''Atherina'' is a

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of fish of

silverside Silverside may refer to: * Silverside (beef), a cut of beef * Silverside (fish), the Atheriniformes, an order of fish * Agnes Silverside (died 1557), one of the Colchester Martyrs * USS ''Silversides'', the name of 2 U.S. Navy submarines * Silvers ...

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Atherinidae The Old World silversides are a family, Atherinidae, of fish in the order Atheriniformes. Atherinidae are abundant and considered bony fish (teleost) that are widespread globally, living in rivers, estuaries, and coastal waters. They occur worldw ...

, found in the temperate and tropic zones. Up to long, they are widespread in the

Mediterranean The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern ...

Black Sea The Black Sea is a marginal sea, marginal Mediterranean sea (oceanography), mediterranean sea lying between Europe and Asia, east of the Balkans, south of the East European Plain, west of the Caucasus, and north of Anatolia. It is bound ...

and

Sea of Azov The Sea of Azov is an inland Continental shelf#Shelf seas, shelf sea in Eastern Europe connected to the Black Sea by the narrow (about ) Strait of Kerch, and sometimes regarded as a northern extension of the Black Sea. The sea is bounded by Ru ...

; in

lagoon A lagoon is a shallow body of water separated from a larger body of water by a narrow landform, such as reefs, barrier islands, barrier peninsulas, or isthmuses. Lagoons are commonly divided into ''coastal lagoons'' (or ''barrier lagoons'') an ...

s such as

Syvash The Syvash or Sivash ( Russian and Ukrainian: ; , Cyrillic: Сываш, "dirt"), also known as the or (, ''Gniloye More''; , ''Hnyle More''; ), is a large area of shallow lagoons on the western edge of the Sea of Azov. Separated from the sea ...

in Ukraine; and

estuaries An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environm ...

. They enter the lower reaches of the

Dnieper The Dnieper or Dnepr ( ), also called Dnipro ( ), is one of the major transboundary rivers of Europe, rising in the Valdai Hills near Smolensk, Russia, before flowing through Belarus and Ukraine to the Black Sea. Approximately long, with ...

Southern Bug The Southern Bug, also called Southern Buh (; ; ; or just ), and sometimes Boh River (; ),

Dniester The Dniester ( ) is a transboundary river in Eastern Europe. It runs first through Ukraine and then through Moldova (from which it more or less separates the breakaway territory of Transnistria), finally discharging into the Black Sea on Uk ...

and

Danube River The Danube ( ; see also other names) is the second-longest river in Europe, after the Volga in Russia. It flows through Central and Southeastern Europe, from the Black Forest south into the Black Sea. A large and historically important riv ...

s. They can also be found in the freshwater Lake Trichonis of

Greece Greece, officially the Hellenic Republic, is a country in Southeast Europe. Located on the southern tip of the Balkan peninsula, it shares land borders with Albania to the northwest, North Macedonia and Bulgaria to the north, and Turkey to th ...

, and there is an isolated population in the

Caspian Sea The Caspian Sea is the world's largest inland body of water, described as the List of lakes by area, world's largest lake and usually referred to as a full-fledged sea. An endorheic basin, it lies between Europe and Asia: east of the Caucasus, ...

Species

There are currently seven recognized species in this genus:> * '' Atherina boyeri'' A. Risso, 1810 (Big-scale sand smelt) * '' Atherina breviceps''

Valenciennes Valenciennes (, also , , ; ; or ; ) is a communes of France, commune in the Nord (French department), Nord Departments of France, department, Hauts-de-France, France. It lies on the Scheldt () river. Although the city and region experienced ...

, 1835 (Cape silverside) * '' Atherina caspia'' Eichwald, 1831 * ''

Atherina harringtonensis ''Atherina'' is a genus of fish of Old World silverside, silverside family (biology), family Atherinidae, found in the temperate and tropic zones. Up to long, they are widespread in the Mediterranean Sea, Mediterranean, Black Sea and Sea of Azov ...

'' Goode, 1877 * '' Atherina hepsetus''

Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...

, 1758 (Mediterranean sand smelt) * '' Atherina lopeziana'' Rossignol & Blache, 1961 * '' Atherina presbyter'' G. Cuvier, 1829 (Sand smelt)

Commercial importance

Species of ''Atherina'' feature in the traditional

Italian Italian(s) may refer to: * Anything of, from, or related to the people of Italy over the centuries ** Italians, a Romance ethnic group related to or simply a citizen of the Italian Republic or Italian Kingdom ** Italian language, a Romance languag ...

, Catalan,

Occitan Occitan may refer to: * Something of, from, or related to the Occitania territory in parts of France, Italy, Monaco and Spain. * Something of, from, or related to the Occitania administrative region of France. * Occitan language, spoken in parts o ...

, south- Ukrainian, Turkish, and

Greek cuisine Greek cuisine is the cuisine of Greece and the Greek diaspora. In common with many other Mediterranean cuisine, cuisines of the Mediterranean, it is founded on the triad of wheat, olive oil, and wine. It uses vegetables, olive oil, grains ...

s in fried form. The fish are lightly powdered with

wheat flour Wheat flour is a powder made from the grinding of common wheat used for human consumption. Wheat varieties are called "soft" or "weak" if gluten content is low, and are called "hard" or "strong" if they have high gluten content. Hard flour, or ...

before being dropped in hot

olive oil Olive oil is a vegetable oil obtained by pressing whole olives (the fruit of ''Olea europaea'', a traditional Tree fruit, tree crop of the Mediterranean Basin) and extracting the oil. It is commonly used in cooking for frying foods, as a cond ...

. In

Ukraine Ukraine is a country in Eastern Europe. It is the List of European countries by area, second-largest country in Europe after Russia, which Russia–Ukraine border, borders it to the east and northeast. Ukraine also borders Belarus to the nor ...

and

, it is commercially important.

Fossil record

*''Atherina austriaca'' - Badenian of Czechia (

Mikulov Mikulov (; ) is a town in Břeclav District in the South Moravian Region of the Czech Republic. It has about 7,600 inhabitants. From the 16th to the 19th century, it was the cultural centre of the Jewish community of Moravia. The historic centr ...

) : 13.82 - 12.8 mya *''Atherina atropatiensis'' -

Tortonian The Tortonian is in the geologic time scale an age or stage of the late Miocene that spans the time between 11.608 ± 0.005 Ma and 7.246 ± 0.005 Ma (million years ago). It follows the Serravallian and is followed by the Messinian. The Tort ...

of Iran *''Atherina cavalloi'' -

Messinian The Messinian is in the geologic timescale the last age or uppermost stage of the Miocene. It spans the time between 7.246 ± 0.005 Ma and 5.333 ± 0.005 Ma (million years ago). It follows the Tortonian and is followed by the Zanclean, the fir ...

of Italy (Emilia-Romagna) *''Atherina gidjakensis'' -

Serravallian The Serravallian is, in the geologic timescale, an List of time periods, age or a stage (stratigraphy), stage in the middle Miocene epoch (geology), Epoch/series (stratigraphy), Series, which spans the time between 13.82 annum, Ma and 11.63 Ma (m ...

of Kazakhstan *''Atherina macrocephala'' -

Ypresian In the geologic timescale the Ypresian is the oldest age (geology), age or lowest stage (stratigraphy), stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by th ...

of Italy *''Atherina suchovi'' - Sarmatian of Moldova : 12.8 - 11.63 mya Also, fossils of the extant ''Atherina boyeri'' have been found at several sites in Italy (Tuscany) and the Greek Islands of Rhodes and Crete, dating back as far as the

References

Atherininae Ray-finned fish genera Taxa named by Carl Linnaeus Taxonomy articles created by Polbot {{Atheriniformes-stub