''Basilotritus'' is a genus of

basilosaurid Basilosauridae is a family of extinct cetaceans. They lived during the middle to the early late Eocene and are known from all continents, including Antarctica. They were probably the first fully aquatic cetaceans.Buono M, Fordyce R.E., Marx F. ...

early whale known from the early late

Middle Eocene The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "d ...

(late

Lutetian The Lutetian is, in the geologic timescale, a stage (stratigraphy), stage or age (geology), age in the Eocene. It spans the time between . The Lutetian is preceded by the Ypresian and is followed by the Bartonian. Together with the Bartonian it ...

Bartonian The Bartonian is, in the ICS's geologic time scale, a stage or age in the middle Eocene Epoch or Series. The Bartonian Age spans the time between . It is preceded by the Lutetian and is followed by the Priabonian Age. Stratigraphic defin ...

, ) Kiev Formation of the

Ukraine Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian invas ...

, Stockletten Formation of

Germany Germany, officially the Federal Republic of Germany (FRG),, is a country in Central Europe. It is the most populous member state of the European Union. Germany lies between the Baltic and North Sea to the north and the Alps to the sou ...

Tongeren Formation The Tongeren Formation ( nl, Formatie van Tongeren; abbreviation: TO) is a geologic formation in the subsurface of the Netherlands. The formation consists of shallow marine, epicontinental and continental sediments, predominantly clay and sand from ...

of the

Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...

, Giushi Formation of

Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a transcontinental country spanning the northeast corner of Africa and southwest corner of Asia via a land bridge formed by the Sinai Peninsula. It is bordered by the Med ...

and the

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...

(

Castle Hayne Formation A castle is a type of fortified structure built during the Middle Ages predominantly by the nobility or royalty and by military orders. Scholars debate the scope of the word ''castle'', but usually consider it to be the private fortified ...

North Carolina North Carolina () is a state in the Southeastern region of the United States. The state is the 28th largest and 9th-most populous of the United States. It is bordered by Virginia to the north, the Atlantic Ocean to the east, Georgia a ...

and

Piney Point Formation The Piney Point Formation is a geologic formation in Virginia. It preserves fossils dating back to the Lutetian Stage of the Eocene Epoch of the Paleogene period. Species The following species are known from this formation. Keep in mind, this fo ...

Virginia Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the East Coast of the United States, Atlantic Coast and the Appalachian Mountains. The geography an ...

''Basilotritus uheni''

''Basilotritus uheni'' has been found in the late Middle Eocene (Bartonian) Kiev Formation at Beloskelevatoye, Ukraine, making it the oldest fossil

cetacean Cetacea (; , ) is an infraorder of aquatic mammals that includes whales, dolphins, and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively carnivorous diet. They propel them ...

from

Eastern Europe Eastern Europe is a subregion of the European continent. As a largely ambiguous term, it has a wide range of geopolitical, geographical, ethnic, cultural, and socio-economic connotations. The vast majority of the region is covered by Russia, wh ...

. Its

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...

, NMNH-P OF-2096, consists of a partial skeleton.

''Basilotritus wardi''

''Basilotritus wardi'' is known from a partial skeleton collected in the late

(~42 Ma) of North Carolina. It was originally described as a species of the

protocetid Protocetidae, the protocetids, form a diverse and heterogeneous group of extinct cetaceans known from Asia, Europe, Africa, South America, and North America. Description There were many genera, and some of these are very well known (e.g., ''R ...

Eocetus ''Eocetus'' is an extinct protocetid early whale known from the early late Eocene (Bartonian, ) Giushi Formation in Gebel Mokattam, (, paleocoordinates ) outside Cairo, Egypt. The specimen was first named by Fraas as ''Mesocetus schweinfurth ...

'', ''E. wardii''. Because this species displays a mix of protocetid and basilosaurid traits, the original generic assignment was questioned by subsequent authors.Geisler, Jonathan H; Sanders, Albert E; Luo, Zhe-Xi (July 2005). "A new protocetid whale (Cetacea, Archaeoceti) from the late middle Eocene of South Carolina". ''American Museum Novitates'' 3480: 1–68. . . This skepticism was confirmed by the discovery of ''B. uheni'', allowing for "E." ''wardi'' to be referred to ''Basilotritus''.

References

Basilosauridae Prehistoric cetacean genera Bartonian life Lutetian life Eocene mammals of Africa Fossils of Egypt Eocene mammals of Europe Paleogene Germany Fossils of Germany Fossils of Ukraine Eocene mammals of North America Fossils of the United States Fossils of North Carolina Fossil taxa described in 2013 Paleontology in Virginia {{paleo-whale-stub