depicted in popular culture since
Elizabethan The Elizabethan era is the epoch in the Tudor period of the history of England during the reign of Queen Elizabeth I (1558–1603). Historians often depict it as the golden age in English history. The Roman symbol of Britannia (a female per ...
times in literature and paintings and more recently in film, television, and comic books.


History

The origin of the Beagle is uncertain. In the 11th century,
William the Conqueror William the Conqueror (Bates ''William the Conqueror'' p. 33– 9 September 1087), sometimes called William the Bastard, was the first Norman king of England (as William I), reigning from 1066 until his death. A descendant of Rollo, he was D ...
brought the St. Hubert Hound and the Talbot hound to Britain. In Britain, both of these strains were then crossed with
Greyhound The English Greyhound, or simply the Greyhound, is a dog breed, breed of dog, a sighthound which has been bred for coursing, greyhound racing and hunting. Some are kept as show dogs or pets. Greyhounds are defined as a tall, muscular, smooth-c ...
s to give them speed and stamina for deer hunting. Beagles are similar to the Harrier and the extinct
Southern Hound The Southern Hound was a breed of dog that existed in Britain probably until sometime in the 19th century, now extinct. The exact date of its extinction is not known; it is likely that it was gradually interbred with other breeds until the genuine ...
, though they are smaller and slower. From the
Middle Ages In the history of Europe, the Middle Ages or medieval period lasted approximately from the 5th to the late 15th centuries, similarly to the post-classical period of global history. It began with the fall of the Western Roman Empire and ...
, ''Beagle'' was used as a generic description for the smaller hounds, though these dogs differed considerably from the modern breed. Miniature breeds of Beagle-type dogs were known from the times of
Edward II Edward II (25 April 1284 – 21 September 1327), also known as Edward of Caernarfon or Caernarvon, was King of England from 1307 until he was deposed in January 1327. The fourth son of Edward I, Edward became the heir to the throne follo ...
and Henry VII, who both had packs of Glove Beagles, so named since they were small enough to fit on a glove, and
Queen Elizabeth I Elizabeth I (7 September 153324 March 1603) was Queen of England and Ireland from 17 November 1558 until her death in 1603. She was the last and longest reigning monarch of the House of Tudor. Her eventful reign, and its effect on history ...
kept a breed known as a Pocket Beagle, which stood at the shoulder. Small enough to fit in a "pocket" or saddlebag, they rode along on the hunt. The larger hounds would run the prey to ground, then the hunters would release the small dogs to continue the chase through underbrush. Elizabeth I referred to the dogs as her ''singing Beagles'' and often entertained guests at her royal table by letting her Pocket Beagles cavort amid their plates and cups. 19th-century sources refer to these breeds interchangeably and it is possible that the two names refer to the same small variety. In George Jesse's ''Researches into the History of the British Dog'' from 1866, the early 17th-century poet and writer
Gervase Markham Gervase (or Jervis) Markham (ca. 1568 – 3 February 1637) was an English poet and writer. He was best known for his work ''The English Huswife, The English Huswife, Containing the Inward and Outward Virtues Which Ought to Be in a Complete Woma ...
is quoted referring to the beagle as small enough to sit on a man's hand and to the: By the 18th century, two breeds had been developed for hunting hare and rabbit: the
Southern Hound The Southern Hound was a breed of dog that existed in Britain probably until sometime in the 19th century, now extinct. The exact date of its extinction is not known; it is likely that it was gradually interbred with other breeds until the genuine ...
and the
North Country Beagle The North Country Beagle, Northern Hound or Northern Beagle was a breed of dog that existed in Britain probably until early in the 19th century. The exact date of its extinction is not known; it is likely that it was gradually interbred with othe ...
(or Northern Hound). The Southern Hound, a tall, heavy dog with a square head, and long, soft ears, was common from south of the
River Trent The Trent is the third Longest rivers of the United Kingdom, longest river in the United Kingdom. Its Source (river or stream), source is in Staffordshire, on the southern edge of Biddulph Moor. It flows through and drains the North Midlands ...
and probably closely related to the Talbot Hound. Though slow, it had stamina and an excellent scenting ability. The North Country Beagle was bred chiefly in
Yorkshire Yorkshire ( ) is an area of Northern England which was History of Yorkshire, historically a county. Despite no longer being used for administration, Yorkshire retains a strong regional identity. The county was named after its county town, the ...
and was common in the northern counties. It was smaller than the Southern Hound, less heavy-set, and with a more pointed muzzle. It was faster than its southern counterpart but its scenting abilities were less well-developed. Standards for the Pocket Beagle were drawn up as late as 1901; these genetic lines are now extinct, although modern breeders have attempted to recreate the variety.


Development of the modern breed

Reverend Phillip Honeywood established a Beagle pack in
Essex Essex ( ) is a Ceremonial counties of England, ceremonial county in the East of England, and one of the home counties. It is bordered by Cambridgeshire and Suffolk to the north, the North Sea to the east, Kent across the Thames Estuary to the ...
in the 1830s; it is believed that this pack formed the basis for the modern breed. Although details of the pack's lineage are not recorded, it is thought that North Country Beagles and Southern Hounds were strongly represented;
William Youatt William Youatt (1776 – 9 January 1847) was an English Veterinarian, veterinary surgeon and animal welfare writer. Life Youatt was the son of a non-conformist minister. He was educated for the Nonconformist (Protestantism), nonconformist ministr ...
suspected that Harriers formed a good majority of the Beagle's bloodline, but the origin of the Harrier is itself obscure. Honeywood's Beagles were small, standing at about at the shoulder, and pure white according to John Mills (writing in ''The Sportsman's Library'' in 1845).
Prince Albert Prince Albert most commonly refers to: *Prince Albert of Saxe-Coburg and Gotha (1819–1861), consort of Queen Victoria *Albert II, Prince of Monaco (born 1958), present head of state of Monaco Prince Albert may also refer to: Royalty * Alb ...
and Lord Winterton also had Beagle packs around this time, and royal favour no doubt led to some revival of interest in the breed, but Honeywood's pack was regarded as the finest of the three. Although credited with the development of the modern breed, Honeywood concentrated on producing dogs for hunting and it was left to Thomas Johnson to refine the breeding to produce dogs that were both attractive and capable hunters. Two strains were developed: the rough-coated and smooth-coated varieties. The rough-coated beagle survived until the beginning of the 20th century, and there were even records of one making an appearance at a dog show as late as 1969, but this variety is now extinct, having probably been absorbed into the standard beagle bloodline. In the 1840s, a standard Beagle type was beginning to develop. The distinction between the North Country Beagle and Southern Hound had been lost, but there was still a large variation in size, character, and reliability among the emerging packs. In 1856, "Stonehenge" (the pseudonym of
John Henry Walsh John Henry Walsh FRCS (21 October 1810 – 12 February 1888) was an English sports writer born in Hackney, London who wrote under the pseudonym "Stonehenge." Walsh was educated in private schools and became a fellow of the Royal College of Surg ...
), writing in the ''Manual of British Rural Sports'', was still dividing beagles into four varieties: the medium Beagle; the dwarf or lapdog beagle; the fox beagle (a smaller, slower version of the Foxhound); and the rough-coated or terrier beagle, which he classified as a cross between any of the other varieties and one of the Scottish
terrier Terrier () is a Dog type, type of dog originally bred to hunt vermin. A terrier is a dog of any one of many Dog breed, breeds or landraces of the terrier Dog type, type, which are typically small, wiry, Gameness, game, and fearless. There are fi ...
breeds. Stonehenge also gives the start of a standard description: By 1887, the threat of extinction was on the wane: there were 18 Beagle packs in England. The Beagle Club was formed in 1890 and the first standard drawn up at the same time. The following year the Association of Masters of Harriers and Beagles was formed. Both organisations aimed to further the best interests of the breed, and both were keen to produce a standard type of Beagle. By 1902, the number of packs had risen to 44.


Export

Beagles were in the United States by the 1840s at the latest, but the first dogs were imported strictly for hunting and were of variable quality. Since Honeywood had only started breeding in the 1830s, it is unlikely these dogs were representative of the modern breed, and the description of them as looking like straight-legged
Dachshund The dachshund ( or ; German: 'badger dog'), also known as the wiener dog or sausage dog, badger dog, doxen and doxie, is a short-legged, long-bodied, hound-type dog breed. The dog may be smooth-haired, wire-haired, or long-haired, with varie ...
s with weak heads has little resemblance to the standard. Serious attempts at establishing a quality bloodline began in the early 1870s when General
Richard Rowett Richard Rowett (November 17, 1830 – July 13, 1887) was a leading political figure of nineteenth-century Illinois, a famous animal breeder and an officer in the Union Army during the American Civil War. He was born in East Looe, Cornwall in 1830 ...
from
Illinois Illinois ( ) is a U.S. state, state in the Midwestern United States, Midwestern United States. It borders on Lake Michigan to its northeast, the Mississippi River to its west, and the Wabash River, Wabash and Ohio River, Ohio rivers to its ...
imported some dogs from England and began breeding. Rowett's Beagles are believed to have formed the models for the first American standard, drawn up by Rowett, L. H. Twadell, and
Norman Ellmore Norman or Normans may refer to: Ethnic and cultural identity * The Normans, a people partly descended from Norse Vikings who settled in the territory of Normandy in France in the 9th and 10th centuries ** People or things connected with the Norma ...
in 1887.


Popularity

On its formation, the Association of Masters of Harriers and Beagles took over the running of a regular show at
Peterborough Peterborough ( ) is a City status in the United Kingdom, cathedral city in the City of Peterborough district in the Ceremonial counties of England, ceremonial county of Cambridgeshire, England. The city is north of London, on the River Nene. A ...
that had started in 1889, and the Beagle Club in the UK held its first show in 1896. The regular showing of the breed led to the development of a uniform type, and the Beagle continued to prove a success up until the outbreak of
World War I World War I or the First World War (28 July 1914 – 11 November 1918), also known as the Great War, was a World war, global conflict between two coalitions: the Allies of World War I, Allies (or Entente) and the Central Powers. Fighting to ...
when all shows were suspended. After the war, the breed was again struggling for survival in the UK: the last of the Pocket Beagles was probably lost during this time, and registrations fell to an all-time low. A few breeders (notably Reynalton Kennels) managed to revive interest in the dog and by
World War II World War II or the Second World War (1 September 1939 – 2 September 1945) was a World war, global conflict between two coalitions: the Allies of World War II, Allies and the Axis powers. World War II by country, Nearly all of the wo ...
, the breed was once again doing well. Registrations dropped again after the end of the war but almost immediately recovered. As purebred dogs, Beagles have always been more popular in the United States and Canada than in their native country England. The National Beagle Club of America was formed in 1888 and by 1901 a Beagle had won a Best in Show title. As in the UK, activity during World War I was minimal, but the breed showed a much stronger revival in the U.S. when hostilities ceased. In 1928 it won a number of prizes at the
Westminster Kennel Club The Westminster Kennel Club Dog Show is an all-breed conformation show, held annually in the New York metropolitan area. The Westminster Kennel Club Dog Show is one of a handful of Bench show, benched shows in the United States. Dogs are require ...
's show and by 1939 a Beagle – Champion Meadowlark Draughtsman – had captured the title of top-winning American-bred dog for the year. On 12 February 2008, a Beagle,
K-Run's Park Me In First Ch. K-Run's Park Me In First, also known as Uno (May 5, 2005 – September 20, 2018) was a beagle from Belleville, Illinois, who won Best in Show in the 2008 Westminster Kennel Club dog show. He was the first beagle to claim the top prize at ...
(Uno), won the Best In Show category at the Westminster Kennel Club show for the first time in the competition's history. In North America they have been consistently in the top-ten most-popular breeds for over 30 years. From 1953 to 1959 the beagle was ranked No. 1 on the list of the
American Kennel Club The American Kennel Club (AKC) is a 501(c)(4) nonprofit breed registry, registry of purebred dog pedigree (animal), pedigrees in the United States. In addition to maintaining its pedigree registry, this kennel club also promotes and sanctions eve ...
's registered breeds; in 2005 and 2006 it ranked 5th out of the 155 breeds registered. In the UK they are not quite so popular, placing 28th and 30th in the rankings of registrations with the Kennel Club in 2005 and 2006 respectively. In the United States the beagle ranked 4th most popular breed in 2012 and 2013, behind the
Labrador Retriever The Labrador Retriever or simply Labrador or Lab is a British list of dog breeds, breed of water dog retriever gun dog. It was developed in the United Kingdom from St. John's water dogs imported from the Newfoundland Colony, colony of Newfoun ...
,
German Shepherd The German Shepherd, also known in Britain as an Alsatian, is a German Dog breed, breed of working dog of medium to large size. The breed was developed by Max von Stephanitz using various Old German herding dogs, traditional German herding dog ...
, and Golden Retriever.


Name

According to the ''
Oxford English Dictionary The ''Oxford English Dictionary'' (''OED'') is the principal historical dictionary of the English language, published by Oxford University Press (OUP), a University of Oxford publishing house. The dictionary, which published its first editio ...
'', the first mention of ''beagle'' in English literature dates from c. 1475 in ''
The Squire of Low Degree ''The Squire of Low Degree'', also known as ''The Squyr of Lowe Degre'', ''The Sqyr of Lowe Degre'' or ''The Sqyr of Lowe Degree'', is an anonymous late Middle English or early Modern English verse romance. There is little doubt that it was inte ...
''. The origin of the word ''beagle'' is uncertain, although it has been suggested that the word derives from the French ''begueule'' which means "gate throat". It is not known why the black and tan
Kerry Beagle The Kerry Beagle () is the only extant scent hound breed native to Ireland. Description Appearance Despite its name, the Kerry Beagle is a small hound. is an Irish word for small, given to the dog because it is smaller than other hounds; ...
, present in Ireland since
Celt The Celts ( , see Names of the Celts#Pronunciation, pronunciation for different usages) or Celtic peoples ( ) were a collection of Indo-European languages, Indo-European peoples. "The Celts, an ancient Indo-European people, reached the apoge ...
ic times, has the ''beagle'' description, since at it is significantly taller than the modern day Beagle, and in earlier times was even larger. Some writers suggest that the Beagle's scenting ability may have come from cross-breeding earlier strains with the Kerry Beagle. Originally used for hunting stags, it is today used for hare and
drag hunting Drag hunting or draghunting is a form of equestrian sport where mounted riders hunt the trail of an artificially laid scent with hounds. Description Drag hunting is an equestrian sport where a field of mounted riders chase a pack of hounds who fo ...
.


Appearance

The general appearance of the Beagle resembles a miniature
Foxhound A foxhound is a type of large hunting hound bred for strong hunting instincts, a keen sense of smell, and their barking, energy, drive, and speed. In fox hunting, the foxhound's namesake, packs of foxhounds track quarry, followed—usually on ho ...
, but the head is broader and the muzzle shorter, the expression completely different and the legs shorter in proportion to the body. They are generally between high at the
withers Withers are the ridge between the shoulder blades of an animal, typically a quadruped. In many species, this ridge is the tallest point of the body. In horses and dogs, it is the standard place to measure the animal's height. In contrast, catt ...
and weigh between , with females being slightly smaller than males on average. They have a smooth, somewhat domed
skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...
with a medium-length, square-cut muzzle, and a black (or occasionally liver) gumdrop nose. The jaw is strong, and the teeth scissor together with the upper teeth fitting perfectly over the lower teeth. Both sets align square to the jaw. The eyes are large, hazel or brown, with a mild, hound-like pleading look. The large ears are long, soft, and low-set, turning towards the cheeks slightly and rounded at the tips. Beagles have a strong, medium-length neck (which is long enough for them to easily bend to the ground to pick up a scent), with little folding in the skin but some evidence of a
dewlap A dewlap is a longitudinal flap of skin or similar flesh that hangs beneath the lower jaw or neck of many vertebrates. More loosely, it can be various similar structures in the neck area, such as those caused by a double chin or the submandibula ...
; a broad chest narrowing to a tapered
abdomen The abdomen (colloquially called the gut, belly, tummy, midriff, tucky, or stomach) is the front part of the torso between the thorax (chest) and pelvis in humans and in other vertebrates. The area occupied by the abdomen is called the abdominal ...
and waist and a long, slightly curved tail (known as the "stern") tipped with white. The white tip, known as the flag, was bred for selectively, as the tail remains easily seen when the dog's head is down following a scent. The tail does not curl over the back, but is held upright when the dog is active. The Beagle has a muscular body and a medium-length, smooth, hard coat. The front legs are straight and carried under the body while the rear legs are muscular and well bent at the stifles. The tricoloured Beaglewhite with large black areas and light brown shadingis the most common. Tricoloured Beagles occur in a number of shades, from the "Classic Tri" with a jet black saddle (also known as "Blackback"), to the "Dark Tri" (where faint brown markings are intermingled with more prominent black markings), to the "Faded Tri" (where faint black markings are intermingled with more prominent brown markings). Some tricoloured dogs have a broken pattern, sometimes referred to as ''pied''. These dogs have mostly white coats with patches of black and brown hair. Tricolour Beagles are almost always born black and white. The white areas are typically set by eight weeks, but the black areas may fade to brown as the puppy matures. (The brown may take between one and two years to fully develop.) Some Beagles gradually change colour during their lives, and may lose their black markings entirely. Two-colour varieties always have a white base colour with areas of the second colour. Tan and white is the most common two-colour variety, but there is a wide range of other colours including lemon, a very light tan; red, a reddish, almost orange, brown; and liver, a darker brown, and black. Liver is not common and is not permitted in some standards; it tends to occur with yellow eyes. Ticked or mottled varieties may be either white or black with different coloured flecks (''ticking''), such as the blue-mottled or bluetick Beagle, which has spots that appear to be a midnight-blue colour, similar to the colouring of the
Bluetick Coonhound The Bluetick Coonhound is a breed of coonhound originating in the United States. The Bluetick Coonhound is known for its friendly personality, cold nose, and deep bawl mouth. It is most commonly used as a raccoon hunting dog, but is also kept as ...
. Some tricolour Beagles also have ticking of various colours in their white areas. Saddle-patterned Beagles that express ancient domino (eA) are called hare-pied. Domino restricts eumelanin production leading to a smaller and faded saddle. Dark hairs in the area of the saddle have a light-coloured hair base. The nose pigment may be lightened down the middle.


Sense of smell

Alongside the
Bloodhound The bloodhound is a large scent hound, originally bred for hunting deer, wild boar, rabbits, and since the Middle Ages, for tracking people. Believed to be descended from hounds once kept at the Abbey of Saint-Hubert, Belgium, in French it is ...
and
Basset Hound The Basset Hound is a short-legged breed of scent hound. The Basset Hound was developed in Great Britain from several now-extinct strains of France, French basset breeds. It was bred primarily for hunting rabbit and hare on foot, moving slowly en ...
, the Beagle has one of the best developed senses of smell of any dog. In the 1950s, John Paul Scott and John Fuller began a 13-year study of canine behaviour. As part of this research, they tested the scenting abilities of various breeds by putting a mouse in a one-acre field and timing how long it took the dogs to find it. The Beagles found it in less than a minute while
Fox Terrier Fox Terriers are two different breeds of the terrier dog type: the Smooth Fox Terrier and the Wire Fox Terrier. Both of these breeds originated in the 19th century from a handful of dogs who are descended from earlier varieties of United Kingdo ...
s took 15 minutes and
Scottish Terrier The Scottish Terrier (; also known as the Aberdeen Terrier), popularly called the Scottie, is a dog breed, breed of dog. Initially one of the highland breeds of terrier that were grouped under the name of ''Skye Terrier'', it is one of five br ...
s failed to find it at all. The long ears and large lips of the Beagle probably assist in trapping the scents close to the nose.


Variations


Breed varieties

The
American Kennel Club The American Kennel Club (AKC) is a 501(c)(4) nonprofit breed registry, registry of purebred dog pedigree (animal), pedigrees in the United States. In addition to maintaining its pedigree registry, this kennel club also promotes and sanctions eve ...
recognises two separate varieties of Beagle: the 13-inch for hounds less than , and the 15-inch for those between . The
Canadian Kennel Club The Canadian Kennel Club (or CKC; ), founded in 1888 and chartered under the Animal Purebred Act, is one of the national kennel clubs of Canada. It maintains breed registries services for those purebred dogs approved for its control by Agricultur ...
recognises a single type, with a height not exceeding . The
Kennel Club (UK) The Royal Kennel Club (KC) is the official kennel club of the United Kingdom. It is the oldest recognised kennel club in the world. Its role is to oversee various canine activities including dog shows, dog agility and working trials. It also op ...
and FCI affiliated clubs recognise a single type, with a height of between . English and American varieties are sometimes mentioned. However, there is no official recognition from any Kennel Club for this distinction. Beagles fitting the American Kennel Club standard – which disallows animals over – are smaller on average than those fitting the Kennel Club standard which allows heights up to . Pocket Beagles are sometimes advertised for sale but while the UK Kennel Club originally specified a standard for the pocket Beagle in 1901, the variety is now not recognised by any Kennel Club. A strain known as Patch Hounds was developed by Willet Randall and his family from 1896 specifically for their rabbit hunting ability. They trace their bloodline back to Field Champion Patch, but do not necessarily have a patchwork marking.


Crossbreeds

In the 1850s,
John Henry Walsh John Henry Walsh FRCS (21 October 1810 – 12 February 1888) was an English sports writer born in Hackney, London who wrote under the pseudonym "Stonehenge." Walsh was educated in private schools and became a fellow of the Royal College of Surg ...
(Stonehenge) recommended a cross between a Beagle and a
Scottish Terrier The Scottish Terrier (; also known as the Aberdeen Terrier), popularly called the Scottie, is a dog breed, breed of dog. Initially one of the highland breeds of terrier that were grouped under the name of ''Skye Terrier'', it is one of five br ...
as a retriever. He found the
crossbreed A crossbreed is an organism with purebred parents of two different breeds, varieties, or populations. A domestic animal of unknown ancestry, where the breed status of only one parent or grandparent is known, may also be called a crossbreed though ...
to be a good worker, silent and obedient, but it had the drawback that it was small and could barely carry a hare. More recently, the trend has been for " designer dogs" and one of the most popular has been the Beagle/
Pug The Pug is a breed of dog with the physically distinctive features of a wrinkly, short-muzzled face, and curled tail. An ancient breed, with roots dating back to 400 B.C., they have a fine, glossy coat that comes in a variety of colors, most ...
cross known as a
puggle A puggle is a dog crossbreed, dog crossbred from a Pug and a Beagle. The breed originated in the 1990s in the United States. History The puggle originated from accidental matings. The first planned cross between a pug and a beagle took place ...
. Some puppies of this cross are less excitable than a Beagle and with a lower exercise requirement, similar to the Pug parent; but many are highly excitable and require vigorous exercise.


Temperament

The Beagle has an even temper and gentle disposition. Described in several breed standards as "merry", they are amiable and typically neither aggressive nor timid, although this depends on the individual. They enjoy company, and although they may initially be standoffish with strangers, they are easily won over. They make poor
guard dogs A guard dog or watchdog is a dog used to watch for and guard people or property against unwanted human or animal intruders. A dog trained to attack intruders is known as an attack dog. History Dogs have been used as guardians since ancient ...
for this reason, although their tendency to bark or
howl Howl most often refers to: * Howling, an animal vocalization in many canine species * "Howl" (poem), a 1956 poem by Allen Ginsberg Howl or The Howl may also refer to: Film * '' The Howl'', a 1970 Italian film * ''Howl'' (2010 film), a 2010 Am ...
when confronted with the unfamiliar makes them good watch dogs. In a 1985 study conducted by Ben and Lynette Hart, the Beagle was given the highest excitability rating, along with the
Yorkshire Terrier The Yorkshire Terrier, also known as a Yorkie, is a British breed of toy dog of terrier type. It is among the smallest of the terriers and indeed of all dog breeds, with a weight of no more than . It originated in the nineteenth century in the ...
,
Cairn Terrier The Cairn Terrier is a terrier breed originating in the Scottish Highlands and recognised as one of Scotland's earliest working dogs. The name "Cairn Terrier" was a compromise suggestion when the breed was brought to official shows in the Unite ...
,
Miniature Schnauzer The Miniature Schnauzer is a dog breed, breed of small dog of the Schnauzer type that originated in Germany in the mid-to-late 19th century. Miniature Schnauzers may have been developed from the smallest specimens of the Standard Schnauzer, or c ...
,
West Highland White Terrier The West Highland White Terrier (commonly shortened to Westie) is a dog breed, breed of dog from Scotland with a distinctive white harsh coat and somewhat soft white undercoat. It is a medium-sized terrier, though with longer legs than other ...
, and
Fox Terrier Fox Terriers are two different breeds of the terrier dog type: the Smooth Fox Terrier and the Wire Fox Terrier. Both of these breeds originated in the 19th century from a handful of dogs who are descended from earlier varieties of United Kingdo ...
. They are ranked 72nd in
Stanley Coren Stanley Coren (born 1942) is a psychology professor, neuropsychological researcher and writer on the intelligence, mental abilities and history of dogs. He works in research and instructs in psychology at the University of British Columbia in V ...
's ''
The Intelligence of Dogs ''The Intelligence of Dogs: A Guide to the Thoughts, Emotions, and Inner Lives of Our Canine Companions'' is a 1994 book on dog intelligence by Stanley Coren, a professor of canine psychology at the University of British Columbia. The book expla ...
'', as Coren places them among the group with the lowest degree of working/obedience intelligence. Coren's scale, however, does not assess understanding, independence, or creativity. Beagles are excellent with children and this is one of the reasons they have become popular family pets. Beagles are pack animals; they are prone to
separation anxiety Separation anxiety disorder (SAD) is an anxiety disorder in which an individual experiences excessive anxiety regarding separation from home and/or from people to whom the individual has a strong emotional attachment (e.g., a parent, caregive ...
, a condition which causes them to destroy things when left unattended. Not all Beagles will howl, but most will bark when confronted with strange situations, and some will bay (also referred to as "speaking", "giving tongue", or "opening") when they catch the scent of potential quarry. They also generally get along well with cats and other dogs. They are not too demanding with regard to exercise; their inbred stamina means they do not easily tire when exercised, but they also do not need to be worked to exhaustion before they will rest. Regular exercise helps ward off the weight gain to which the breed is prone.


Health

A 2024 study in the UK found a life expectancy of 12.5 years compared to an average of 12.7 for purebreeds and 12 for
crossbreeds A crossbreed is an organism with purebred parents of two different breeds, varieties, or populations. A domestic animal of unknown ancestry, where the breed status of only one parent or grandparent is known, may also be called a crossbreed though ...
. A 2024 Italian study found a life expectancy of 11 years for the breed compared to 10 years overall. Beagles may be prone to
epilepsy Epilepsy is a group of Non-communicable disease, non-communicable Neurological disorder, neurological disorders characterized by a tendency for recurrent, unprovoked Seizure, seizures. A seizure is a sudden burst of abnormal electrical activit ...
, but this can often be controlled with medication.
Hypothyroidism Hypothyroidism is an endocrine disease in which the thyroid gland does not produce enough thyroid hormones. It can cause a number of symptoms, such as cold intolerance, poor ability to tolerate cold, fatigue, extreme fatigue, muscle aches, co ...
and a number of types of
dwarfism Dwarfism is a condition of people and animals marked by unusually small size or short stature. In humans, it is sometimes defined as an adult height of less than , regardless of sex; the average adult height among people with dwarfism is . '' ...
occur in Beagles. Two conditions in particular are unique to the breed: "Funny Puppy", in which the puppy is slow to develop and eventually develops weak legs, a crooked back and although normally healthy, is prone to a range of illnesses; and Musladin-Lueke syndrome (MLS) in which the eyes are slanted and the outer toes are underdeveloped but otherwise development is as normal.
Hip dysplasia Hip dysplasia is an abnormality of the hip joint where the socket portion does not fully cover the ball portion, resulting in an increased risk for joint dislocation. Hip dysplasia may occur at birth or develop in early life. Regardless, it doe ...
, common in Harriers and in some larger breeds, is rarely considered a problem in Beagles. Beagles are considered a chondrodystrophic breed, meaning that they are prone to types of disk diseases. In rare cases, Beagles may develop immune mediated polygenic arthritis (where the immune system attacks the joints) even at a young age. The symptoms can sometimes be relieved by
steroid A steroid is an organic compound with four fused compound, fused rings (designated A, B, C, and D) arranged in a specific molecular configuration. Steroids have two principal biological functions: as important components of cell membranes t ...
treatments. Another rare disease in the breed is neonatal cerebellar cortical degeneration. Affected puppies are slow, have lower co-ordination, fall more often, and do not have a normal gait. It has an estimated carrier rate of 5% and affected rate of 0.1%. A genetic test is available. Their long floppy ears can mean that the inner ear does not receive a substantial air flow or that moist air becomes trapped, and this can lead to ear infections. Beagles may also be affected by a range of eye problems; two common ophthalmic conditions in beagles are
glaucoma Glaucoma is a group of eye diseases that can lead to damage of the optic nerve. The optic nerve transmits visual information from the eye to the brain. Glaucoma may cause vision loss if left untreated. It has been called the "silent thief of ...
and
corneal dystrophy Corneal dystrophy is a group of rare hereditary disorders characterised by bilateral abnormal deposition of substances in the transparent front part of the eye called the cornea. Signs and symptoms Corneal dystrophy may not significantly affect ...
. "
Cherry eye Cherry eye is a disorder of the nictitating membrane (NM), also called the third eyelid, present in the eyes of dogs and cats.Gelatt, K. N. (2000). ''Essentials of Veterinary Ophthalmology''. Baltimore: Lippincott Williams & Wilkins. Cherry eye i ...
", a prolapse of the gland of the
third eyelid The nictitating membrane (from Latin ''wikt:nicto, nictare'', to blink) is a transparent or translucent third eyelid present in some animals that can be drawn across the eye from the Medial (anatomy), medial canthus to protect and moisten it while ...
, and
distichiasis A distichia is an eyelash that arises from an abnormal part of the eyelid. This abnormality, attributed to a genetic mutation, is known to affect dogs and humans. Distichiae usually exit from the duct of the meibomian gland at the eyelid margin. ...
, a condition in which eyelashes grow into the eye causing irritation, sometimes exist; both these conditions can be corrected with surgery. They can suffer from several types of retinal atrophy. Failure of the nasolacrimal drainage system can cause dry eye or leakage of tears onto the face. The cause of primary open angle glaucoma in the Beagle is an
autosomal recessive In genetics, dominance is the phenomenon of one variant (allele) of a gene on a chromosome masking or overriding the Phenotype, effect of a different variant of the same gene on Homologous chromosome, the other copy of the chromosome. The firs ...
mutation in the
ADAMTS10 A disintegrin and metalloproteinase with thrombospondin motifs 10 is an enzyme that in humans is encoded by the ''ADAMTS10'' gene. This gene belongs to the ADAMTS (a disintegrin and metalloproteinase domain with thrombospondin type-1 motifs) fami ...
gene. As field dogs they are prone to minor injuries such as cuts and sprains, and, if inactive, obesity is a common problem as they will eat whenever food is available and rely on their owners to regulate their weight. When working or running free they are also likely to pick up parasites such as
flea Flea, the common name for the order (biology), order Siphonaptera, includes 2,500 species of small flightless insects that live as external parasites of mammals and birds. Fleas live by hematophagy, ingesting the blood of their hosts. Adult f ...
s,
tick Ticks are parasitic arachnids of the order Ixodida. They are part of the mite superorder Parasitiformes. Adult ticks are approximately 3 to 5 mm in length depending on age, sex, and species, but can become larger when engorged. Ticks a ...
s,
harvest mite Trombiculidae (), commonly referred to in North America as chiggers and in Britain as harvest mites, but also known as berry bugs, bush-mites, red bugs or scrub-itch mites, are a family of mites. Chiggers are often confused with Tunga penetrans ...
s, and
tapeworm Eucestoda, commonly referred to as tapeworms, is the larger of the two subclasses of flatworms in the class Cestoda (the other subclass being Cestodaria). Larvae have six posterior hooks on the scolex (head), in contrast to the ten-hooked Ce ...
s, and irritants such as grass seeds can become trapped in their eyes, soft ears, or paws. Beagles may exhibit a behaviour known as
reverse sneezing Reverse sneezing, also known as inspiratory paroxysmal respiration, is a clinical event that occurs in dogs and cats. It is possibly caused by a muscle spasm at the back of the animal's mouth, more specifically where the muscle and throat meet. ...
, in which they sound as if they are choking or gasping for breath, but are actually drawing air in through the mouth and nose. The exact cause of this behaviour is not known, but it can be a common occurrence and is not harmful to the dog. Canine juvenile polyarteritis syndrome, also known as juvenile polyarteritis syndrome of beagle dogs is a multisystemic necrotising vasculitis of the small arteries. The condition is characterised by a reoccurrent acute fever and cervicalgia over a period of 3-7 days. The Beagle is predisposed with most reports concerning the breed — whence the name 'Beagle pain syndrome'.


Hunting

Beagles were developed primarily for hunting rabbit or hare, an activity known as
beagling Beagling is mainly the hunting of hares and rabbits by beagles using their strong sense of smell. A beagle pack (10 or more hounds) is usually followed on foot, but in a few cases mounted. Beagling is often enjoyed by 'retired' fox hunters who ...
. They were seen as ideal hunting companions for the elderly who could follow on horseback without exerting themselves, for young hunters who could keep up with them on ponies, and for the poorer hunters who could not afford to maintain a stable of good hunting horses. Before the advent of the fashion for foxhunting in the 19th century, hunting was an all day event where the enjoyment was derived from the chase rather than the kill. In this setting, the tiny Beagle was well matched to the hare, as unlike Harriers they would not quickly finish the hunt, but because of their excellent scent-tracking skills and stamina they were almost guaranteed to eventually catch the hare. The Beagle packs would run closely together ("so close that they might be covered with a sheet") which was useful in a long hunt, as it prevented stray dogs from obscuring the trail. In thick undergrowth they were also preferred to
spaniel A spaniel is a Dog type, type of gun dog. Spaniels were especially bred to flush game out of denser brush. By the late 17th century, spaniels had been specialized into water and land breeds. The extinct English Water Spaniel was used to retriev ...
s when hunting
pheasant Pheasants ( ) are birds of several genera within the family Phasianidae in the order Galliformes. Although they can be found all over the world in introduced (and captive) populations, the pheasant genera's native range is restricted to Eura ...
. With the fashion for faster hunts, the beagle fell out of favour for chasing hare, but was still employed for rabbit hunting. In ''Anecdotes of Dogs'' (1846),
Edward Jesse Edward Jesse (14 January 1780 – 28 March 1868) was an English writer on natural history. As a surveyor for the Office of Works, he was responsible for much of the restoration and presentation of Hampton Court Palace after it was opened to ...
says: In the United States they appear to have been employed chiefly for hunting rabbits from the earliest imports. Hunting hare with Beagles became popular again in Britain in the mid-19th century and continued until it was made illegal in Scotland by the
Protection of Wild Mammals (Scotland) Act 2002 The Protection of Wild Mammals (Scotland) Act was an Act of the Scottish Parliament passed in February 2002, making Scotland the first part of the United Kingdom to ban traditional fox hunting and hare coursing. It was repealed in 2023. Passage ...
, and in England and Wales by the
Hunting Act 2004 The Hunting Act 2004 (c. 37) is an Act of the Parliament of the United Kingdom which bans the hunting of most wild mammals (notably foxes, deer, hares and mink) with dogs in England and Wales, subject to some strictly limited exemptions; the ...
. Under this legislation, Beagles may still pursue rabbits with the landowner's permission.
Drag hunting Drag hunting or draghunting is a form of equestrian sport where mounted riders hunt the trail of an artificially laid scent with hounds. Description Drag hunting is an equestrian sport where a field of mounted riders chase a pack of hounds who fo ...
is popular where hunting is no longer permitted or for those owners who do not wish to participate in hunting a live animal, but still wish to exercise their dog's innate skills. The traditional foot pack consists of up to 40 Beagles, marshaled by a Huntsman who directs the pack and who is assisted by a variable number of whippers-in whose job is to return straying hounds to the pack. The Master of the Hunt is in overall day-to-day charge of the pack, and may or may not take on the role of Huntsman on the day of the hunt. As hunting with Beagles was seen as ideal for young people, many of the British
public schools Public school may refer to: *Public school (government-funded), a no-fee school, publicly funded and operated by the government *Public school (United Kingdom), certain elite fee-charging private schools in England and Wales *Great Public Schools, ...
traditionally maintained Beagle packs. Protests were lodged against Eton's use of Beagles for hunting as early as 1902 but the pack is still in existence today. In 2001, the Wye College beagle pack was taken by the
Animal Liberation Front The Animal Liberation Front (ALF) is a Far-left politics, far-left international, Leaderless resistance, leaderless, decentralized movement that emerged in Britain in the 1970s, evolving from the Bands of Mercy. It operates without a formal lead ...
. School and university packs are still maintained by Eton,
Marlborough Marlborough or the Marlborough may refer to: Places Australia * Marlborough, Queensland * Principality of Marlborough, a short-lived micronation in 1993 * Marlborough Highway, Tasmania; Malborough was an historic name for the place at the sou ...
,
Radley Radley is a village and civil parish about northeast of the centre of Abingdon, Oxfordshire. The parish includes the hamlet of Lower Radley on the River Thames. It was part of Berkshire until the 1974 boundary changes transferred it to Oxfor ...
, the
Royal Agricultural University The Royal Agricultural University (RAU), formerly the Royal Agricultural College, is a public university in Cirencester, Gloucestershire, England. Established in 1845, it was the first agricultural college in the English-speaking world. ...
, and
Christ Church, Oxford Christ Church (, the temple or house, ''wikt:aedes, ædes'', of Christ, and thus sometimes known as "The House") is a Colleges of the University of Oxford, constituent college of the University of Oxford in England. Founded in 1546 by Henry V ...
. In addition to organised beagling, Beagles have been used for hunting or flushing to guns (often in pairs) a wide range of game including snowshoe hare, cottontail rabbits, game birds, roe deer, red deer, bobcat, coyote, wild boar, and foxes, and have even been recorded as being used to hunt stoat. In most of these cases, the beagle is employed as a gun dog, flushing game for hunter's guns.


Detection

Beagles are used as detection dogs in the Beagle Brigade of the United States Department of Agriculture. These dogs are used to detect food items in luggage being taken into the United States. After trialling several breeds, Beagles were chosen because they are relatively small and unintimidating for people who are uncomfortable around dogs, easy to care for, intelligent and work well for rewards. They are also used for this purpose in a number of other countries including by the Ministry of Agriculture and Forestry (New Zealand), Ministry of Agriculture and Forestry in New Zealand, the Australian Quarantine and Inspection Service, and in Canada, Japan, and the People's Republic of China. Larger breeds are generally used for detection of explosives as this often involves climbing over luggage and on large conveyor belts, work for which the smaller Beagle is not suited.


Use in animal testing

Beagles are one of the dog breeds most often used in animal testing, due to their size, temperament, and historical utilisation. In the United States, as many as 65,000 Beagles are used every year for medical, cosmetic, beauty, and other chemical tests. Most are bred specifically for this purpose, by companies such as Inotiv, Envigo, and live inside controlled laboratory facilities. Beagles are used in a range of research procedures: fundamental biological research, applied human medicine, applied veterinary medicine, and protection of man, animals, or the environment. Of the 8,018 dogs used in testing in the UK in 2004, 7,799 were Beagles (97.3%). In the UK, the Animals (Scientific Procedures) Act 1986 gave special status to primates, equids, cats and dogs and in 2005 the Animal Procedures Committee (set up by the act) ruled that testing on mice was preferable, even though a greater number of individual animals were involved. In 2005, beagles were involved in less than 0.3% of the total experiments on animals in the UK, but of the 7,670 experiments performed on dogs 7406 involved Beagles (96.6%). In the UK, companies breeding animals for research must be licensed under the Animals (Scientific Procedures) Act.


Bans and activism against Beagle testing

Testing of cosmetic products on animals is banned in the member states of the European Community, although France protested the ban and has made efforts to have it lifted. It is permitted in the United States but is not mandatory if safety can be ascertained by other methods, and the test species is not specified by the Food and Drug Administration (FDA). When testing toxicity of food additives, food contaminants, and some drugs and chemicals the FDA uses Beagles and miniature pigs as surrogates for direct human testing. The Beagle Freedom Project has successfully advocated for Beagles to be adopted or released from labs. Minnesota was the first state to enact a Beagle Freedom adoption law in 2014, mandating that dogs and cats are allowed to be adopted once they have completed research testing. Anti-Animal testing, vivisection groups have reported on abuse of animals inside testing facilities. In 1997 footage secretly filmed by a freelance journalist inside Huntingdon Life Sciences in the UK showed staff punching and screaming at Beagles. Consort Kennels, a UK-based breeder of Beagles for testing, closed down in 1997 after pressure from animal rights groups. There are various examples of activists utilising Freedom of Information Act (United States), Freedom of Information Act (FOIA) requests to obtain information about taxpayer funding of animal testing. The White Coat Waste Project reported that the National Institute of Allergy and Infectious Diseases funded experiments in which 28 Beagles were infected by disease-causing parasites and that dogs taking part in the experiments were "vocalizing in pain." In response, People for the Ethical Treatment of Animals, People for the Ethical Treatment of Animals (PETA) called on all members of the National Institute of Health to resign and stated that there is a "need to find a new NIH director to replace the outgoing Francis Collins who will shut down research that violates the dignity of nonhuman animals."


Animal welfare research

A 2023 research article report lower levels of the stress hormone cortisol in beagles kept in larger areas with toys. A 2024 article researches the use of environmental enrichment in lab beagles by adding toys to their cages.


Other roles

Although bred for hunting, Beagles are versatile and are nowadays employed for various other roles in detection, therapy, and as family pets. Beagles are used as sniffer dogs for termite detection in Australia, and have been mentioned as possible candidates for drug and explosive detection. Because of their gentle nature and unimposing build, they are also frequently used in Animal-assisted therapy, pet therapy, visiting the sick and elderly in hospital. In June 2006, a trained Beagle assistance dog was credited with saving the life of its owner after using her owner's mobile phone to dial an emergency number. In the aftermath of the 2010 Haiti earthquake, a Beagle search and rescue dog with a Colombian rescue squad was credited with locating the owner of the Hôtel Montana, who was subsequently rescued after spending 100 hours buried in the rubble. Beagles were hired by New York City to help with bedbug detection, although some have expressed doubts about the role of such dogs in this type of detection.


In popular culture

*Anthropomorphic animal, Anthropomorphic Beagles appeared in comic strips and animated cartoons since the 1950s with the ''Peanuts'' character Snoopy, who was billed as "the world's most famous Beagle". The clay animation, ''Wallace & Gromit'', also features a beagle – the titular ''Gromit'' – in its cast. *Former US President Lyndon Baines Johnson had several Beagles, and caused an outcry when he picked up one of them by its ears during an official greeting on the White House lawn. *The ship on which Charles Darwin made Second voyage of HMS Beagle, the voyage which provided much of the inspiration for ''On the Origin of Species'' was named HMS Beagle, HMS ''Beagle'' after the breed, and, in turn, lent its name to the ill-fated British Mars, Martian lander ''Beagle 2''. *An American bred 15 inch male Beagle with the registered name of Ch
K-Run's Park Me In First Ch. K-Run's Park Me In First, also known as Uno (May 5, 2005 – September 20, 2018) was a beagle from Belleville, Illinois, who won Best in Show in the 2008 Westminster Kennel Club dog show. He was the first beagle to claim the top prize at ...
and the pet name of "Uno" won the 2008 Westminster Kennel Club Dog Show. *A Canadian bred 15 inch female Beagle with the registered name of Gr Ch Tashtins Lookin For Trouble and the pet name of "Miss P" won the 2015 Westminster Kennel Club Dog Show. *Beagle is also featured as an animal actor in Inspector Gadget films named Brian.


Notable Beagles

* K-Run's Park Me In First, Uno, who in 2008 became the first Beagle to win the Westminster Kennel Club Dog Show * Tashtins Lookin For Trouble, Miss P, winner of the 2015 Westminster Kennel Club Dog Show


References

Informational notes a. In this article "Beagle" (with a capital B) is used to distinguish the modern breed from other beagle-type dogs. c. The Harts posed the following question to a panel of 96 experts, half of which were veterinary surgeons and the other half dog obedience trial judges:
d. The specific references in each of the author's works are as follows:
Shakespeare: "''Sir Toby Belch'': She's a beagle, true-bred, and one that adores me: what o' that?" ''Twelfth Night'' (c. 1600) Act II Scene III
Webster: "''Mistress Tenterhook: You are a sweet beagle" ''Westward Ho (play), Westward Ho'' (1607) Act III Scene IV:2
Dryden: "The rest in shape a beagle's whelp throughout, With broader forehead and a sharper snout" ''The Cock and the Fox'', and again: "About her feet were little beagles seen" in ''Palamon and Arcite'' both from ''Fables, Ancient and Modern'' (1700)
Tickell: "Here let me trace beneath the purpled morn, The deep-mouth'd beagle, and the sprightly horn" ''To a Lady before Marriage'' (published posthumously in 1749)
Fielding: "'What the devil would you have me do?' cries the Squire, turning to Blifil, 'I can no more turn her, than a beagle can turn an old hare.'" ''The History of Tom Jones, a Foundling'' (1749) Chapter 7.
Cowper: "For persevering chase and headlong leaps, True beagle as the staunchest hound he keeps" ''The Progress of Error'' (1782)
Pope: "Thus on a roe the well-breath'd beagle flies, And rends his hide fresh-bleeding with the dart" ''The Iliad of Homer'' (1715–20) Book XV:697–8 Citations Bibliography * * * * * * * * * * * * * * * *


External links

{{Authority control Companion dogs Dog breeds originating in England FCI breeds Hunting with hounds Scent hounds