Aythya Affinis
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''Aythya'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
diving duck The diving ducks, commonly called pochards or scaups, are a category of duck which feed by diving beneath the surface of the water. They are part of Anatidae, the diverse and very large family that includes ducks, geese, and swans. The diving ...
s, with twelve species currently accepted. The genus was described in 1822 by the German zoologist
Friedrich Boie Friedrich Boie (4 June 1789 – 3 March 1870) was a German entomologist, herpetologist, ornithologist, and lawyer.Beolens, Bo; Watkins, Michael; Grayson, Michael (2011). ''The Eponym Dictionary of Reptiles''. Baltimore: Johns Hopkins Universi ...
, with the
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
being
greater scaup The greater scaup (''Aythya marila''), just scaup in Europe or, colloquially, "bluebill" in North America, is a mid-sized diving duck, larger than the closely related lesser scaup and tufted duck. It spends the summer months breeding in Iceland ...
. The name ''Aythya'' comes from the Ancient Greek word (), which referred to an unknown diving-bird. The species are plump, compact, medium-sized ducks ranging from 37–61 cm long, 60–84 cm wingspan, and weighing 410–1600 g, with
canvasback The canvasback (''Aythya valisineria'') is a species of diving duck, the largest found in North America. Taxonomy Scottish-American naturalist Alexander Wilson described the canvasback in 1814. The genus name is derived from Greek ''aithuia'', ...
the largest, and
ring-necked duck The ring-necked duck (''Aythya collaris'') is a diving duck from North America commonly found in freshwater ponds and lakes. The scientific name is derived from Greek , an unidentified seabird mentioned by authors including Hesychius and Arist ...
and
ferruginous duck The ferruginous duck (''Aythya nyroca''), also known as ferruginous pochard, common white-eye or white-eyed pochard, is a medium-sized diving duck from Eurosiberia. The scientific name is derived from the Ancient Greek word, (), an unknown se ...
marginally the smallest. The body plumage is variably white, grey, red-brown, or black, often with a finely
vermiculated Vermiculation is a surface pattern of dense but irregular lines, so called from the Latin meaning "little worm" because the shapes resemble worms, worm casts, or worm tracks in mud or wet sand. The word may be used in a number of contexts for ...
pattern; in several species, the flanks are white or pale grey, the back darker grey to black, and the breast and tail black. The heads are strongly coloured in the males, orange-red in some species, and black with a green to purple sheen (
structural colour Structural coloration in animals, and a few plants, is the production of colour by microscopically structured surfaces fine enough to interfere with visible light instead of pigments, although some structural coloration occurs in combination wi ...
) in good light; the sheen colour varies with both species and angle of light incidence – in for example
greater scaup The greater scaup (''Aythya marila''), just scaup in Europe or, colloquially, "bluebill" in North America, is a mid-sized diving duck, larger than the closely related lesser scaup and tufted duck. It spends the summer months breeding in Iceland ...
, the head has a green sheen in direct light, but a purple sheen when backlit. In females, the heads are browner, sometimes with white patterning on the face. The eye colour is also variable in males, from white in
ferruginous duck The ferruginous duck (''Aythya nyroca''), also known as ferruginous pochard, common white-eye or white-eyed pochard, is a medium-sized diving duck from Eurosiberia. The scientific name is derived from the Ancient Greek word, (), an unknown se ...
(leading to its archaic name of "white-eyed pochard"), through yellow in most species, to deep red in
common pochard The common pochard (; ''Aythya ferina''), known simply as pochard in the United Kingdom, is a medium-sized diving duck in the family Anatidae. It is widespread across the Palearctic. It breeds primarily in the steppe regions of Scandinavia and Si ...
and
canvasback The canvasback (''Aythya valisineria'') is a species of diving duck, the largest found in North America. Taxonomy Scottish-American naturalist Alexander Wilson described the canvasback in 1814. The genus name is derived from Greek ''aithuia'', ...
; in females, the eye is brown in all species. The bills are short, fairly broad, and pale blue-grey to black, usually with a small black 'nail' at the tip; the pattern of blue-grey and black is important in species identification. In flight, the wings are dark grey to black, with a white to pale grey wingbar along the primary and secondary feathers; the pattern of the wingbar (whether all-white, or all-grey, or white on the secondaries and grey on the primaries) is an important identification feature. The
webbed feet The webbed foot is a specialized limb with interdigital membranes (webbings) that aids in aquatic locomotion, present in a variety of tetrapod vertebrates. This adaptation is primarily found in semiaquatic species, and has convergently evolved ...
, used for propulsion in both swimming and diving, are 5–7 cm long, large for the size of the birds, and dark grey to blackish in all the species. Svensson, L., Mullarney, K., & Zetterström, D. (2022) ''
Collins Bird Guide The ''Collins Bird Guide'' is a field guide to the birds of the Western Palearctic. Its authors are Lars Svensson, Killian Mullarney, Dan Zetterström and Peter J. Grant, and it is illustrated by Killian Mullarney and Dan Zetterström (wit ...
'', ed. 3. , pages 30-35
The species occur throughout Asia, Europe, North America, Australia, New Zealand, and Madagascar, and also in the northern half of Africa primarily in winter. Small numbers also reach the far north of South America in winter. In the breeding season, they are restricted to well-vegetated freshwater lakes, while in the winter they use both freshwater lakes and sheltered saltwater bays and inlets.


''Aythya'' species

The genus contains 12 species; all are monotypic except for ''A. marila'', which has two subspecies in the Old and New Worlds, respectively.


Hybrids

The species in the genus are all closely related, and are more prone to hybridisation than most other bird genera. Hybrids regularly seen in Europe include tufted duck × common pochard, tufted duck × ring-necked duck, greater scaup × tufted duck, and common pochard × ferruginous duck; while in North America, ring-necked duck × greater scaup, greater scaup × tufted duck, and canvasback × redhead are frequent. These hybrids can often resemble, and be mistaken for, other species in the genus; for example tufted duck × common pochard hybrids are easily mistaken for lesser scaup. Usually only male hybrids are evident; female hybrids are less obvious and even more difficult to identify.


Diet

The diet, mostly obtained by diving to depths of 0.5–6 m (exceptionally 10 m), but also at times from the surface without diving, consists of a mixture of plant material (including seeds, leaves and roots of water plants) and bottom-dwelling invertebrates (including worms, molluscs, insects). In urban situations, several species have learnt to take bread or birdseed fed to ducks by people.


Breeding

The nests are hidden in dense waterside vegetation, for preference on islets which give greater security from land predators. Incubation of the
The dash is a punctuation mark consisting of a long horizontal line. It is similar in appearance to the hyphen but is longer and sometimes higher from the baseline. The most common versions are the endash , generally longer than the hyphen ...
–10 18eggs takes 3–4 weeks, and is done entirely by the female. The ducklings fledge at around 6–7 weeks old, and are cared for primarily by the female, though the male may assist with guarding the ducklings. 'Dump nesting', where more than one female lays eggs in a nest, is common, and likely accounts for nests containing more than ten eggs.


Prehistory

''Aythya shihuibas'' was described from the
Late Miocene The Late Miocene (also known as Upper Miocene) is a sub-epoch of the Miocene epoch (geology), Epoch made up of two faunal stage, stages. The Tortonian and Messinian stages comprise the Late Miocene sub-epoch, which lasted from 11.63 Ma (million ye ...
of
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
, but probably belongs outside
crown group In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor ...
''Aythya''. Zelenkov (2016) transferred the species ''
Anas ''Anas'' is a genus of dabbling ducks. It includes the pintails, most teals, and the mallard and its close relatives. It formerly included additional species but following the publication of a molecular phylogenetic study in 2009 the genus was s ...
denesi'' Kessler (2013), known from the late Miocene of
Hungary Hungary is a landlocked country in Central Europe. Spanning much of the Pannonian Basin, Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia and ...
, to the genus ''Aythya''. An undescribed prehistoric species is known only from
Early Pleistocene The Early Pleistocene is an unofficial epoch (geology), sub-epoch in the international geologic timescale in chronostratigraphy, representing the earliest division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently esti ...
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
remains found at Dursunlu,
Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
; it might however be referrable to a
paleosubspecies A chronospecies is a species derived from a sequential development pattern that involves continual and uniform changes from an extinct ancestral form on an evolutionary scale. The sequence of alterations eventually produces a population that is p ...
of an extant species considering its age (see also
Greater scaup The greater scaup (''Aythya marila''), just scaup in Europe or, colloquially, "bluebill" in North America, is a mid-sized diving duck, larger than the closely related lesser scaup and tufted duck. It spends the summer months breeding in Iceland ...
). Subfossils have also been found on
Réunion Réunion (; ; ; known as before 1848) is an island in the Indian Ocean that is an overseas departments and regions of France, overseas department and region of France. Part of the Mascarene Islands, it is located approximately east of the isl ...
; this
Réunion pochard The Réunion pochard (''Aythya'' sp.), also known as the Mascarene Islands pochard, is an undescribed extinct species of diving duck that lived on the island of Réunion. In 1999 there were two carpometacarpi subfossils found by Cécile Mourer ...
awaits formal description, and may prove to have been a population of the
Madagascar pochard The Madagascar pochard or Madagascan pochard (''Aythya innotata''; ) is an extremely rare diving duck of the genus ''Aythya''. Thought to be extinct in the late 1990s, specimens of the species were rediscovered at Lake Matsaborimena near Bemanevi ...
. The
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
''"Aythya" arvernensis'' is now placed in '' Mionetta,'' while ''"Aythya" chauvirae'' seems to contain the remains of two species, at least one of which does not seem to be a diving duck. Worthy, Trevor; Tennyson, A.J.D.; Jones, C.; McNamara, J.A. & Douglas, B.J. (2007): Miocene waterfowl and other birds from central Otago, New Zealand. '' J. Syst. Palaeontol.'' 5(1): 1–39. (HTML abstract)


See also

*
Late Quaternary prehistoric birds Late Quaternary prehistoric birds are Bird, avian taxa that became extinct during the Late Quaternary – the Late Pleistocene or Early Holocene – and before recorded history, specifically before they could be studied alive by orni ...
*
List of fossil bird genera Birds evolved from certain feathered theropod dinosaurs, and there is no real dividing line between birds and non-avian dinosaurs except that some of the former survived the Cretaceous–Paleogene extinction event while the latter did not. For ...


Gallery

File:Canvasback pair2.jpg,
Canvasback The canvasback (''Aythya valisineria'') is a species of diving duck, the largest found in North America. Taxonomy Scottish-American naturalist Alexander Wilson described the canvasback in 1814. The genus name is derived from Greek ''aithuia'', ...
(''Aythya valisineria'') File:Aythya ferina (Marek Szczepanek).jpg,
Common pochard The common pochard (; ''Aythya ferina''), known simply as pochard in the United Kingdom, is a medium-sized diving duck in the family Anatidae. It is widespread across the Palearctic. It breeds primarily in the steppe regions of Scandinavia and Si ...
(''Aythya ferina'') File:Redhead in Central Park (15369).jpg,
Redhead Red hair, also known as ginger hair, is a human hair color found in 2–6% of people of Northern or Northwestern European ancestry and lesser frequency in other populations. It is most common in individuals homozygous for a recessive alle ...
(''Aythya americana'') File:Aythya-collaris-001.jpg,
Ring-necked duck The ring-necked duck (''Aythya collaris'') is a diving duck from North America commonly found in freshwater ponds and lakes. The scientific name is derived from Greek , an unidentified seabird mentioned by authors including Hesychius and Arist ...
(''Aythya collaris'') File:Aythya australis female - Hurstville Golf Course.jpg, Female
Hardhead The hardhead (''Aythya australis''), also known as the white-eyed duck, is the only true diving duck found in Australia. The common name "hardhead" has nothing to do with the density of the bird's skull, instead referring to the difficulty encou ...
(''Aythya australis'') File:Aythya baeri cropped.jpg,
Baer's pochard Baer's pochard (''Aythya baeri'') is a diving duck found in eastern Asia. It is a bird in North and Central China, and formerly bred in southeast Russia and Northeast China, migrating in winter to southern China, Vietnam, Japan, and India. Baer ...
(''Aythya baeri'') File:Moretta Tabaccata - Ferruginous Duck - Aythya nyroca.jpg,
Ferruginous duck The ferruginous duck (''Aythya nyroca''), also known as ferruginous pochard, common white-eye or white-eyed pochard, is a medium-sized diving duck from Eurosiberia. The scientific name is derived from the Ancient Greek word, (), an unknown se ...
(''Aythya nyroca'') File:AythyaInnotata.svg,
Madagascar pochard The Madagascar pochard or Madagascan pochard (''Aythya innotata''; ) is an extremely rare diving duck of the genus ''Aythya''. Thought to be extinct in the late 1990s, specimens of the species were rediscovered at Lake Matsaborimena near Bemanevi ...
(''Aythya innotata'') File:NZ Scaup 01.jpg,
New Zealand scaup The New Zealand scaup (''Aythya novaeseelandiae''), also known as the black teal (or in Māori language, Māori), is a diving duck species of the genus ''Aythya'' endemic (ecology), endemic to New Zealand. They weigh around and measure around ...
(''Aythya novaeseelandiae'') File:Tufted-Duck-male-female.jpg,
Tufted duck The tufted duck (or tufted pochard) (''Aythya fuligula'') is a small diving duck with a population of nearly one million birds, found in northern Eurasia. They are partially migratory. The scientific name is derived from Ancient Greek , an unide ...
(''Aythya fuligula'') File:2017-03-24 Aythya marila, pair, Killingworth Lake, Northumberland 01.jpg,
Greater scaup The greater scaup (''Aythya marila''), just scaup in Europe or, colloquially, "bluebill" in North America, is a mid-sized diving duck, larger than the closely related lesser scaup and tufted duck. It spends the summer months breeding in Iceland ...
(''Aythya marila'') File:AythyaAffins 2109.JPG,
Lesser scaup The lesser scaup (''Aythya affinis'') is a small North American diving duck that migrates south as far as Central America in winter. It is colloquially known as the little bluebill or broadbill because of its distinctive blue bill. The origin of ...
(''Aythya affinis'')


References

{{Taxonbar, from=Q214300 Diving ducks Bird genera Taxa named by Friedrich Boie