''Avalonnectes'' is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of small-bodied rhomaleosaurid known from the

Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic� ...

period Period may refer to: Common uses * Period (punctuation) * Era, a length or span of time *Menstruation, commonly referred to as a "period" Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (o ...

(most likely earliest

Hettangian The Hettangian is the earliest age and lowest stage of the Jurassic Period of the geologic timescale. It spans the time between 201.3 ± 0.2 Ma and 199.3 ± 0.3 Ma (million years ago). The Hettangian follows the Rhaetian (part of the Triass ...

stage) of the United Kingdom. It contains a single species, ''A. arturi''.

Discovery

''Avalonnectes'' is known from the

holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...

specimen

NHMUK The Natural History Museum in London is a museum that exhibits a vast range of specimens from various segments of natural history. It is one of three major museums on Exhibition Road in South Kensington, the others being the Science Museum and ...

14550, which consists of the posterior portion of a

skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...

, and a nearly complete, three-dimensionally preserved and articulated

postcrania The postcranium ("behind the cranium"; plural: postcrania) or postcranial skeleton in zoology and vertebrate paleontology is the skeleton apart from the skull. The postcranium encompasses the axial skeleton, which includes the entirety of the verte ...

skeleton A skeleton is the structural frame that supports the body of most animals. There are several types of skeletons, including the exoskeleton, which is a rigid outer shell that holds up an organism's shape; the endoskeleton, a rigid internal fra ...

. Another partial postcranial skeleton which was referred to it is AGT uncatalogued. Both specimens were collected at

Street A street is a public thoroughfare in a city, town or village, typically lined with Building, buildings on one or both sides. Streets often include pavements (sidewalks), pedestrian crossings, and sometimes amenities like Street light, streetligh ...

, of

Somerset Somerset ( , ), Archaism, archaically Somersetshire ( , , ) is a Ceremonial counties of England, ceremonial county in South West England. It is bordered by the Bristol Channel, Gloucestershire, and Bristol to the north, Wiltshire to the east ...

, from the Pre-''Planorbis'' beds of the

Blue Lias Formation The Blue Lias is a geological formation in southern, eastern and western England and parts of South Wales, part of the Lias Group. The Blue Lias consists of a sequence of limestone and shale layers, laid down in latest Triassic and early Jurassi ...

of the

Lower Lias Group The Lias Group or Lias is a lithostratigraphy, lithostratigraphic unit (a sequence of rock strata) found in a large area of western Europe, including the British Isles, the North Sea, the Low Countries and the north of Germany. It consists of Sed ...

. These beds likely occur below the first occurrence of the ammonite '' Psiloceras planorbis''. Thus, they probably fall within the earliest Hettangian ''P. tilmanni'' Chronozone, which is about 199.6-198 million years old, immediately following the

Triassic The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ...

–

Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...

Boundary. Plesiosaurs fossils which were discovered at Street represent the earliest known occurrence of the

Plesiosauria The Plesiosauria or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared in the latest Triassic Period, possibly in the Rhaetian stage, about 203 million year ...

. Hence, ''Avalonnectes'' is one of the oldest plesiosaurs to date.

Description

''Avalonnectes'' is a small-bodied rhomaleosaurid, with a trunck length of and a neck length of more than . The number of

dorsal vertebrae In vertebrates, thoracic vertebrae compose the middle segment of the vertebral column, between the cervical vertebrae and the lumbar vertebrae. In humans, there are twelve thoracic vertebra (anatomy), vertebrae of intermediate size between the ce ...

are about 18–19; this low number represents its

autapomorphy In phylogenetics, an autapomorphy is a distinctive feature, known as a Synapomorphy, derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or Outgroup (cladistics), outgroup taxa, not ...

Classification

phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...

analysis performed by Benson ''et al.'' (2012) found it to be a basal rhomaleosaurid. The

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

below shows ''Avalonnectes'' phylogenetic position among other plesiosaurs following Benson ''et al.'' (2012).

Etymology

''Avalonnectes'' was first described and named by Roger B. J. Benson, Mark Evans and Patrick S. Druckenmiller in

2012 2012 was designated as: *International Year of Cooperatives *International Year of Sustainable Energy for All Events January *January 4 – The Cicada 3301 internet hunt begins. * January 12 – Peaceful protests begin in the R ...

and the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

is ''Avalonnectes arturi''. The generic name is derived from

Avalon Avalon () is an island featured in the Arthurian legend. It first appeared in Geoffrey of Monmouth's 1136 ''Historia Regum Britanniae'' as a place of magic where King Arthur's sword Excalibur was made and later where Arthur was taken to recove ...

, an island from the legend of

King Arthur According to legends, King Arthur (; ; ; ) was a king of Great Britain, Britain. He is a folk hero and a central figure in the medieval literary tradition known as the Matter of Britain. In Wales, Welsh sources, Arthur is portrayed as a le ...

, often identified with

Glastonbury Glastonbury ( , ) is a town and civil parish in Somerset, England, situated at a dry point on the low-lying Somerset Levels, south of Bristol. The town had a population of 8,932 in the 2011 census. Glastonbury is less than across the River ...

, near Street, and from

Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...

''nectes'', meaning "swimmer". The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

honors the paleontologist Arthur Cruickshank (1932–2011), who with M. A. Taylor initiated the restudy of British Lower Jurassic plesiosaurians in the 1990s. It is also a reference to the legendary King Arthur of British folklore.

References

{{Taxonbar, from1=Q18558972, from2=Q367526 Early Jurassic plesiosaurs of Europe Fossil taxa described in 2012 Fossils of Great Britain Rhomaleosauridae Sauropterygian genera

Discovery

Description

Classification

Etymology

See also

References