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Australidelphia is a
superorder Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized ...
of marsupials encompassing about three-quarters of all living marsupial species, including all those native to
Australasia Australasia is a subregion of Oceania, comprising Australia, New Zealand (overlapping with Polynesia), and sometimes including New Guinea and surrounding islands (overlapping with Melanesia). The term is used in a number of different context ...
and one South American species, the monito del monte. Unlike other American marsupials, which belong to the Ameridelphia, Australidelphia’s lineage emerged in South America, with genetic evidence (
retrotransposon Retrotransposons (also called Class I transposable elements) are mobile elements which move in the host genome by converting their transcribed RNA into DNA through reverse transcription. Thus, they differ from Class II transposable elements, or ...
insertion sites) showing the monito del monte as its most ancient branch. Within this superorder, the Australian members form a distinct group (
clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
) named Eomarsupialia, though their internal relationships (branching order) is yet to be determined. Studies suggest Australidelphia originated in South America alongside other major marsupial groups, likely dispersing to Australia via
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. ...
in a single event after the monito’s lineage ( Microbiotheria) diverged, leaving other South American orders (
Didelphimorphia Opossums () are members of the marsupial order Didelphimorphia () endemic to the Americas. The largest order of marsupials in the Western Hemisphere, it comprises 126 species in 18 genera. Opossums originated in South America and entered North A ...
and Paucituberculata) as more basal.


Phylogeny

The following cladogram is a phylogeny of Australidelphia based on the work of May-Collado, Kilpatrick & Agnarsson 2015, with extinct clades from Black et al. 2012.


Taxonomy

The
order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...
s within this group are listed below: * Genus †'' Djarthia'' Godthelp, Wroe & Archer 1999 * Order Dasyuromorphia (71 species) ** Family Dasyuridae:
antechinus ''Antechinus'' (// ('ant-echinus')) is a genus of small dasyurid marsupial endemic to Australia. They resemble mice with the bristly fur of shrews. Names They are also sometimes called 'broad-footed marsupial mice', 'pouched mice', or 'Ante ...
es,
quoll Quolls (; genus ''Dasyurus'') are carnivorous marsupials native to Australia and New Guinea. They are primarily nocturnal, and spend most of the day in a den. Of the six species of quoll, four are found in Australia and two in New Guinea. Anot ...
s, dunnarts,
Tasmanian devil The Tasmanian devil (''Sarcophilus harrisii''; palawa kani: ''purinina'') is a carnivorous marsupial of the family Dasyuridae. It was formerly present across mainland Australia, but became extinct there around 3,500 years ago; it is now con ...
, and allies ** Family Myrmecobiidae: numbat ** Family †
Thylacinidae Thylacinidae is an extinct family (biology), family of carnivorous marsupials from the order Dasyuromorphia. The only species to survive into modern times was the thylacine (''Thylacinus cynocephalus''), which became Extinction, extinct in 1936. ...
:
thylacine The thylacine (; binomial name ''Thylacinus cynocephalus''), also commonly known as the Tasmanian tiger or Tasmanian wolf, was a carnivorous marsupial that was native to the Mainland Australia, Australian mainland and the islands of Tasmani ...
* Order
Diprotodontia Diprotodontia (, from Greek language, Greek "two forward teeth") is the largest extant order (biology), order of marsupials, with about 155 species, including the kangaroos, Wallaby, wallabies, Phalangeriformes, possums, koala, wombats, and many ...
(117 species) ** Family
Acrobatidae The Acrobatidae are a small family of flying and gliding animals, gliding marsupials containing two genera, each with a single species, the feathertail glider (''Acrobates pygmaeus'') from Australia and feather-tailed possum (''Distoechurus penna ...
:
feathertail glider The feathertail glider (''Acrobates pygmaeus''), also known as the pygmy gliding possum, pygmy glider, pygmy phalanger, flying phalanger and flying mouse, is a species of marsupial native to eastern Australia. It is the world's smallest gliding ...
and
feather-tailed possum The feather-tailed possum (''Distoechurus pennatus'') is a species of marsupial in the family Acrobatidae. It is found in West Papua and Papua New Guinea. It is not to be confused with the feathertail glider, the only other species in the family ...
** Family Burramyidae: pygmy possums ** Family Hypsiprymnodontidae: musky rat-kangaroo ** Family
Macropodidae Macropodidae is a Family (biology), family of marsupials that includes kangaroos, Wallaby, wallabies, tree-kangaroos, wallaroos, pademelons, quokkas, and several other groups. These genera are allied to the suborder Macropodiformes, containing ...
: kangaroos, wallabies, and allies ** Family Petauridae:
striped possum The striped possum or common striped possum (''Dactylopsila trivirgata'') is a member of the marsupial family Petauridae. It is found mainly in New Guinea. The species is black with three white stripes running head to tail, and its head has whi ...
,
Leadbeater's possum Leadbeater's possum (''Gymnobelideus leadbeateri'') is a critically endangered Phalangeriformes, possum largely restricted to small pockets of Eucalyptus delegatensis, alpine ash, Eucalyptus regnans, mountain ash, and Eucalyptus pauciflora, sno ...
, yellow-bellied glider,
sugar glider The sugar glider (''Petaurus breviceps'') is a small, omnivorous, arboreal, and nocturnal gliding possum. The common name refers to its predilection for sugary foods such as sap and nectar and its ability to glide through the air, much lik ...
,
mahogany glider The mahogany glider (''Petaurus gracilis'') is an endangered gliding possumsquirrel glider ** Family Phalangeridae: brushtail possums and cuscuses ** Family Phascolarctidae:
koala The koala (''Phascolarctos cinereus''), sometimes inaccurately called the koala bear, is an arboreal herbivorous marsupial native to Australia. It is the only Extant taxon, extant representative of the Family (biology), family ''Phascolar ...
** Family Potoridae: potoroos, rat kangaroos, bettongs ** Family Pseudocheiridae: ringtailed possums and allies ** Family Tarsipedidae: honey possum ** Family Vombatidae: wombats ** Family †
Diprotodontidae Diprotodontidae is an extinct family of large herbivorous marsupials, endemic to Australia and New Guinea during the Oligocene through Pleistocene periods from 28.4 million to 40,000 years ago. Description The family primarily consisted of lar ...
: giant wombats ** Family † Palorchestidae: marsupial tapirs ** Family † Thylacoleonidae: marsupial lions * Order Microbiotheria (1 species) ** Family Microbiotheriidae: monito del monte * Order Notoryctemorphia (2 species) ** Family Notoryctidae: marsupial moles * Order Peramelemorphia (21 species) ** Family
Peramelidae The marsupial family Peramelidae contains the extant bandicoots. They are found throughout Australia and New Guinea, with at least some species living in every available habitat, from rainforest to desert. Four fossil peramelids are described. O ...
: bandicoots and allies ** Family Thylacomyidae: bilbies ** Family †Chaeropodidae: pig-footed bandicoots * Order † Yalkaparidontia Archer, Hand & Godthelp 1988 ** Family † Yalkaparidontidae Archer, Hand & Godthelp 1988


References

{{Taxonbar, from=Q631975 Fauna of the Australasian realm Mammal superorders Extant Paleocene first appearances Marsupials of Australia Marsupials Metatheria Paleocene mammals of Australia Paleocene mammals of Oceania