Balaena

''Balaena'' is a genus of cetacean (whale) in the family Balaenidae. ''Balaena'' is considered a monotypic genus, as it has only a single extant species, the bowhead whale (''B. mysticetus''). It was named in 1758 by Linnaeus, who at the time considered all of the right whales (and the bowhead) as a single species. Historically, both the family Balaenidae and genus ''Balaena'' were known by the common name, "right whales", however ''Balaena'' are now known as bowhead whales. "The taxonomy is not in doubt.... Concerning common names, the species was once commonly known in the North Atlantic and adjacent Arctic as the Greenland Right Whale. However, the common name Bowhead Whale is now generally used for the species."

Throughout history, the family Balaenidae has been the subject of great taxonomic debate. Authorities have repeatedly recategorized the three populations of right whale plus the bowhead whale, as one, two, three or four species, either in a single genus or in two separate genera. In the early whaling days, they were all thought to be a single species, ''Balaena mysticetus''. Eventually, it was recognized that bowheads and right whales were in fact different. Later, morphological factors such as differences in the skull shape of northern and southern right whales indicated at least two species of right whale—one in the Northern Hemisphere, the other in the Southern Ocean. As recently as 1998, Dale Rice, in his comprehensive and otherwise authoritative classification, ''Marine mammals of the world: systematics and distribution'', listed just two species: ''Balaena glacialis'' (the right whales) and ''Balaena mysticetus'' (the bowheads).

A DNA study by Rosenbaum in 2000, and another study by Churchill in 2007 finally provided clear evidence to conclude that the ''three'' living right whale species do comprise a phylogenetic lineage, ''distinct from the bowhead'', and that the bowhead and the right whales are rightly classified into two separate genera. The right whales, therefore, are now officially in the genus ''Eubalaena''.

The fossil record of ''Balaena'', dating to the late Miocene, encompasses ten species known from finds in Europe, North America, and South America.The Paleobiology Database ''Balaena'' entry
accessed on 21 December 2010 ''Balena'', described by Scopoli in 1777, and ''Leiobalaena'', described by Eschricht in 1849, are junior synonyms of ''Balaena''.

Taxonomy

* ''Balaena'' Bowhead whales
** † '' Balaena affinis'' (Pliocene, UK)
** † '' Balaena montalionis'' (Piacenzian; Casina, Italy).
** ''Balaena mysticetus'', Bowhead whale (Late Pleistocene to Modern)
** † '' Balaena ricei'' (Piacenzian; Rice's Pit, Yorktown Formation, Virginia, USA).

Dubious species

* † '' Balaena arcuata'' (Miocene/Pliocene; Belgium)
* † '' Balaena forsythmajori'' (Zanclean; Case il Poggio. Italy)
* † '' Balaena larteti'' (Neogene; Landes, France)
* † '' Balaena macrocephalus'' (Langhian; Sos, France)
* † '' Balaena simpsoni'' (Miocene/Pliocene; Ancud, Chile) (possibly a member of NeobalaenidaeR. Donoso-Barros. 1975. Contribucion al conocimiento de los cetaceos vivientes y fosiles del territorio de Chile. Gayana Zoologica 36:1-127)

Species inquirenda

* † '' Balaena pampaea'' ( Calabrian; Pampeana Formation, Bahía Blanca, Argentina)
† = extinct species

References

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Balaenidae
Mammal genera with one living species
Extant Miocene first appearances
Taxa named by Carl Linnaeus
Cetacean genera