''Banksia squarrosa'', commonly known as pingle, is a species of prickly shrub that is

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

to Western Australia. It has linear to narrow lance-shaped leaves with up to ten sharply-pointed teeth on each side, yellow flowers in heads of about sixty and later, up to seven oblong to egg-shaped follicles in each head.

Description

''Banksia squarrosa'' is a shrub that typically grows to a height of but does not form a

lignotuber A lignotuber is a woody swelling of the root crown possessed by some plants as a protection against destruction of the plant stem, such as by fire. Other woody plants may develop basal burls as a similar survival strategy, often as a respons ...

. It has

sessile Sessility, or sessile, may refer to: * Sessility (motility), organisms which are not able to move about * Sessility (botany), flowers or leaves that grow directly from the stem or peduncle of a plant * Sessility (medicine), tumors and polyps that ...

linear to narrow lance-shaped leaves that are long and wide with up to ten sharply-pointed, triangular teeth on each side. The flowers are yellow and are arranged in heads of between fifty and seventy with narrow triangular to linear

involucral bracts In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves in size, color, shape or texture. They also look ...

long at the base of each head. The

perianth The perianth (perigonium, perigon or perigone in monocots) is the non-reproductive part of the flower. It is a structure that forms an envelope surrounding the sexual organs, consisting of the calyx (sepals) and the corolla (petals) or tepal ...

is long and the

pistil Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl (botany), whorl of a flower; it consists ...

long and straight. Flowering occurs from June to November and the follicles are oblong to egg-shaped, long and more or less

glabrous Glabrousness () is the technical term for a lack of hair, down, setae, trichomes, or other such covering. A glabrous surface may be a natural characteristic of all or part of a plant or animal, or be due to loss because of a physical condition, ...

. Up to seven follicles form in each head.

Taxonomy and naming

This species was first collected from near

King George Sound King George Sound (Mineng ) is a sound (geography), sound on the south coast of Western Australia. Named King George the Third's Sound in 1791, it was referred to as King George's Sound from 1805. The name "King George Sound" gradually came in ...

in 1829 by William Baxter, and its description was published by

Robert Brown Robert Brown may refer to: Robert Brown (born 1965), British Director, Animator and author Entertainers and artists * Washboard Sam or Robert Brown (1910–1966), American musician and singer * Robert W. Brown (1917–2009), American printmaker ...

in '' Supplementum primum Prodromi florae Novae Hollandiae'' as ''Dryandra squarrosa'' the following year. In 1839

John Lindley John Lindley Fellow of the Royal Society, FRS (5 February 1799 – 1 November 1865) was an English botanist, gardener and orchidology, orchidologist. Early years Born in Old Catton, Catton, near Norwich, England, John Lindley was one of four c ...

described ''D. carduacea'' from specimens collected in the vicinity of the Swan River. In 1996, Alex George reduced ''D. carduacea'' to a

synonym A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...

of ''D. squarrosa''. In 1996, George described two subspecies of ''D. squarrosa'': * ''Dryandra squarrosa'' subsp. ''argillacea'' that has a perianth long with a glabrous limb; * ''Dryandra squarrosa'' subsp. ''squarrosa'' that has a perianth long with a hairy limb. In 2007

Austin Mast Austin R. Mast is a research botanist. Born in 1972, he obtained a Ph.D. from the University of Wisconsin–Madison in 2000. He is currently a professor within the Department of Biological Science at Florida State University (FSU), and has been ...

and

Kevin Thiele Kevin R. Thiele is currently an adjunct associate professor at the University of Western Australia and the director of Taxonomy Australia. He was the curator of the Western Australian Herbarium from 2006 to 2015. His research interests include ...

transferred all dryandras to the genus ''Banksia'' and renamed this species ''Banksia squarrosa'' and the two subspecies ''argillacea'' and ''squarrosa'' respectively. The names of the subspecies are accepted by the

Australian Plant Census The Australian Plant Census (APC) provides an online interface to currently accepted, published, scientific names of the vascular flora of Australia, as one of the output interfaces of the national government Integrated Biodiversity Information Sys ...

Distribution and habitat

Pingle is widely distributed in the south-west of Western Australia, occurring between Bindoon, the

Whicher Range Whicher Range, also known as Whicher Scarp, is a range in the South West region of Western Australia. The range has an average elevation of above sea level. Bounded by the Swan Coastal Plain to the west and the south, the Darling Scarp to t ...

and Albany, growing in woodland and forest. Subspecies ''argillacea'' has a limited distribution near the western side of the Whicher Range. Subspecies ''squarrosa'' occurs throughout the species' range, except near the Whicher Range.

Conservation status

This banksia is classified as "not threatened" by the Western Australian Government

Department of Parks and Wildlife The Department of Parks and Wildlife (DPaW) was the department of the Government of Western Australia responsible for managing lands described in the ''Conservation and Land Management Act 1984'' and implementing the state's conservation and en ...

, but subspecies ''argillacea'' is listed as " Threatened Flora (Declared Rare Flora — Extant)" by the

Department of Environment and Conservation (Western Australia) The Department of Environment and Conservation (DEC) was a department of the Government of Western Australia that was responsible for implementing the state's conservation and environment legislation and regulations. It was formed on 1 July 2006 ...

References

* {{Taxonbar, from=Q4856732 squarrosa Taxa named by Robert Brown (botanist, born 1773) Plants described in 1830