Aucasaurus Garridoi By Paleocolour
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''Aucasaurus'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of medium-sized
abelisaurid Abelisauridae (meaning "Abel's lizards") is a family (or clade) of ceratosaurian theropod dinosaurs. Abelisaurids thrived during the Cretaceous period, on the ancient southern supercontinent of Gondwana, and today their fossil remains are foun ...
theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
from
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
that lived during the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
(
Santonian The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya ( million years ago) and 83.6 ± 0.7 m ...
to
Campanian The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
stage Stage, stages, or staging may refer to: Arts and media Acting * Stage (theatre), a space for the performance of theatrical productions * Theatre, a branch of the performing arts, often referred to as "the stage" * ''The Stage'', a weekly Brit ...
) of the
Anacleto Formation The Anacleto Formation is a geologic formation with outcrops in the Argentina, Argentine Patagonian provinces of Mendoza Province, Mendoza, Río Negro Province, Río Negro, and Neuquén Province, Neuquén. It is the youngest formation within the ...
. It was smaller than the related ''
Carnotaurus ''Carnotaurus'' (; ) is a genus of Theropoda, theropod dinosaur that lived in South America during the Late Cretaceous period, probably sometime between 72 and 69 million years ago. The only species is ''Carnotaurus sastrei''. Known from a si ...
'', although more derived in some ways, such as its extremely reduced arms and almost total lack of fingers. The type skeleton is complete to the thirteenth
caudal vertebra Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spinal ...
, and so is relatively well understood, and was the most complete
abelisaurid Abelisauridae (meaning "Abel's lizards") is a family (or clade) of ceratosaurian theropod dinosaurs. Abelisaurids thrived during the Cretaceous period, on the ancient southern supercontinent of Gondwana, and today their fossil remains are foun ...
known when described in 2002. However, the skull is damaged, causing some
paleontologist Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
s to speculate that it was involved in a fight prior to death.


Discovery

The holotype of ''Aucasaurus'' is known from finds in the Río Colorado Subgroup, a
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
group comprising the
Anacleto Formation The Anacleto Formation is a geologic formation with outcrops in the Argentina, Argentine Patagonian provinces of Mendoza Province, Mendoza, Río Negro Province, Río Negro, and Neuquén Province, Neuquén. It is the youngest formation within the ...
in the
Neuquén Basin Neuquén Basin () is a sedimentary basin covering most of Neuquén Province in Argentina. The basin originated in the Jurassic and developed through alternating continental and marine conditions well into the Tertiary. The basin bounds to the wes ...
of
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
that has yielded many
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
s. Numerous
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
egg An egg is an organic vessel grown by an animal to carry a possibly fertilized egg cell (a zygote) and to incubate from it an embryo within the egg until the embryo has become an animal fetus that can survive on its own, at which point the ...
s are also known from this deposit. The type specimen belongs to a mature individual of at least eleven years old.


Description

''Aucasaurus'' was a medium-sized abelisaur, measuring long and weighing approximately . Its skull was not as short or as deep-snouted as that of ''
Carnotaurus ''Carnotaurus'' (; ) is a genus of Theropoda, theropod dinosaur that lived in South America during the Late Cretaceous period, probably sometime between 72 and 69 million years ago. The only species is ''Carnotaurus sastrei''. Known from a si ...
''. Also, instead of horns, it had a pair of low ridges above each eye.


Postcranial skeleton

The small arms of ''Aucasaurus'' were also like that of its horned relative, but were proportionally longer due to its small size, and the bones lacked the bony processes and some unusual proportions present in ''Carnotaurus''. The
hand A hand is a prehensile, multi-fingered appendage located at the end of the forearm or forelimb of primates such as humans, chimpanzees, monkeys, and lemurs. A few other vertebrates such as the Koala#Characteristics, koala (which has two thumb#O ...
of ''Aucasaurus'' was unusual: four
metacarpals In human anatomy, the metacarpal bones or metacarpus, also known as the "palm bones", are the appendicular skeleton, appendicular bones that form the intermediate part of the hand between the phalanges (fingers) and the carpal bones (wrist, wris ...
were present, but the first and fourth lacked fingers. The second and third had fingers, but they were quite short and had no claws. Traits found in the caudal vertebrae suggests a significant caudal musculature and rigidity, possibly indicating that it could have been a fast runner.


Braincase

A study was done on the braincase of ''Aucasaurus'' in 2015 by Ariana Paulina-Carabajal and Cecilia Succar, in which the skull material was scanned using a medical CT machine. Virtual three-dimensional inner ear and cranial endocasts were obtained and visualized using the imagine software at the
University of Alberta The University of Alberta (also known as U of A or UAlberta, ) is a public research university located in Edmonton, Alberta, Canada. It was founded in 1908 by Alexander Cameron Rutherford, the first premier of Alberta, and Henry Marshall Tory, t ...
. A latex cranial
endocast An endocast is the internal cast of a hollow object, often referring to the cranial vault in the study of brain development in humans and other organisms. Endocasts can be artificially made for examining the properties of a hollow, inaccessible ...
was also made. The forebrain, midbrain, and hindbrain resemble the morphology described for the abelisaurids ''
Majungasaurus ''Majungasaurus'' (; ) is a genus of abelisaurid theropod dinosaur that lived in Madagascar from 70 to 66 million years ago, at the end of the Cretaceous Period, making it one of the last-known non-avian dinosaurs that went extinct during th ...
'' and ''
Indosaurus ''Indosaurus'' () is a genus of dubious carnivorous theropod dinosaur that lived in what is now India, about 69 to 66 million years ago during the Maastrichtian division of the Late Cretaceous. Discovery and naming The now-lost holotype wa ...
''. However, ''Aucasaurus'' exhibits a floccular process that is relatively larger than that of ''Majungasaurus''. In ''Aucasaurus'' the
flocculus The flocculus (Latin: ''tuft of wool'', diminutive) is a small lobe of the cerebellum at the posterior border of the middle cerebellar peduncle anterior to the biventer lobule. Like other parts of the cerebellum, the flocculus is involved in moto ...
is enclosed in an 8-shaped floccular recess, similar in shape and size to that observed in ''Abelisaurus'', suggesting that the two Patagonian taxa were capable of a slightly wider range of movements of the head. The labyrinth of the inner ear is similar in shape and size to the semicircular canals of ''Majungasaurus'', although the lateral semicircular canal is shorter in ''Aucasaurus''.


Pathology

The 5th and 6th holotype tail vertebrae of ''Aucasaurus'' provide evidence of failed developmental vertebral segmentation, so two of the vertebrae are fused. Baiano ''et al''. (2024) interpret this as a
congenital disorder A birth defect is an abnormal condition that is present at childbirth, birth, regardless of its cause. Birth defects may result in disability, disabilities that may be physical disability, physical, intellectual disability, intellectual, or dev ...
(birth defect) called
block vertebrae Congenital vertebral anomalies are a collection of malformations of the spine. Most, around 85%, are not clinically significant, but they can cause compression of the spinal cord by deforming the vertebral canal or causing instability. This condi ...
. This represents the earliest known occurrence of this
pathology Pathology is the study of disease. The word ''pathology'' also refers to the study of disease in general, incorporating a wide range of biology research fields and medical practices. However, when used in the context of modern medical treatme ...
in the non-avian theropod fossil record.


Classification

In 2009, Novas suggested that ''Aucasaurus garridoi'' might be a
junior synonym In taxonomy, the scientific classification of living organisms, a synonym is an alternative scientific name for the accepted scientific name of a taxon. The botanical and zoological codes of nomenclature treat the concept of synonymy differently. ...
of ''
Abelisaurus ''Abelisaurus'' (; "Abel's lizard") is a genus of predatory abelisaurid theropod dinosaur alive during the Late Cretaceous Period (geology), Period (Campanian) of what is now South America. It was a bipedal carnivore that probably reached about ...
comahuensis''. In 2010,
Gregory S. Paul Gregory Scott Paul (born December 24, 1954) is an American freelance researcher, author and illustrator who works in paleontology. He is best known for his work and research on theropoda, theropod dinosaurs and his detailed illustrations, both l ...
renamed ''Aucasaurus garridoi'' into ''Abelisaurus garridoi''. Despite their similarities, other researchers have placed both genera as separate genera, and subsequent studies suggest that ''Aucasaurus'' was more closely related to other taxa such as ''Carnotaurus'', with some including them in the
Carnotaurini Brachyrostra (meaning "short snouts") is a clade within the theropod dinosaur family Abelisauridae. It includes the famous genera ''Carnotaurus'', '' Aucasaurus'', potentially '' Abelisaurus'' as well as their close relatives from the Cretaceous ...
. Below is a cladogram by Canalle ''et al.'' in 2009.


See also

*
Timeline of ceratosaur research This timeline of ceratosaur research is a chronological listing of events in the History of paleontology, history of paleontology focused on the ceratosaurs, a group of relatively primitive, often horned, predatory theropod dinosaurs that became ...


References

{{Portal bar, Dinosaurs Abelisauridae Dinosaur genera Santonian dinosaurs Anacleto Formation Fossil taxa described in 2002 Taxa named by Rodolfo Coria Taxa named by Luis M. Chiappe Dinosaurs of Argentina