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Ayres's hawk-eagle (''Hieraaetus ayresii''), also referred to as Ayres' eagle,Newman, K (1998) Newman's Birds of Southern Africa. Halfway House: Southern Book Publishers. . is a medium-sized
bird of prey Birds of prey or predatory birds, also known as (although not the same as) raptors, are hypercarnivorous bird species that actively predation, hunt and feed on other vertebrates (mainly mammals, reptiles and smaller birds). In addition to speed ...
in the family
Accipitridae The Accipitridae () is one of the four families within the order Accipitriformes, and is a family of small to large birds of prey with strongly hooked bills and variable morphology based on diet. They feed on a range of prey items from insects ...
. It is native to African woodlands. Its name honours
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
n
ornithologist Ornithology, from Ancient Greek ὄρνις (''órnis''), meaning "bird", and -logy from λόγος (''lógos''), meaning "study", is a branch of zoology dedicated to the study of birds. Several aspects of ornithology differ from related discip ...
Thomas Ayres.


Description

The adult male has blackish upper parts that are mottled with white and usually has a white forehead and
supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also k ...
. The upper-wing
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are small feathers behind t ...
are similar. The tail is ashy grey with a broad black tip and three to four narrower dark bars.
Primary feathers Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tai ...
and
secondary feathers Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the Bird wing, wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those ...
are black. The underparts are white, with heavy dark brown spots and blotches on the breast and belly, becoming sparser on the thighs and vent. The legs are well-feathered and pure white. The under-wing coverts are brown and marked with white, and the underside of flight feathers is dark and heavily barred lacking any noticeable grey patch. The eyes are yellow to orange, the
cere The beak, bill, or Rostrum (anatomy), rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for pecking, wikt:grasp#Verb, grasping, and holding (in wikt:probe ...
and feet yellow, and the bill is bluish horn coloured becoming paler towards the base, with a black tip. This species has a small but at times pronounced crest. The males are smaller than the females, darker and usually more densely spotted on the underparts, and have a smaller amount of white on the forehead and supercilium. There are two
phases Phase or phases may refer to: Science *State of matter, or phase, one of the distinct forms in which matter can exist *Phase (matter), a region of space throughout which all physical properties are essentially uniform *Phase space, a mathematica ...
, the normal as described above and a
melanistic Melanism is the congenital excess of melanin in an organism resulting in dark pigment. Pseudomelanism, also called abundism, is another variant of pigmentation, identifiable by dark spots or enlarged stripes, which cover a large part of the bod ...
phase, which is mostly black with white markings. The juvenile Ayres's hawk-eagle is dark brown on the back and coverts, similarly dark on the crown, slightly paler necked, largely white below from the throat to the crissum and legs with sparse dark brown markings variably along the chest, belly, and flanks. When maturing, the young hawk-eagles often manifest a duskier, browner colour with somewhat of a rufous cast from below before moulting in its mature, adult
plumage Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...
. Mostly the species is considered confusable with the
African hawk-eagle The African hawk-eagle (''Aquila spilogaster'') is a large bird of prey. Like all eagles, it belongs to the family ''Accipitridae''. This species' feathered legs mark it as a member of the Aquilinae subfamily. The African hawk-eagle breeds in tr ...
which is larger and lankier with a more protruding head, a longer tail, more sparsely marked underparts, and more heavily marked wings with a white window above. It shares white "landing lights" at the fore of the wings with the similarly sized
booted eagle The booted eagle (''Hieraaetus pennatus'', also classified as ''Aquila pennata'') is a medium-sized mostly migratory bird of prey with a wide distribution in the Palearctic and southern Asia, wintering in the tropics of Africa and Asia, with a ...
but that species is usually rather differently marked overall (either paler and browner or all dark in dark morphs) and also similarly sized
black sparrowhawk The black sparrowhawk (''Astur melanoleucus''), sometimes known as the black goshawk or great sparrowhawk, is a bird of prey belonging to the family (biology), family Accipitridae. It was formerly placed in the genus ''Accipiter''. It occurs mai ...
s are mentioned as a confusion species but sparrowhawks are rather differently proportioned with bare, dull green-yellow legs. The Ayres's hawk-eagle is a medium-sized raptor but quite a small eagle, about the weight of a
peregrine falcon The peregrine falcon (''Falco peregrinus''), also known simply as the peregrine, is a Cosmopolitan distribution, cosmopolitan bird of prey (raptor) in the family (biology), family Falconidae renowned for its speed. A large, Corvus (genus), cro ...
albeit with a slightly greater length and
wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the opposite wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingsp ...
more akin to a
common buzzard The common buzzard (''Buteo buteo'') is a medium-to-large bird of prey which has a large range. It is a member of the genus '' Buteo'' in the family Accipitridae. The species lives in most of Europe and extends its breeding range across much of ...
, with a total length of and has a wingspan of .Kemp, A. C., P. F. D. Boesman, and J. S. Marks (2020). ''Ayres's Hawk-Eagle (Hieraaetus ayresii)'', version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. A small sample of males have been known to weigh from and females weigh from . The average weight of four adult males was while that of 10 adult females was , juvenile Ayres's being some 5% lighter in spite of slightly greater wing sizes.Hartley, R. R., & Mundy, P. J. (2003). Morphometrics and status of Ayres's Hawk-eagle in Zimbabwe. Journal of Raptor Research, 37(1), 44-54. The wing chord is and from in males and females, respectively. The tail and tarsus length is from and from in males and and from in females. The species thus manifests quite strong
sexual dimorphism Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...
in favor of the female, which does not overlap in standard measurements with the smaller male. Bill length is from , averaging in males and in females. The enlarged hallux claw on the rear toe is from , averaging in males and in females, quite large for the small size of the eagle.Parry, S. J. (2001). ''The booted eagles (Aves: Accipitridae): perspectives in evolutionary biology''. University of London, University College London (United Kingdom).


Distribution and habitat

Ayres's hawk-eagle has a patchy sub-Saharan distribution ranging from
Sierra Leone Sierra Leone, officially the Republic of Sierra Leone, is a country on the southwest coast of West Africa. It is bordered to the southeast by Liberia and by Guinea to the north. Sierra Leone's land area is . It has a tropical climate and envi ...
east to
Somalia Somalia, officially the Federal Republic of Somalia, is the easternmost country in continental Africa. The country is located in the Horn of Africa and is bordered by Ethiopia to the west, Djibouti to the northwest, Kenya to the southwest, th ...
, and south to northern
Namibia Namibia, officially the Republic of Namibia, is a country on the west coast of Southern Africa. Its borders include the Atlantic Ocean to the west, Angola and Zambia to the north, Botswana to the east and South Africa to the south; in the no ...
and northeast
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
. Its central range is from the southern
Democratic Republic of the Congo The Democratic Republic of the Congo (DRC), also known as the DR Congo, Congo-Kinshasa, or simply the Congo (the last ambiguously also referring to the neighbouring Republic of the Congo), is a country in Central Africa. By land area, it is t ...
and southern
Uganda Uganda, officially the Republic of Uganda, is a landlocked country in East Africa. It is bordered to the east by Kenya, to the north by South Sudan, to the west by the Democratic Republic of the Congo, to the south-west by Rwanda, and to the ...
down almost throughout
Kenya Kenya, officially the Republic of Kenya, is a country located in East Africa. With an estimated population of more than 52.4 million as of mid-2024, Kenya is the 27th-most-populous country in the world and the 7th most populous in Africa. ...
,
Tanzania Tanzania, officially the United Republic of Tanzania, is a country in East Africa within the African Great Lakes region. It is bordered by Uganda to the northwest; Kenya to the northeast; the Indian Ocean to the east; Mozambique and Malawi to t ...
,
Zambia Zambia, officially the Republic of Zambia, is a landlocked country at the crossroads of Central Africa, Central, Southern Africa, Southern and East Africa. It is typically referred to being in South-Central Africa or Southern Africa. It is bor ...
and
Mozambique Mozambique, officially the Republic of Mozambique, is a country located in Southeast Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi and Zambia to the northwest, Zimbabwe to the west, and Eswatini and South Afr ...
as well as much of
Angola Angola, officially the Republic of Angola, is a country on the west-Central Africa, central coast of Southern Africa. It is the second-largest Portuguese-speaking world, Portuguese-speaking (Lusophone) country in both total area and List of c ...
to northeastern
Namibia Namibia, officially the Republic of Namibia, is a country on the west coast of Southern Africa. Its borders include the Atlantic Ocean to the west, Angola and Zambia to the north, Botswana to the east and South Africa to the south; in the no ...
, northern
Botswana Botswana, officially the Republic of Botswana, is a landlocked country in Southern Africa. Botswana is topographically flat, with approximately 70 percent of its territory part of the Kalahari Desert. It is bordered by South Africa to the sou ...
and northern
Zimbabwe file:Zimbabwe, relief map.jpg, upright=1.22, Zimbabwe, relief map Zimbabwe, officially the Republic of Zimbabwe, is a landlocked country in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Bots ...
. In the rainy season moves out of denser and taller deciduous
woodland A woodland () is, in the broad sense, land covered with woody plants (trees and shrubs), or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunli ...
s of central Africa into more open albeit treed
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...
habitat further south, and probably into coastal East Africa; as the rains cause the leaves to emerge turning woodland into
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
s and tree savanna into woodland. It may then enter towns in South Africa to prey mainly on
doves Columbidae is a bird family consisting of doves and pigeons. It is the only family in the order Columbiformes. These are stout-bodied birds with small heads, relatively short necks and slender bills that in some species feature fleshy ceres. ...
and feral pigeons. A similar north to south movement is expected in west Africa, where species has been recorded as a
vagrant Vagrancy is the condition of wandering homelessness without regular employment or income. Vagrants usually live in poverty and support themselves by travelling while engaging in begging, scavenging, or petty theft. In Western countries, ...
west to
Senegal Senegal, officially the Republic of Senegal, is the westernmost country in West Africa, situated on the Atlantic Ocean coastline. It borders Mauritania to Mauritania–Senegal border, the north, Mali to Mali–Senegal border, the east, Guinea t ...
and
Gambia The Gambia, officially the Republic of The Gambia, is a country in West Africa. Geographically, The Gambia is the List of African countries by area, smallest country in continental Africa; it is surrounded by Senegal on all sides except for ...
, albeit unconfirmed. It tends to occur around woodlands of ''
Brachystegia ''Brachystegia'' is a genus of tree of the subfamily Detarioideae that is native to tropical Africa. Trees of the genus are commonly known as miombo, and are dominant in the miombo woodlands of central and southern tropical Africa. The Zambezia ...
'' and '' Baikiaea'' in the core south-central African part of their distribution. It is not typically a forest dwelling species and prefers mature woodland among often irregular,
rocky ''Rocky'' is a 1976 American independent film, independent sports drama film directed by John G. Avildsen and written by and starring Sylvester Stallone. It is the first installment in the Rocky (film series), ''Rocky'' franchise and also star ...
terrain. Occasionally but not regularly they may habituate to ''
Acacia ''Acacia'', commonly known as wattles or acacias, is a genus of about of shrubs and trees in the subfamily Mimosoideae of the pea family Fabaceae. Initially, it comprised a group of plant species native to Africa, South America, and Austral ...
''
plantation Plantations are farms specializing in cash crops, usually mainly planting a single crop, with perhaps ancillary areas for vegetables for eating and so on. Plantations, centered on a plantation house, grow crops including cotton, cannabis, tob ...
s and
riparian zone A riparian zone or riparian area is the interface between land and a river or stream. In some regions, the terms riparian woodland, riparian forest, riparian buffer zone, riparian corridor, and riparian strip are used to characterize a ripari ...
s. Ayres's hawk-eagles tend to be rare in hyper
arid Aridity is the condition of geographical regions which make up approximately 43% of total global available land area, characterized by low annual precipitation, increased temperatures, and limited water availability.Perez-Aguilar, L. Y., Plata ...
and, of course, treeless areas.


Biology

Ayres's hawk-eagle is a bird hunter, almost to the exclusion of any other type of prey, especially
dove Columbidae is a bird family consisting of doves and pigeons. It is the only family in the order Columbiformes. These are stout-bodied birds with small heads, relatively short necks and slender bills that in some species feature fleshy ceres. ...
s and
pigeon Columbidae is a bird family consisting of doves and pigeons. It is the only family in the order Columbiformes. These are stout-bodied birds with small heads, relatively short necks and slender bills that in some species feature fleshy ceres. ...
s, it soars high above the ground to search for prey. It is unique for a
Aquilinae The Aquilinae are a subfamily of eagles of the family Accipitridae. The general common name used for members of this subfamily is "booted eagle", although this is also the common name of a member of the subfamily.Lerner, H., Christidis, L., Gamau ...
eagle in its
falcon Falcons () are birds of prey in the genus ''Falco'', which includes about 40 species. Some small species of falcons with long, narrow wings are called hobbies, and some that hover while hunting are called kestrels. Falcons are widely distrib ...
-like highly aerial method for hunting birds on the wing, an aptitude it shares with the Asian
rufous-bellied eagle The rufous-bellied eagle or rufous-bellied hawk-eagle (''Lophotriorchis kienerii'') is a bird of prey in the family Accipitridae that is found in the forested regions of tropical Asia. Relatively small for eagles and contrastingly patterned like ...
. Favoured in diet in some areas are
laughing dove The laughing dove (''Spilopelia senegalensis'') is a small pigeon that is a resident breeder in Africa, the Middle East, South Asia, and Western Australia where it has established itself in the wild after being released from Perth Zoo in 1898. Th ...
s and
red-eyed dove The red-eyed dove (''Streptopelia semitorquata'') is a dove that is widespread and common in Sub-Saharan Africa. It has been listed as Least Concern on the IUCN Red List since 2004. Taxonomy The red-eyed dove was formally described by the Germ ...
s. Once a bird has been singled out, the eagle stoops to intercept it in mid air. Other than birds it has been recorded as catching a few mammals including
bush squirrel The African bush squirrels are a genus of squirrels, ''Paraxerus'', in the subfamily Xerinae. They are only found in Africa. The 11 species in this genus are: *Alexander's bush squirrel (''P. alexandri'') * Boehm's bush squirrel (''P. boehmi'') * ...
s, and
fruit bat Megabats constitute the family Pteropodidae of the order Chiroptera. They are also called fruit bats, Old World fruit bats, or—especially the genera ''Acerodon'' and ''Pteropus''—flying foxes. They are the only member of the superfamil ...
s. They are an exceptional agile and fast eagle and can overtake many birds in flight. The Ayres's hawk-eagle predominantly hunt quite small birds, typically weighing from , and can sometimes include nestling birds in their diet. However, it quite commonly takes heavier birds than in the aforementioned weight range, including
rock pigeon The rock dove (''Columba livia''), also sometimes known as "rock pigeon" or "common pigeon", is a member of the bird family Columbidae (doves and pigeons). In common usage, it is often simply referred to as the "pigeon", although the rock dov ...
s and African grey hornbills. Sometimes they will overtake birds up to over twice their size including
white-faced whistling duck The white-faced whistling duck (''Dendrocygna viduata'') is a whistling duck that breeds in sub-Saharan Africa and much of South America. This species is gregarious, and at favoured sites, the flocks of a thousand or more birds arriving at dawn ...
s, assorted
francolin Francolins are birds in the tribe Gallini that traditionally have been placed in the genus ''Francolinus'', but now commonly are divided into multiple genera. As previously defined, they were paraphyletic as the genus '' Pternistis'', which wa ...
s and
guineafowl Guinea fowl () (or guineahen) are birds of the family Numididae in the order Galliformes. They are endemic to Africa and rank among the oldest of the gallinaceous birds. Phylogenetically, they branched off from the core Galliformes after the C ...
s and even other birds of prey including
gabar goshawk The gabar goshawk (''Micronisus gabar'') is a small species of African and Arabian bird of prey in the family Accipitridae. Description The gabar goshawk is polymorphic and occurs in two distinct forms which fluctuate in relative abundance acro ...
s and
shikra The shikra (''Tachyspiza badia'') is a small bird of prey in the family Accipitridae found widely distributed in Asia and Africa where it is also called the little banded goshawk. The African forms may represent a separate species but have usual ...
s. Ayres's hawk-eagle is a monogamous, territorial solitary nester. The nest consists of a large platform of a twigs and sticks, lined with green leaves and typically concealed in the fork within a well-leafed, large tree, usually from above the ground. The nest can be from across. Eggs are laid from April to September, peaking in April and May. A single egg is laid, measuring on average . The female is almost solely responsible for incubation which takes about 43 days, the male brings her food every two to three days. The chick is fed almost daily, fledging at about 73 days old and becoming fully independent from its parents approximately three months after fledging.Dewhurst, C. E., & van Someren, C. (1988). Observation on the breeding ecology of Ayres' hawk eagle Hieraaetus ayresii at Karen, Nairobi, Kenya. GABAR., 3(2), 85-93.


Conservation status

Although Ayres's hawk-eagle is an uncommon bird throughout its range, it is classified as
Least Concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
(LC) by the IUCN, due to its large range and its numbers, while small, appearing stable at the present time. The estimated global population is only 1000 to 10,000 individuals. The main threat faced by the species is
cutting Cutting is the separation or opening of a physical object, into two or more portions, through the application of an acutely directed force. Implements commonly used for wikt:cut, cutting are the knife and saw, or in medicine and science the sca ...
of woodland. Furthermore, it is persecuted at times for hunting down
domestic pigeon The domestic pigeon (''Columba livia domestica'' or ''Columba livia'' Form (zoology), forma ''domestica'') is a pigeon subspecies that was derived from the rock dove, rock dove or rock pigeon. The rock pigeon is the world's oldest domesticated ...
s.


References


External links

* (Ayres's hawk-eagle = ) Ayres' eagle
Species text in The Atlas of Southern African Birds
{{Taxonbar, from=Q1260400
Ayres's hawk-eagle Ayres's hawk-eagle (''Hieraaetus ayresii''), also referred to as Ayres' eagle,Newman, K (1998) Newman's Birds of Southern Africa. Halfway House: Southern Book Publishers. . is a medium-sized bird of prey in the family Accipitridae. It is native t ...
Birds of prey of Sub-Saharan Africa
Ayres's hawk-eagle Ayres's hawk-eagle (''Hieraaetus ayresii''), also referred to as Ayres' eagle,Newman, K (1998) Newman's Birds of Southern Africa. Halfway House: Southern Book Publishers. . is a medium-sized bird of prey in the family Accipitridae. It is native t ...
Eagles
Ayres's hawk-eagle Ayres's hawk-eagle (''Hieraaetus ayresii''), also referred to as Ayres' eagle,Newman, K (1998) Newman's Birds of Southern Africa. Halfway House: Southern Book Publishers. . is a medium-sized bird of prey in the family Accipitridae. It is native t ...
Taxonomy articles created by Polbot