Australian Warblers
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Acanthizidae—sometimes called Australian warblers—are a
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
of
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
birds which includes
gerygone ''Gerygone'' (), the gerygones or peep-warblers, is a genus of bird in the family Acanthizidae. The genus ranges from Southeast Asia through New Guinea and Australia to New Zealand and the Chatham Islands. Most of the species are found in Austr ...
s, thornbills ''
Acanthiza ''Acanthiza'' is a genus of perching bird, passeriform birds, most endemic to Australia, but with two species (''A. murina'' and ''A. cinerea'') restricted to New Guinea. These birds are commonly known as thornbills. They are not closely relat ...
'', and scrubwrens ''
Sericornis ''Sericornis'' is a genus of small, mainly insectivorous birds, the scrubwrens in the family Acanthizidae. Despite the similarity in shape and habits, the true wrens (Troglodytidae) are a quite unrelated group of passerines. The genus previous ...
''. The family Acanthizidae consists of small to medium passerine birds, with a total length varying between . They have short rounded wings, slender bills, long legs, and a short tail. Most species have olive, grey, or brown plumage, although some have patches of a brighter yellow. The
weebill The weebill (''Smicrornis brevirostris'') is a species of bird in the family Acanthizidae. It is an insectivorous passerine that is found throughout mainland Australia. At long, it is Australia's smallest bird. It was originally described by Joh ...
is the smallest species of acanthizid, and the smallest Australian passerine; the largest is the pilotbird.


Taxonomy and systematics

Following the Sibley-Ahlquist taxonomy (1990) they were previously regarded as subfamily Acanthizinae within the family Pardalotidae. More recent molecular genetic studies do not support this arrangement. The Dasyornithidae (which include the bristlebirds) are variously seen either as subfamily Dasyornithinae within the family Acanthizidae or Pardalotidae or as own family (Schodde & Mason 1999). A
molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...
study published in 2019 found that the family Acanthizidae is
sister A sister is a woman or a girl who shares parents or a parent with another individual; a female sibling. The male counterpart is a brother. Although the term typically refers to a familial relationship, it is sometimes used endearingly to ref ...
to the pardalotes in the small family Pardalotidae. The pardalotes are native to Australia.


List of genera

The family contains 67 species divided in 15 genera. *''
Pachycare The goldenface (''Pachycare flavogriseum'') is a species of passerine bird endemic to New Guinea. It is the only species (monotypic) within the genus ''Pachycare''. Taxonomy and systematics The placement of this species and genus within the pas ...
'' – Goldenface ''Pachycare flavogriseum'' *'' Oreoscopus'' – Fernwren ''Oreoscopus gutturalis'' *'' Acanthornis'' – Scrubtit ''Acanthornis magna'' *'' Smicrornis'' – Weebill ''Smicrornis brevirostris'' *'' Calamanthus'' – Fieldwrens (three species) *'' Hylacola'' – Heathwrens (two species) *'' Pycnoptilus'' – Pilotbird ''Pycnoptilus floccosus'' *''
Pyrrholaemus ''Pyrrholaemus'' is a genus of birds in the family Acanthizidae. The genus was introduced by the English ornithologist and bird artist John Gould in 1841. The name ''Pyrrholaemus'' is from classical Greek ''purrhos'' meaning "flame-coloured" or ...
'' – Redthroat and Speckled Warbler (two species) *''
Origma ''Origma'' is a genus of passerine birds in the family Acanthizidae. A molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gai ...
'' – Rockwarbler and two species of Mouse-Warblers (three species) *'' Neosericornis'' – Yellow-throated Scrubwren ''Neosericornis citreogularis'' *''
Aethomyias ''Aethomyias'' is a genus of passerine birds in the family Acanthizidae that are endemic to New Guinea. A molecular phylogenetic study of the scrubwrens and mouse-warblers published in 2018 led to a substantial revision of the taxonomic classifi ...
'' – Scrubwrens and Bicoloured Mouse-warbler (six species) *''
Sericornis ''Sericornis'' is a genus of small, mainly insectivorous birds, the scrubwrens in the family Acanthizidae. Despite the similarity in shape and habits, the true wrens (Troglodytidae) are a quite unrelated group of passerines. The genus previous ...
'' – Scrubwrens (eight species) *''
Gerygone ''Gerygone'' (), the gerygones or peep-warblers, is a genus of bird in the family Acanthizidae. The genus ranges from Southeast Asia through New Guinea and Australia to New Zealand and the Chatham Islands. Most of the species are found in Austr ...
'' – Gerygones (20 species including one now extinct) *''
Acanthiza ''Acanthiza'' is a genus of perching bird, passeriform birds, most endemic to Australia, but with two species (''A. murina'' and ''A. cinerea'') restricted to New Guinea. These birds are commonly known as thornbills. They are not closely relat ...
'' – Thornbills (14 species) *''
Aphelocephala ''Aphelocephala'' is a genus of bird in the thornbill family Acanthizidae. The three species are collectively known as whitefaces. They are endemic to Australia, generally occurring in the arid centre of the continent. They are a striking looking ...
'' – Whitefaces (three species)


Distribution and habitat

Acanthizids are native to Australia, Indonesia, New Zealand, and the southwest Pacific. The greatest diversity is found in Australia, 35
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
species, then New Guinea with 15. A species lives in
Vanuatu Vanuatu ( or ; ), officially the Republic of Vanuatu (; ), is an island country in Melanesia located in the South Pacific Ocean. The archipelago, which is of volcanic origin, is east of northern Australia, northeast of New Caledonia, east o ...
,
New Caledonia New Caledonia ( ; ) is a group of islands in the southwest Pacific Ocean, southwest of Vanuatu and east of Australia. Located from Metropolitan France, it forms a Overseas France#Sui generis collectivity, ''sui generis'' collectivity of t ...
and the
Solomon Islands Solomon Islands, also known simply as the Solomons,John Prados, ''Islands of Destiny'', Dutton Caliber, 2012, p,20 and passim is an island country consisting of six major islands and over 1000 smaller islands in Melanesia, part of Oceania, t ...
, and a further three in the
New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...
region, including endemic species in the
Chatham Islands The Chatham Islands ( ; Moriori language, Moriori: , 'Misty Sun'; ) are an archipelago in the Pacific Ocean about east of New Zealand's South Island, administered as part of New Zealand, and consisting of about 10 islands within an approxima ...
and
Norfolk Island Norfolk Island ( , ; ) is an States and territories of Australia, external territory of Australia located in the Pacific Ocean between New Zealand and New Caledonia, directly east of Australia's Evans Head, New South Wales, Evans Head and a ...
. In Asia two species are restricted to Indonesia and another is found in the
Philippines The Philippines, officially the Republic of the Philippines, is an Archipelagic state, archipelagic country in Southeast Asia. Located in the western Pacific Ocean, it consists of List of islands of the Philippines, 7,641 islands, with a tot ...
and on mainland Asia. Most species are sedentary, except the gerygones. The family occupies a range of habitats from
rainforest Rainforests are forests characterized by a closed and continuous tree Canopy (biology), canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforests can be generally classified as tropi ...
s to arid regions.


Behaviour and ecology

Most species are terrestrial, feeding primarily on insects, although also eating some seeds. In particular the whitefaces consume large numbers of seeds, and other species will take fruits. The secretions of sap-sucking insects are favoured by some species, as are the insects themselves. Some species are less terrestrial, such as the
weebill The weebill (''Smicrornis brevirostris'') is a species of bird in the family Acanthizidae. It is an insectivorous passerine that is found throughout mainland Australia. At long, it is Australia's smallest bird. It was originally described by Joh ...
, which forages in the treetops, or the rock-dwelling
rockwarbler The rockwarbler (''Origma solitaria''), is a bird in the family Acanthizidae. It is the only bird species endemic to the mainland of the state of New South Wales in Australia. Taxonomy and systematics English artist and naturalist John Lewin d ...
. Rainforest species lay one to two eggs in a clutch, and species in the deserts and Tasmania lay three to four. Acanthizids are unusual for passerines in their long incubation periods, which rival those of large songbirds like the
Corvidae Corvidae is a Cosmopolitan distribution, cosmopolitan Family (biology), family of Songbird, oscine passerine birds that contains the crows, ravens, Rook (bird), rooks, magpies, jackdaws, jays, treepies, choughs, and Nutcracker (bird), nutcrackers ...
. Also, despite their long incubation period hatching is completely synchronous and within-brood mortality completely absent. Acanthizids are relatively long-lived, with many species living to over ten years of age in the wild and
cooperative breeding Cooperative breeding is a social system characterized by alloparental care: offspring receive care not only from their parents, but also from additional group members, often called helpers. Cooperative breeding encompasses a wide variety of group s ...
is found in the weebill and with a lesser degree of development in all whitefaces and most species of ''Sericornis'' and ''Acanthiza''.Se
“Old endemics and new invaders: alternative strategies of passerines for living in the Australian environment”
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Status and conservation

Most taxa are considered as
least concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
. One species – the
Lord Howe gerygone The Lord Howe gerygone or Lord Howe gerygone flyeater (''Gerygone insularis'') was a small bird in the family Acanthizidae, brown and greyish in color. Its head was brown apart from a pale grey eye-ring and a grey throat and chin, many parts of ...
(''Gerygone insularis'') – became extinct by
rat Rats are various medium-sized, long-tailed rodents. Species of rats are found throughout the order Rodentia, but stereotypical rats are found in the genus ''Rattus''. Other rat genera include '' Neotoma'' (pack rats), '' Bandicota'' (bandicoo ...
predation in the early 1930s. The Norfolk Island gerygone (''Gerygone modesta'') is vulnerable and the
chestnut-breasted whiteface The chestnut-breasted whiteface (''Aphelocephala pectoralis'') is a species of bird in the family Acanthizidae. It is endemic to Australia. Its natural habitat is subtropical or tropical dry shrubland. It is threatened by habitat loss. Referenc ...
(''Aphelocephala pectoralis'') is near threatened.


Gallery

File:Scrub wren for wiki.jpg,
White-browed scrubwren The white-browed scrubwren (''Sericornis frontalis'') is a passerine bird found on the New England Tablelands and coastal areas of Australia. Placed in the family Pardalotidae in the Sibley-Ahlquist taxonomy, this has met with opposition and in ...
(''Sericornis frontalis'') File:Large-billed gerygone portland08.JPG, Large-billed gerygone (''Gerygone magnirostris'') File:Brown Gerygone lam08.jpg, Brown gerygone (''Gerygone mouki'') File:Smicrornis brevirostris.jpg,
Weebill The weebill (''Smicrornis brevirostris'') is a species of bird in the family Acanthizidae. It is an insectivorous passerine that is found throughout mainland Australia. At long, it is Australia's smallest bird. It was originally described by Joh ...
(''Smicrornis brevirostris'') File:Speckeld warbler.jpg,
Speckled warbler The speckled warbler (''Pyrrholaemus sagittatus'') is a species of bird in the family (biology), family Acanthizidae. It is Endemism, endemic to eastern Australia. Its natural habitat is temperate forests. Taxonomy The speckled warbler was first ...
(''Chthonicola sagittatus'') File:Hylacola cauta Wilpena Pound.JPG, Shy heathwren (''Hylacola cauta'') File:Yellowthornbill.jpg, Yellow thornbill (''Acanthiza nana'')


References


Further reading

* Christidis, L., and W.E. Boles. 1994. The taxonomy and species of Birds of Australia and its territories. R.A.O.U. Monograph 2: 1–112. * del Hoyo, J.; Elliot, A. & Christie D. (editors). (2007). Handbook of the Birds of the World. Volume 12: Picathartes to Tits and Chickadees. Lynx Edicions. * Mason, Ian J. & Schodde, Richard. 1999. The Directory of Australian Birds: Passerines. * Sibley, C.G., and J.E. Ahlquist. 1990. Phylogeny and Classification of Birds: A Study in Molecular Evolution. Yale Univ. Press, New Haven, CT. {{Taxonbar, from=Q782355 Bird families