Banksia Nobilis
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''Banksia nobilis'', commonly known as the golden dryandra, great dryandra or kerosene bush, is a shrub of the family
Proteaceae The Proteaceae form a family (biology), family of flowering plants predominantly distributed in the Southern Hemisphere. The family comprises 83 genus, genera with about 1,660 known species. Australia and South Africa have the greatest concentr ...
which is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to
Western Australia Western Australia (WA) is the westernmost state of Australia. It is bounded by the Indian Ocean to the north and west, the Southern Ocean to the south, the Northern Territory to the north-east, and South Australia to the south-east. Western Aust ...
. It occurs on
lateritic Laterite is a soil type rich in iron and aluminium and is commonly considered to have formed in hot and wet tropical areas. Nearly all laterites are of rusty-red coloration, because of high iron oxide content. They develop by intensive and prolo ...
rises from Eneabba to Katanning in the state's Southwest Botanic Province. With large pinnatifid leaves with triangular lobes, and a golden or reddish pink inflorescence, it is a popular garden plant. It was known as ''Dryandra nobilis'' until 2007, when all ''Dryandra'' species were transferred to ''
Banksia ''Banksia'' is a genus of around 170 species of flowering plants in the family Proteaceae. These Australian wildflowers and popular garden plants are easily recognised by their characteristic flower spikes, and woody fruiting "cones" and head ...
'' by
Austin Mast Austin R. Mast is a research botanist. Born in 1972, he obtained a Ph.D. from the University of Wisconsin–Madison in 2000. He is currently a professor within the Department of Biological Science at Florida State University (FSU), and has been ...
and
Kevin Thiele Kevin R. Thiele is currently an adjunct associate professor at the University of Western Australia and the director of Taxonomy Australia. He was the curator of the Western Australian Herbarium from 2006 to 2015. His research interests include ...
. There are two subspecies, ''B. nobilis'' subsp. ''nobilis'' and ''B. nobilis'' subsp. ''fragrans''.


Description

''Banksia nobilis'' grows as a shrub up to four metres high. Its leaves are pinnatifid, with 14 to 32 triangular lobes on each side from eight to 22 centimetres long, five to 25 millimetres wide, on a
petiole Petiole may refer to: *Petiole (botany), the stalk of a leaf, attaching the blade to the stem *Petiole (insect anatomy) In entomology, petiole is the technical term for the narrow waist of some hymenopteran insects, especially ants, bees, and ...
five to 15 millimetres long. Inflorescences occur on short lateral branchlets; this species bears a great many inflorescences, often carrying an inflorescence in almost every
axil A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, fl ...
. Flowers are golden or reddish pink, with a greenish cream limb. After flowering, heads can bear up to 5 follicles each.


Taxonomy

Specimens of ''B. nobilis'' were first collected in the 1830s by James Drummond from the vicinity of the
Swan River Colony The Swan River Colony, also known as the Swan River Settlement, or just ''Swan River'', was a British colony established in 1829 on the Swan River, in Western Australia. This initial settlement place on the Swan River was soon named Perth, an ...
. The species was published under the name ''Dryandra nobilis'' by
John Lindley John Lindley Fellow of the Royal Society, FRS (5 February 1799 – 1 November 1865) was an English botanist, gardener and orchidology, orchidologist. Early years Born in Old Catton, Catton, near Norwich, England, John Lindley was one of four c ...
in his 1840 ''
A Sketch of the Vegetation of the Swan River Colony "A Sketch of the Vegetation of the Swan River Colony", also known by its standard botanical abbreviation ''Sketch Veg. Swan R.'', is an 1839 article by John Lindley on the flora of the Swan River Colony. Nearly 300 new species were published in it ...
'', where he referred to it as "a most splendid plant in the way of ''D. longifolia'' and ''tenuifolia'', with leaves from a foot to a foot and half long". Lindley did not specify his
type Type may refer to: Science and technology Computing * Typing, producing text via a keyboard, typewriter, etc. * Data type, collection of values used for computations. * File type * TYPE (DOS command), a command to display contents of a file. * ...
material, and there is no type at the
University of Cambridge The University of Cambridge is a Public university, public collegiate university, collegiate research university in Cambridge, England. Founded in 1209, the University of Cambridge is the List of oldest universities in continuous operation, wo ...
Herbarium, where most of Lindley's type specimens are lodged. However most of ''A Sketch of the Vegetation of the Swan River Colony'' is based upon the collections of Drummond, and one of Drummond's specimens has since been selected as
lectotype In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to anchor or centralizes ...
for the species. Lindley also proffered no
etymology Etymology ( ) is the study of the origin and evolution of words—including their constituent units of sound and meaning—across time. In the 21st century a subfield within linguistics, etymology has become a more rigorously scientific study. ...
for the specific epithet, but it is accepted that it comes from the
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
''nobilis'' ("noble, imposing") in reference to the purportedly noble appearance of the plant. In addition to plant specimens, Drummond also sent
seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
s of ''D. nobilis'' to England. Plants were raised from Drummond's seeds, and in 1852 one of them flowered, making ''D. nobilis'' one of only two dryandras known to have flowered in
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
from Drummond's seed. The flowering specimen was about seven years old at the time, and about four feet high. It became the basis for a
lithographed Lithography () is a planographic method of printing originally based on the immiscibility of oil and water. The printing is from a stone (lithographic limestone) or a metal plate with a smooth surface. It was invented in 1796 by the German a ...
plate by
Walter Hood Fitch Walter Hood Fitch (28 February 1817 – 14 January 1892) was a botanical illustrator, born in Glasgow, Scotland, who executed some 10,000 drawings for various publications. His work in colour lithograph, including 2700 illustrations for ''C ...
, which was featured in Volume 78 of ''
Curtis's Botanical Magazine ''The Botanical Magazine; or Flower-Garden Displayed'', is an illustrated publication which began in 1787. The longest running botanical magazine, it is widely referred to by the subsequent name ''Curtis's Botanical Magazine''. Each of the issue ...
''. By this time, however, the enthusiasm for
Proteaceae The Proteaceae form a family (biology), family of flowering plants predominantly distributed in the Southern Hemisphere. The family comprises 83 genus, genera with about 1,660 known species. Australia and South Africa have the greatest concentr ...
that prevailed among horticulturalists in the 1840s had waned, and ''D. nobilis'' would be the last dryandra to feature in ''Curtis's Botanical Magazine''. Text accompanying Fitch's plate by
William Jackson Hooker Sir William Jackson Hooker (6 July 178512 August 1865) was an English botany, botanist and botanical illustrator, who became the first director of Kew Gardens, Kew when in 1841 it was recommended to be placed under state ownership as a botan ...
suggested that
Carl Meissner Carl Daniel Friedrich Meissner (1 November 1800 – 2 May 1874) was a Swiss botanist. Biography Born in Bern, Switzerland on 1 November 1800, he was christened Meisner but later changed the spelling of his name to Meissner. For most of his 40 ...
considered their specimen to be the then unpublished species ''Dryandra runcinata'', but this was a mis-identification: ''D. runcinata'' is now considered a synonym of ''
Banksia squarrosa ''Banksia squarrosa'', commonly known as pingle, is a species of prickly shrub that is Endemism, endemic to Western Australia. It has linear to narrow lance-shaped leaves with up to ten sharply-pointed teeth on each side, yellow flowers in hea ...
'' rather than this species. The species has had a fairly uneventful taxonomic history since publication, with only three events of taxonomic interest since publication. In 1891,
Otto Kuntze Carl Ernst Otto Kuntze (23 June 1843 – 27 January 1907) was a German botanist. Biography Otto Kuntze was born in Leipzig. An apothecary in his early career, he published an essay entitled ''Pocket Fauna of Leipzig''. Between 1863 and 1866, he ...
made an unsuccessful attempt to transfer the genus ''Dryandra'' into the name ''Josephia'', in the process publishing the name ''Josephia nobilis'' for this species. That name is now considered a nomenclatural synonym of ''B. nobilis''. In 1996 Alex George published a subspecies, ''D. nobilis'' subsp. ''fragrans'' (now ''B. nobilis'' subsp. ''fragrans''), therefore also invoking the
autonym Autonym may refer to: * Autonym, the name used by a person to refer to themselves or their language; see Exonym and endonym * Autonym (botany), an automatically created infrageneric or infraspecific name See also * Nominotypical subspecies, in zo ...
''D. nobilis'' subsp. ''nobilis'' (now ''B. nobilis'' subsp. ''nobilis'') to encompass the type material. Finally, in 2007 the genus ''Dryandra'' was transferred into ''
Banksia ''Banksia'' is a genus of around 170 species of flowering plants in the family Proteaceae. These Australian wildflowers and popular garden plants are easily recognised by their characteristic flower spikes, and woody fruiting "cones" and head ...
'' by
Austin Mast Austin R. Mast is a research botanist. Born in 1972, he obtained a Ph.D. from the University of Wisconsin–Madison in 2000. He is currently a professor within the Department of Biological Science at Florida State University (FSU), and has been ...
and
Kevin Thiele Kevin R. Thiele is currently an adjunct associate professor at the University of Western Australia and the director of Taxonomy Australia. He was the curator of the Western Australian Herbarium from 2006 to 2015. His research interests include ...
, resulting in the publication of the current name for this species, ''Banksia nobilis'' (Lindl.) A.R.Mast & K.R.Thiele.


Distribution and habitat

''Banksia nobilis'' occurs on
lateritic Laterite is a soil type rich in iron and aluminium and is commonly considered to have formed in hot and wet tropical areas. Nearly all laterites are of rusty-red coloration, because of high iron oxide content. They develop by intensive and prolo ...
rises from Eneabba in the north to Katanning in the south. Much of its distribution roughly follows the boundary between the
Jarrah Forest Jarrah Forest, also known as the Southwest Australia woodlands, is an interim Australian bioregion and ecoregion located in the south west of Western Australia.
and
Avon Wheatbelt The Avon Wheatbelt is a bioregion in Western Australia. It has an area of . It is considered part of the larger Southwest Australia savanna ecoregion. Geography The Avon Wheatbelt bioregion is mostly a gently undulating landscape with low rel ...
biogeographic Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
regions, but at its northern limits it extends into the
Swan Coastal Plain The Swan Coastal Plain in Western Australia is the geographic feature which contains the Swan River as it travels west to the Indian Ocean. The coastal plain continues well beyond the boundaries of the Swan River and its tributaries, as a geol ...
and
Geraldton Sandplains Geraldton Sandplains is an Interim Biogeographic Regionalisation for Australia, interim Australian bioregion of Western Australia. It has an area of . The Geraldton Sandplains is part of the larger Southwest Australia savanna ecoregion, as asses ...
. For most of its range it grows in eucalypt woodland and tall
shrubland Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominance (ecology), dominated by shrubs, often also including grasses, herbaceous plant, herbs, and geophytes. Shrubland may either occur naturally o ...
, but at its northern extents it grows among thick
kwongan Kwongan is a plant community found in south-western Western Australia. The name is a Bibulman Aboriginal term of wide geographical use defined by Beard (1976) as Kwongan has replaced other terms applied by European botanists such as sand-h ...
.


Conservation

''Banksia nobilis'' subsp. ''nobilis'' is not currently endangered, and is currently exploited in the wild by the
cut flower industry Cut flowers are flowers and flower buds (often with some stem and leaf) that have been cut from the plant bearing it. It is removed from the plant for decorative use. Cut greens are leaves with or without stems added to the cut flowers for con ...
. ''Banksia nobilis'' subsp. ''fragrans'' has been listed as Priority Three - Poorly Known Taxa on the Western Australian Department of Environment and Conservation's
Declared Rare and Priority Flora List The Declared Rare and Priority Flora List is the system by which Western Australia's conservation flora are given a priority. Developed by the Government of Western Australia's Department of Environment and Conservation, it was used extensively ...
. An assessment of the potential
impact of climate change Effects of climate change are well documented and growing for Earth's natural environment and human societies. Changes to the climate system include an Instrumental temperature record, overall warming trend, Effects of climate change on the ...
on this species found that severe change is likely to lead to a contraction of its range by up to 80% by 2080; but under less severe change scenarios the distribution may contract slightly or even grow, depending on how effectively it can migrate into newly habitable areas.


Cultivation

''Banksia nobilis'' subsp. ''nobilis'' has long been a popular garden plant because of its prolific flowering. It prefers heavy, gravelly soil with excellent drainage, in a sunny position. It is drought tolerant, and frost tolerant down to -7 °C. Plants become straggly and untidy with age, so should be pruned when young. Propagation is by seed, as the hairy stems of this plant make it virtually impossible to strike stem cuttings, and heel cuttings have been met with limited success. Germination takes three to four weeks, and has an excellent success rate. Flowering usually begins at three to four years from seed. ''Banksia nobilis'' subsp. ''fragrans'' is new to cultivation and there is little information available for it. It probably requires a similar aspect to subspecies ''nobilis'', but would be better suited to warmer areas.


References


External links

* * * {{Taxonbar, from=Q4856654
nobilis ''Nobilis'' is a contemporary fantasy tabletop role-playing game created by Jenna K. Moran. Early editions were published under Moran's previous names, Rebecca Sean Borgstrom and R. Sean Borgstrom. The player characters are "Sovereign Powers" ...
Eudicots of Western Australia Endemic flora of Southwest Australia