Baurusuchus Salgadoensis
   HOME

TheInfoList



OR:

''Baurusuchus'' is an
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of baurusuchid
mesoeucrocodylia Mesoeucrocodylia is the clade that includes Eusuchia and crocodyliforms formerly placed in the paraphyletic group Mesosuchia. The group appeared during the Early Jurassic, and continues to the present day. Diagnosis It was long known that Me ...
n, which lived in
Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
from 90 to 83.5 million years ago, in the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
period. It was a terrestrial predator, estimated to reach up to in weight. ''Baurusuchus'' lived during the
Turonian The Turonian is, in the International Commission on Stratigraphy, ICS' geologic timescale, the second age (geology), age in the Late Cretaceous epoch (geology), Epoch, or a stage (stratigraphy), stage in the Upper Cretaceous series (stratigraphy), ...
to
Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya ( million years ago) and 83.6 ± 0.7 m ...
stages of the Late Cretaceous Period, in
Adamantina Formation The Adamantina Formation is a geological Formation (geology), formation in the Bauru Basin of western São Paulo (state), São Paulo state in southeastern Brazil. Its strata date back to the Late Cretaceous epoch of the Cretaceous, Cretaceous Per ...
,
Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
. It gets its name from the Brazilian Bauru Group ("Bauru crocodile"). It was related to the earlier-named '' Cynodontosuchus rothi'', which was smaller, with weaker dentition. The three species are ''B. pachecoi'', named after Eng Joviano Pacheco, its discoverer, ''B. salgadoensis'' (named after General Salgado County in São Paulo, Brazil) and ''B. albertoi'' (named after Alberto Barbosa de Carvalho, Brazilian paleontologist). The latter species is disputed (see phylogeny section). Its relatives include the similarly sized '' Stratiotosuchus'' from the
Adamantina Formation The Adamantina Formation is a geological Formation (geology), formation in the Bauru Basin of western São Paulo (state), São Paulo state in southeastern Brazil. Its strata date back to the Late Cretaceous epoch of the Cretaceous, Cretaceous Per ...
, and ''
Pabweshi ''Pabwehshi'' (meaning "Pab ormationbeast Urdu.html" ;"title="wehshi" in Urdu">wehshi" in Urdu) is an extinct genus of mesoeucrocodylian. It is holotype, based on Geological Survey of Pakistan, GSP-UM 2000, a partial snout and correspondi ...
'', from the
Pakistan Pakistan, officially the Islamic Republic of Pakistan, is a country in South Asia. It is the List of countries and dependencies by population, fifth-most populous country, with a population of over 241.5 million, having the Islam by country# ...
i Pab Formation.


Paleoecology

''B. salgadoensis'' is seen as a terrestrial predator, living in a hot and
arid climate The desert climate or arid climate (in the Köppen climate classification ''BWh'' and ''BWk'') is a dry climate sub-type in which there is a severe excess of evaporation over precipitation. The typically bald, rocky, or sandy surfaces in desert ...
. The position of the external nares was unsuited for an amphibious lifestyle like in modern crocodilians and the snout and teeth are laterally compressed like in
theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
s. Both of this supports the terrestrial hypothesis. The hot environment hypothesis is based on the lifestyle of modern crocodilians and the
stratigraphy Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks. Stratigraphy has three related subfields: lithost ...
of ''Baurusuchus''. ''B. salgadoensis'' was found in fine massive sandstones which are interpreted as a floodplain area in a hot and arid climate. ''Baurusuchus'' was likely able to dig holes for finding water in dry seasons or, like modern
alligator An alligator, or colloquially gator, is a large reptile in the genus ''Alligator'' of the Family (biology), family Alligatoridae in the Order (biology), order Crocodilia. The two Extant taxon, extant species are the American alligator (''A. mis ...
s do, for thermoregulation. The occurrence of very complete skeletons in correlated stratigraphic levels supports this. Such a strategy would have made it less water-bound than most modern crocodiles, allowing it to live in more
continental climate Continental climates often have a significant annual variation in temperature (warm to hot summers and cold winters). They tend to occur in central and eastern parts of the three northern-tier continents (North America, Europe, and Asia), typi ...
. The strongly bent
pterygoid Pterygoid, from the Greek for 'winglike', may refer to: * Pterygoid bone, a bone of the palate of many vertebrates * Pterygoid processes of the sphenoid bone ** Lateral pterygoid plate ** Medial pterygoid plate * Lateral pterygoid muscle * Medial ...
s suggest a powerful bite and that ''Baurusuchus'' could close its jaw very quickly. The skull and tooth
morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
indicates that the biting strategies of ''Baurusuchus'' were similar to a
Komodo dragon The Komodo dragon (''Varanus komodoensis''), also known as the Komodo monitor, is a large reptile of the monitor lizard family Varanidae that is endemic to the Indonesian islands of Komodo (island), Komodo, Rinca, Flores, Gili Dasami, and Gili ...
which include ambushing the prey, biting it and pulling back the serrated, blade-like teeth. ''Baurusuchus'' likely played an important role in its ecosystem, competing with the
abelisaurid Abelisauridae (meaning "Abel's lizards") is a family (or clade) of ceratosaurian theropod dinosaurs. Abelisaurids thrived during the Cretaceous period, on the ancient southern supercontinent of Gondwana, and today their fossil remains are foun ...
s for food.


Classification

''Baurusuchus'' is the type genus of the family
Baurusuchidae Baurusuchidae is a Gondwanan family of mesoeucrocodylians that lived during the Late Cretaceous. It is a group of terrestrial hypercarnivorous crocodilians from South America (Argentina and Brazil) and possibly Pakistan. Baurusuchidae has been, ...
, a family consisting of crocodilians with elongated and laterally compressed skulls. Other members of that family from the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
of South America include '' Stratiotosuchus'' and '' Cynodontosuchus'', but baurusuchids are also known from the Cretaceous of
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
(
Pakistan Pakistan, officially the Islamic Republic of Pakistan, is a country in South Asia. It is the List of countries and dependencies by population, fifth-most populous country, with a population of over 241.5 million, having the Islam by country# ...
) and the
Tertiary Tertiary (from Latin, meaning 'third' or 'of the third degree/order..') may refer to: * Tertiary period, an obsolete geologic period spanning from 66 to 2.6 million years ago * Tertiary (chemistry), a term describing bonding patterns in organic ch ...
of
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
. A study in 2011 erected a new subfamily called Baurusuchinae. Seven diagnostic features for the group were described which include the moderate size and the broad frontals. The paper referred only ''Stratiotosuchus maxhechti'' and ''Baurusuchus'' to the subfamily, making ''Stratiotosuchus'' ''Baurusuchus closest relative so far. However, a study in the year 2014 referred a new species called '' Aplestosuchus sordidus'' to the subfamily, but supported a closer relationship of ''Baurususchus'' with ''Stratiotosuchus'' than with it. The species ''B. albertoi'' is an exception. The paper does not support its affiliation to ''Baurusuchus'' and views it as a close relative of ''Aplestosuchus''. This is the cladogram they presented:


Sources

* ''In the Shadow of the Dinosaurs: Early Mesozoic Tetrapods'' by Nicholas C. Fraser and
Hans-Dieter Sues Hans-Dieter Sues (born 1956) is a German-born American palaeontologist who is a Senior Research Geologist and Curator of Vertebrate Paleontology at the National Museum of Natural History of the Smithsonian Institution in Washington, DC. Career ...
* ''The Osteology of the Reptiles'' by
Alfred Sherwood Romer Alfred Sherwood Romer (December 28, 1894 – November 5, 1973) was an American paleontologist and biologist and a specialist in vertebrate evolution. Biography Alfred Romer was born in White Plains, New York, the son of Harry Houston Romer an ...


References

{{Taxonbar, from=Q137262 Late Cretaceous crocodylomorphs of South America Terrestrial crocodylomorphs Baurusuchidae Adamantina Formation Prehistoric pseudosuchian genera Fossil taxa described in 1945 Taxa named by Llewellyn Ivor Price