''Badamia exclamationis'', commonly known as the brown awl or narrow-winged awl,
[Markku Savela's website on Lepidoptera. Page on genu]
[TOL web page on genu]
''Badamia''
/ref> is a butterfly
Butterflies are insects in the macrolepidopteran clade Rhopalocera from the order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The group comprises ...
belonging to the family Hesperiidae
Skippers are a family of the Lepidoptera (moths and butterflies) named the Hesperiidae. Being diurnal, they are generally called butterflies. They were previously placed in a separate superfamily, Hesperioidea; however, the most recent taxonomy ...
. It is found in south
South is one of the cardinal directions or compass points. The direction is the opposite of north and is perpendicular to both east and west.
Etymology
The word ''south'' comes from Old English ''sūþ'', from earlier Proto-Germanic ''*sunþa ...
and southeast Asia
Southeast Asia, also spelled South East Asia and South-East Asia, and also known as Southeastern Asia, South-eastern Asia or SEA, is the geographical United Nations geoscheme for Asia#South-eastern Asia, south-eastern region of Asia, consistin ...
, Australia, and Oceania
Oceania (, , ) is a region, geographical region that includes Australasia, Melanesia, Micronesia, and Polynesia. Spanning the Eastern Hemisphere, Eastern and Western Hemisphere, Western hemispheres, Oceania is estimated to have a land area of ...
.
Range
The brown awl is found in Sri Lanka
Sri Lanka (, ; si, ශ්රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
, India
India, officially the Republic of India ( Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the ...
, Myanmar
Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
, Cambodia
Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailan ...
, South Yunnan
Yunnan , () is a landlocked province in the southwest of the People's Republic of China. The province spans approximately and has a population of 48.3 million (as of 2018). The capital of the province is Kunming. The province borders the ...
, Australia and Japan. This butterfly is found throughout the Indian subcontinent
The Indian subcontinent is a physiographical region in Southern Asia. It is situated on the Indian Plate, projecting southwards into the Indian Ocean from the Himalayas. Geopolitically, it includes the countries of Bangladesh, Bhutan, India ...
and in the Andaman islands
The Andaman Islands () are an archipelago in the northeastern Indian Ocean about southwest off the coasts of Myanmar's Ayeyarwady Region. Together with the Nicobar Islands to their south, the Andamans serve as a maritime boundary between th ...
. The type locality is South India
South India, also known as Dakshina Bharata or Peninsular India, consists of the peninsular southern part of India. It encompasses the States and union territories of India, Indian states of Andhra Pradesh, Karnataka, Kerala, Tamil Nadu, and T ...
.
Status
As per William Harry Evans
Brigadier William Harry Evans CSI CIE DSO (born 22 July 1876 in Shillong – died 13 November 1956, Church Whitfield ) was a lepidopterist and British Army officer who served in India. He documented the butterfly fauna of India, Burma and C ...
(1932), the butterfly is common in India and rare in the Andaman islands
The Andaman Islands () are an archipelago in the northeastern Indian Ocean about southwest off the coasts of Myanmar's Ayeyarwady Region. Together with the Nicobar Islands to their south, the Andamans serve as a maritime boundary between th ...
. Mark Alexander Wynter-Blyth
Mark Alexander Wynter-Blyth (15 August 1906 – 16 April 1963 Leysin, Switzerland) was an English schoolteacher and amateur naturalist who wrote one of the first field guides to the butterflies of the Indian region. He was also involved in censuses ...
(1957) records it as "Not Rare" and "Locally Common". Krushnamegh Kunte (2000) reports it as common in deciduous forests during the monsoon months and the evergreen forests in the following months.
Habits
A forest butterfly, the brown awl favours openings and edges of deciduous and evergreen forests while its caterpillars are to be found in moist deciduous and semi-evergreen forests. It flies about either late or early in the morning in the shade of the jungles. It can be sometimes seen in bright sunlight visiting flowers, such as ''Glycosmis
''Glycosmis'' is a genus of flowering plants in the citrus family, Rutaceae and tribe Clauseneae.[''Glyc ...]
'', ''Buddleia
''Buddleja'' (; ''Buddleia''; also historically given as ''Buddlea'') is a genus comprising over 140 species of flowering plants endemic to Asia, Africa, and the Americas. The generic name bestowed by Linnaeus posthumously honoured the Revere ...
'', ''Chromolaena
''Chromolaena'' is a genus of about 165 species of perennials and shrubs in the family Asteraceae. The name is derived from the Greek words (), meaning "color", and () or () meaning "cloak". It refers to the colored phyllaries of some specie ...
'' and ''Lantana
''Lantana'' () is a genus of about 150 species of perennial flowering plants in the verbena family, Verbenaceae. They are native to tropical regions of the Americas and Africa but exist as an introduced species in numerous areas, especially ...
'', but is very wary and energetic at such times, moving jerkily and rapidly between flowers or across inflorescence
An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed ...
s. It can also be seen mud-puddling
Mud-puddling, or simply puddling, is a behaviour most conspicuous in butterflies, but occurs in other animals as well, mainly insects; they seek out nutrients in certain moist substances such as rotting plant matter, mud and carrion and they suc ...
or at bird droppings.
During a population explosion, like those of the common banded awl (''Hasora chromus
''Hasora chromus'', the common banded awl,Markku Savela's website on Lepidoptera – page on genu is a butterfly belonging to the family Hesperiidae which is found in the Indian subcontinent, Southeast Asia and Australia.
Range
The common ba ...
''), the caterpillars of the brown awl may strip away all their food supply forcing the butterflies to migrate to other places where a fresh supply of host plants is available and even to other habitats such as shrubs, grasslands and gardens.
The brown awl flies as low as 6 feet over the bushes or as high as 60 to 75 feet in the canopy. The adults feed at lower levels on flowers of shrubs and small trees, but ascend to higher reaches of the vegetation to lay eggs or to bask, which it does very occasionally, holding its wings flat with the forewings covering the hindwings thus giving an arrowhead
An arrowhead or point is the usually sharpened and hardened tip of an arrow, which contributes a majority of the projectile mass and is responsible for impacting and penetrating a target, as well as to fulfill some special purposes such as sign ...
effect. The flight of the butterfly is fast and bounding with an audible wing beat.
When inactive, it rests on the undersides of leaves in shady forest spots, with the head pointing downwards. If disturbed it will generally buzz around energetically before returning to the same spot to rest.
Description
The brown awl is a non-descript brown butterfly, darker above and lighter below. The sexes are alike, except for three to four semi-transparent spots on the forewing which cannot be differentiated in the field. The skipper has a light-brown abdomen with black bands across it. The dry-season form is usually smaller, paler, and may not have the forewing spots.
This skipper is unmistakable because of its long and narrow wings. It has the longest wings in proportion to breadth of all Indian butterflies.
Detailed description
Edward Yerbury Watson
Edward Yerbury Watson (27 July 1864 – 8 November 1897) was an English entomologist who specialised in Lepidoptera most notably Hesperiidae.
Yerbury Watson was born in British India to Joanna and George Edward Watson. He joined the North Lancas ...
(1891) gives a detailed description:
Genus characters
Species description
Similar species
Unlike the other awls, the brown awl lacks the narrow white wing bands on the hindwings. The very distinctive characteristics of the brown awl are the characteristic shape of the body and the narrower wings than the other awls.[
]
Life cycle
Eggs
The brown awl lays many eggs on a single plant, one at a time, on the tips of fresh shoots. The dome-shaped egg is pale green with longitudinal ridges having fine beadings; a total 13 ridges in all.
Larva
The larva is a pale violaceous (violet) yellow, with numerous black transverse dorsal lines; the prolegs are whitish encircled with black.[ The head is yellow, approximately heart shaped, with a black band and many tiny black spots.][
On hatching the larva webs the edges of leaves together with silk] to form a roomy cell from a leaf in which it resides throughout the larval stage. When disturbed, it can move quite briskly and even drop off. The caterpillars of the brown awl grow faster than most of those of other families, and have moist, sticky droppings.[
At the time of pupation they descend close to the ground, looking for suitable spots to pupate. The caterpillar constructs a tubular cell from a leaf by drawing the edges together with thick strands of silk.] In this cell, the caterpillar prepares an extensive silk bed on which it sits awaiting pupation. The freshly formed pupa clings onto the silken pad almost immediately.[
]
Pupa
The pupa is stubby, with protruding eyes and a prominent projection on the head in between them.[ The pupa may be light brown][ or violaceous.][ The body tapers away from the shoulders towards the rear. The abdomen is creamish with a row of four black spots on each side. The pupa is shiny, but plastered with a white powder.][
]
Host plants
The caterpillars have been recorded on the following deciduous and semi-evergreen forest plants, mostly from the family Combretaceae
The Combretaceae, often called the white mangrove family, are a family of flowering plants in the order Myrtales. The family includes about 530 species of trees, shrubs, and lianas in ca 10 genera. The family includes the leadwood tree, '' Combr ...
:[
* Large climbing shrubs of '']Combretum
''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to tropical and southern Africa, about 5 to Madagascar, but the ...
'' such as ''Combretum albidum
''Combretum'', the bushwillows or combretums, make up the type genus of the family Combretaceae. The genus comprises about 272 species of trees and shrubs, most of which are native to Tropics, tropical and southern Africa, about 5 to Madagascar, ...
'',[ '' Combretum latifolium''][ and '' Combretum ovalifolium''.]
* Large forest trees of genus '' Terminalia'' such as '' Terminalia bellerica'',[ '']Terminalia catappa
''Terminalia catappa'' is a large tropical tree in the leadwood tree family, Combretaceae, native to Asia, Australia, the Pacific, Madagascar and Seychelles. Common names in English include country almond, Indian almond, Malabar almond, sea almon ...
'' and ''Terminalia oblongata
''Terminalia oblongata'', commonly known as yellow-wood or rosewood, is a species of plant in the Combretaceae family. It is native to Australia
Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign count ...
''.
* ''