Bat Falcon
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The bat falcon (''Falco rufigularis'') is a
bird of prey Birds of prey or predatory birds, also known as (although not the same as) raptors, are hypercarnivorous bird species that actively predation, hunt and feed on other vertebrates (mainly mammals, reptiles and smaller birds). In addition to speed ...
in the family
Falconidae The falcons and caracaras are around 65 species of Diurnality, diurnal birds of prey that make up the family Falconidae (representing all extant species in the order (biology), order Falconiformes). The family likely originated in South America d ...
, the falcons and caracaras. It is found in
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
,
Central America Central America is a subregion of North America. Its political boundaries are defined as bordering Mexico to the north, Colombia to the southeast, the Caribbean to the east, and the Pacific Ocean to the southwest. Central America is usually ...
,
Trinidad Trinidad is the larger, more populous island of the Republic of Trinidad and Tobago, the country. The island lies off the northeastern coast of Venezuela and sits on the continental shelf of South America. It is the southernmost island in ...
, and every mainland
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
n country except
Chile Chile, officially the Republic of Chile, is a country in western South America. It is the southernmost country in the world and the closest to Antarctica, stretching along a narrow strip of land between the Andes, Andes Mountains and the Paci ...
and
Uruguay Uruguay, officially the Oriental Republic of Uruguay, is a country in South America. It shares borders with Argentina to its west and southwest and Brazil to its north and northeast, while bordering the Río de la Plata to the south and the A ...
.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. 30 January 2023. Species Lists of Birds for South American Countries and Territories. https://www.museum.lsu.edu/~Remsen/SACCCountryLists.htm retrieved January 30, 2023


Taxonomy and systematics

The bat falcon was long known as ''Falco albigularis''; the names ''Falco fusco-coerulescens'' or ''Falco fuscocaerulescens'', long used for the
aplomado falcon The aplomado falcon (''Falco femoralis'') is a medium-sized falcon of the Americas. The species' largest continuous range is in South America, but not in the deep interior Amazon Basin. It was long known as ''Falco fusco-coerulescens'' or ''Falco ...
(''F. femoralis''), are now believed to refer to the present species. The bat falcon has these three subspecies: *''F. r. petoensis'' Chubb, 1918 *''F. r. rufigularis'' Daudin, 1800 *''F. r. ophryophanes'' ( Salvadori, 1895) Some authors maintain that ''F. r. petoensis'' and ''F. r. ophryophanes'' are not subspecies but clinal variations in plumage. Others add a fourth subspecies ''F. r. petrophilus'' that is usually included in ''petoensis''. Yet others assign only two subspecies, the
nominate Nomination is part of the process of selecting a candidate for either election to a public office, or the bestowing of an honor or award. A collection of nominees narrowed from the full list of candidates is a short list. Political office In th ...
''F. r. rufigularis'' and ''F. r. petrophilus''.Bierregaard, R. O. and G. M. Kirwan (2020). Bat Falcon (''Falco rufigularis''), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.batfal1.01 retrieved February 15, 2023 The bat falcon and the
orange-breasted falcon The orange-breasted falcon (''Falco deiroleucus'') is a Near Threatened bird of prey in the family Falconidae, the falcons and caracaras. It is found in southern Mexico, Belize, Guatemala, Panama, and either definitely or probably in every Sout ...
(''F. deiroleucus'') share plumage and vocal characteristics and may be
sister species In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 30 January 2023. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved January 30, 2023 Those two appear to be closely related to the
aplomado falcon The aplomado falcon (''Falco femoralis'') is a medium-sized falcon of the Americas. The species' largest continuous range is in South America, but not in the deep interior Amazon Basin. It was long known as ''Falco fusco-coerulescens'' or ''Falco ...
(''F. femoralis'').


Description

The bat falcon is long. Males weigh and have a wingspan of . Females weigh and have a wingspan of . They have long wings and a longish tail with a square tip. The sexes have similar plumage. Adults have blue-black head and upperparts with grayish edges on the feathers from the upper back to the uppertail
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are small feathers behind t ...
. Their throat, upper breast, and sides of the neck are white to buff, sometimes with some cinnamon; the rest of their breast is black with fine white bars. Their belly, thighs, and undertail coverts are chestnut-rufous. Their tail is blackish with thin white or grayish bars and a white or buff tip. The underside of their wings is black with fine white bars. Their
cere The beak, bill, or Rostrum (anatomy), rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for pecking, wikt:grasp#Verb, grasping, and holding (in wikt:probe ...
and bare skin around the eye are bright yellow, their iris black-brown, and their legs and feet orange-yellow. Juveniles are duller and browner than adults, with a buffier throat, a tawny tinge to the breast's barring, and black bars or spots on the undertail coverts. The three subspecies are similar, differing mainly in the tone of their plumage colors.


Distribution and habitat

The subspecies of the bat falcon are found thus: *''F. r. petoensis'', from northern Mexico south through all of Central America and west of the Andes of Colombia, Ecuador, and extreme northwestern Peru *''F. r. rufigularis'', Trinidad and from eastern Colombia east through Venezuela and
the Guianas The Guianas, also spelled Guyanas or Guayanas, are a geographical region in north-eastern South America. Strictly, the term refers to the three Guianas: Guyana, Suriname, and French Guiana, formerly British, Dutch, and French Guiana respectiv ...
and south through eastern Ecuador, eastern Peru, northern and eastern Bolivia, southern Brazil, and northern Argentina *''F. r. ophryophanes'', central Brazil and adjacent eastern Bolivia, Paraguay, and northeastern Argentina A juvenile male of subspecies ''F. r. petoensis'' wandered to
Santa Ana National Wildlife Refuge Santa Ana National Wildlife Refuge is a National Wildlife Refuge situated along the banks of the Rio Grande, south of Alamo in the Lower Rio Grande Valley, in Hidalgo County, Texas. This sub-tropical sanctuary harbors one of the highest wildl ...
in Alamo, Texas, in December 2021, for the only U.S. record of the species. It stayed in the area into early March. ''F. r. rufigularis'' is resident on Trinidad and has been recorded as a
vagrant Vagrancy is the condition of wandering homelessness without regular employment or income. Vagrants usually live in poverty and support themselves by travelling while engaging in begging, scavenging, or petty theft. In Western countries, ...
on Tobago. The bat falcon inhabits tropical forest. It favors unbroken mature forest but also occurs at the forest edge, in
gallery forest A gallery forest is one formed as a corridor along rivers or wetlands, projecting into landscapes that are otherwise only sparsely treed such as savannas, grasslands, or deserts. The gallery forest maintains a more temperate microclimate above th ...
, on wooded savanna, on cleared land with some remaining trees, and even in suburban and urban areas. In elevation it mostly ranges from sea level to about with a single record in Bolivia at about .


Behavior


Movement

The bat falcon is apparently mostly sedentary. However, the records in Texas, at high elevation in Bolivia, on Tobago, and on islands off the
Yucatán Peninsula The Yucatán Peninsula ( , ; ) is a large peninsula in southeast Mexico and adjacent portions of Belize and Guatemala. The peninsula extends towards the northeast, separating the Gulf of Mexico to the north and west of the peninsula from the C ...
, Honduras, and Panama show a pattern of wandering.


Feeding

Bat falcons perch conspicuously on high, open snags, from which they launch aerial attacks on their prey. They also hunt in sustained flight. Most hunting is around dawn or dusk and often continues well past sunset. Their diet is eclectic; they hunt
bat Bats are flying mammals of the order Chiroptera (). With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most birds, flying with their very long spread-out ...
s,
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s (such as
swift Swift or SWIFT most commonly refers to: * SWIFT, an international organization facilitating transactions between banks ** SWIFT code * Swift (programming language) * Swift (bird), a family of birds It may also refer to: Organizations * SWIF ...
s,
swallow The swallows, martins, and saw-wings, or Hirundinidae are a family of passerine songbirds found around the world on all continents, including occasionally in Antarctica. Highly adapted to aerial feeding, they have a distinctive appearance. The ...
s,
hummingbird Hummingbirds are birds native to the Americas and comprise the Family (biology), biological family Trochilidae. With approximately 366 species and 113 genus, genera, they occur from Alaska to Tierra del Fuego, but most species are found in Cen ...
s,
parakeet A parakeet is any one of many small- to medium-sized species of parrot, in multiple genera, that generally has long tail feathers. Etymology and naming The name ''parakeet'' is derived from the French word ''perroquet'', which is reflect ...
s,
tanager The tanagers (singular ) comprise the bird family (biology), family Thraupidae, in the order Passeriformes. The family has a Neotropical distribution and is the second-largest family of birds. It represents about 4% of all avian species and 12 ...
s and small
water bird A water bird, alternatively waterbird or aquatic bird, is a bird that lives on or around water. In some definitions, the term ''water bird'' is especially applied to birds in freshwater ecosystems, although others make no distinction from seabi ...
s), small
rodent Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...
s,
snake Snakes are elongated limbless reptiles of the suborder Serpentes (). Cladistically squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales much like other members of the group. Many species of snakes have s ...
s,
lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...
s,
frog A frog is any member of a diverse and largely semiaquatic group of short-bodied, tailless amphibian vertebrates composing the order (biology), order Anura (coming from the Ancient Greek , literally 'without tail'). Frog species with rough ski ...
s, and large
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s such as
dragonflies A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of dragonflies are known. Most are tropical, with fewer species in temperate regions. Loss of wetland habitat threate ...
,
butterflies Butterflies are winged insects from the lepidopteran superfamily Papilionoidea, characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The oldest butterfly fossi ...
,
moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyletic with respect to butterflies (s ...
s,
grasshopper Grasshoppers are a group of insects belonging to the suborder Caelifera. They are amongst what are possibly the most ancient living groups of chewing herbivorous insects, dating back to the early Triassic around 250 million years ago. Grassh ...
s,
true bug Hemiptera (; ) is an order of insects, commonly called true bugs, comprising more than 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, assassin bugs, bed bugs, and shield bugs. They range in size from to a ...
s,
beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
s, and
hymenopteran Hymenoptera is a large order of insects, comprising the sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are parasitic. Females typi ...
s. In most areas vertebrates contribute the most to biomass consumed, varying from 66% to 96% in various studies, and birds made up between 32% and 85% of the vertebrates. Invertebrates dominate numerically, and young are fed large numbers of them. Bats generally contribute less than 14% of prey biomass but some pairs seem to specialize in them.


Breeding

The bat falcon's breeding season varies geographically. It spans from February to June in Mexico and northern Central America. It nests in February on Trinidad and probably between October and February in Argentina. Eggs have been noted in March in Venezuela, in April in Guyana, and in August near
Manaus Manaus () is the List of capitals of subdivisions of Brazil, capital and largest city of the States of Brazil, Brazilian state of Amazonas (Brazilian state), Amazonas. It is the List of largest cities in Brazil, seventh-largest city in Brazil, w ...
, Brazil. Most nests are in tree cavities, either natural or excavated by parrots, and between above the ground. Others have been noted in abandoned trogon nest cavities in arboreal termite nests, on cliffs, and on pre-Columbian ruins. The clutch size is two to four eggs. The incubation period is thought to be about 30 days, fledging occurs about 35 to 40 days after hatch, and young are dependent on the parents for at least 12 weeks after fledging. Both sexes incubate the eggs with the female doing at least 75% of the effort and in some areas all of it. Males provide food for incubating females and also most of the food for nestlings.


Vocalization

The bat falcon is vocal "in breeding season, especially near nest, in contacts with other raptors, and even when attacking prey." Its main call is a "rapid shrill screaming" ''kee-kee-kee..'' or ''kew-kew-kew..''; the male's is higher pitched than the female's. Other calls include "an even higher and thinner “tsee-tsee-tsee...”", a "low-intensity “chit” given in contact", and by the female, "a whining or high-pitched wailing call with a wavering quality".


Status

The
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
has assessed the bat falcon as being of Least Concern. It has an extremely large range and an estimated population of at least a half million mature individuals, though the latter is believed to be decreasing. No immediate threats have been identified. It is " despread and generally not uncommon in appropriate habitat, it being one of the most widely distributed New World falcons." But it "no longer breeds in several areas where forest extensively transformed to agriculture" and "pesticides
ave is a Latin word, used by the Roman Empire, Romans as a salutation (greeting), salutation and greeting, meaning 'wikt:hail, hail'. It is the singular imperative mood, imperative form of the verb , which meant 'Well-being, to be well'; thus on ...
unquestionably affected breeding success" in Mexico and Central America.


References


External links

* (for
Suriname Suriname, officially the Republic of Suriname, is a country in northern South America, also considered as part of the Caribbean and the West Indies. It is a developing country with a Human Development Index, high level of human development; i ...
) with RangeMap
Bat Falcon photo gallery
VIREO

oiseaux {{Taxonbar, from=Q844220 Falco (genus) Birds of Central America Birds of Mexico Birds of the Yucatán Peninsula Birds of South America Birds of the Amazon rainforest Birds of the Guiana Shield Birds described in 1800